Diagnostic figures of generalised Copidognathus, from Bartsch (2006).

Belongs within: Halacaridae.

Copidognathus is a large genus of primarily marine mites (though a few live in fresh water). Copidognathus species are found on a wide variety of substrates.

Characters (from Bartsch 2006): Idiