Purslane Portulaca oleracea, photographed by Glazunov Lena.

Belongs within: Caryophyllales.
Contains: Opuntia.

The Cactineae is a clade of flowering plants strongly supported by molecular analyses. Among its members are the cacti (Cactaceae), the purslanes (Portulacaceae) and the Madagascar-endemic Didereaceae. Many members of the clade are succulent to some degree.

    |--Basellaceae AW03
    |    |--Basella rubra PT01
    |    `--Anredera cordifolia H06
    |--Hectorellaceae T00
    |    |--Hectorella caespitosa T00, A27
    |    `--Lyallia T00
    |--Didiereaceae AW03
    |    |--Portulacarioideae [Portulacarieae] AW03
    |    |    |--Portulacaria Jacquin 1787 AW03
    |    |    `--Ceraria Pearson & Stephens 1912 AW03
    |    |         |--C. longipedunculata CV06
    |    |         `--C. namaquensis AW03
    |    `--+--Calyptrotheca Gilg 1897 [Calyptrothecoideae] AW03
    |       `--Didiereoideae AW03
    |            |--Didierea Baillon 1880 AW03
    |            |--Alluaudia (Drake) Drake 1903 AW03
    |            |--Alluaudiopsis Humbert & Choux 1934 AW03
    |            `--Decaryia Choux 1934 AW03
    |--Cactaceae AW03
    |    |  i. s.: Mammillaria leucocentra PT01
    |    |         Epiphyllum oxypetalum P88
    |    |         Pilosocereus tillianus K06
    |    |         Stenocereus griseus K06
    |    |         Subpilocereus repandus K06
    |    |         Schlumbergera × buckleyi MH98
    |    |         Lophocereus schotti GE05
    |    |         Rhipsalis baccifera H06
    |    |         Echinopsis spachiana H06
    |    |         Cereus jamacaru H06
    |    |         Harrisia martinii H06
    |    |         Echinocactus wislizenni LW70
    |    |--Pereskia [Pereskioideae] T00
    |    |    `--P. aculeata H06
    |    |--Maihuenia [Maihuenioideae] T00
    |    |--Cactoideae T00
    |    `--Opuntia T00
    `--Portulacaceae AW03
         |--Montia G95
         |    |--M. australasica GK00
         |    `--M. fontana Linnaeus 1753 [incl. M. lamprosperma Cham. 1831] G95
         |--Talinum G95
         |    |--T. paniculatum (Jacq.) Gaertn. 1791 (see below for synonymy) G95
         |    `--T. triangulare (Jacq.) Willdenow in Linnaeus 1799 (see below for synonymy) G95
         |--Claytonia PT01
         |    |--C. perfoliata PT01
         |    `--C. virginica CTP01
         |--Calandrinia OS04
         |    |--C. calyptrata OS04
         |    |--C. composita GK00
         |    |--C. eremaea G04
         |    |--C. granulifera GK00
         |    `--C. umbellata PT01
         |--Portulaca G95
         |    |  i. s.: P. meridiana VB02
         |    |--P. sect. Portulaca G95
         |    |    |--P. subsect. Portulaca G95
         |    |    |    |--P. lutea Forst. ex Seem. 1865 G95
         |    |    |    `--P. oleracea Linnaeus 1753 G95
         |    |    |         |--P. o. var. oleracea C55a
         |    |    |         `--P. o. var. sylvestris C55a
         |    |    `--P. (subsect. Stellulatotuberculatae) pilosa Linnaeus 1753 (see below for synonymy) G95
         |    `--P. (sect. Neossia) quadrifida Linnaeus 1767 G95
         |--Anacampseros filamentosa CV06
         |    |--A. f. ssp. filamentosa CV06
         |    `--A. f. ssp. tomentosa CV06
         `--Avonia dinteri [=Anacampseros dinteri] CV06

Portulaca pilosa Linnaeus 1753 [incl. P. samoensis Poelln. 1933, P. tuberosa Roxburgh 1814] G95

Talinum paniculatum (Jacq.) Gaertn. 1791 [=Portulaca paniculatum Jacq. 1760; incl. T. patens (Linnaeus) Willdenow 1799] G95

Talinum triangulare (Jacq.) Willdenow in Linnaeus 1799 [=Portulaca triangulare Jacq. 1760; incl. P. racemosa Linnaeus 1771] G95


[A27] Andersen, J. C. 1927. Popular names of New Zealand plants. Transactions and Proceedings of the New Zealand Institute 57: 905-977.

[AW03] Applequist, W. L., & R. S. Wallace. 2003. Expanded circumscription of Didiereaceae and its division into three subfamilies. Adansonia 25 (1): 13-16.

[C55] Candolle, A. de. 1855. Géographie Botanique Raisonée: Ou exposition des faits principaux et des lois concernant la distribution géographique des plantes de l’époque actuelle vol. 1. Librairie de Victor Masson: Paris.

[CV06] Craven, P., & P. Vorster. 2006. Patterns of plant diversity and endemism in Namibia. Bothalia 36 (2): 175-189.

[CTP01] Cubas, P., H. Tahiri & C. Pardo. 2001. Karyological and taxonomic notes on Cytisus Desf. sect. Spartopsis Dumort. and sect. Alburnoides DC. (Genisteae, Leguminosae) from the Iberian Peninsula and Morocco. Botanical Journal of the Linnean Society 135 (1): 43-50.

[G04] Gibson, N. 2004. Flora and vegetation of the Eastern Goldfields Ranges: part 6. Mt Manning Range. Journal of the Royal Society of Western Australia 87 (2): 35-47.

[GK00] Gibson, N., & G. J. Keighery. 2000. Flora and vegetation of the Byenup-Muir reserve system, south-west Western Australia. CALMScience 3 (3): 323-402.

[G95] Gideon, O. 1995. Portulacaceae. In Handbooks of the Flora of Papua New Guinea vol. 3 (B. J. Conn, ed.) pp. 212-220. Melbourne University Press: Carlton (Australia).

[GE05] Grimaldi, D., & M. S. Engel. 2005. Evolution of the Insects. Cambridge University Press: New York.

[H06] Henderson, L. 2006. Comparisons of invasive plants in southern Africa originating from southern temperate, northern temperate and tropical regions. Bothalia 36 (2): 201-222.

[K06] Kwiecinski, G. G. 2006. Phyllostomus discolor. Mammalian Species 801: 1-11.

[LW70] Lowe, C. H., J. W. Wright, C. J. Cole & R. L. Bezy. 1970. Natural hybridization between the teiid lizards Cnemidophorus sonorae (parthenogenetic) and Cnemidophorus tigris (bisexual). Systematic Zoology 19 (2): 114-127.

[MH98] Morikawa, H., A. Higaki, M. Nohno, M. Takahashi, M. Kamada, M. Nakata, G. Toyohara, Y. Okamura, K. Matsui, S. Kitani, K. Fujita, K. Irifune & N. Goshima. 1998. More than a 600-fold variation in nitrogen dioxide assimilation among 217 plant taxa. Plant, Cell and Environment 21: 180-190.

[OS04] Obbens, F. J., & L. W. Sage. 2004. Vegetation and flora of a diverse upland remnant of the Western Australian wheatbelt (Nature Reserve A21064). Journal of the Royal Society of Western Australia 87 (1): 19-28.

[PT01] Pemberton, L. M. S., S.-L. Tsai, P. H. Lovell & P. J. Harris. 2001. Epidermal patterning in seedling roots of eudicotyledons. Annals of Botany 87: 649-654.

[P88] Polunin, I. 1988. Plants and Flowers of Malaysia. Times Editions: Singapore.

[T00] Thorne, R. F. 2000. The classification and geography of the flowering plants: Dicotyledons of the class Angiospermae (subclasses Magnoliidae, Ranunculidae, Caryophyllidae, Dilleniidae, Rosidae, Asteridae, and Lamiidae). The Botanical Review 66: 441-647.

[VB02] Vijay, S. K., & T. N. Bhardwaja. 2002. Vegetation and phenodynamics of wetlands of central Rajasthan. Journal of the Bombay Natural History Society 99 (3): 573-581.

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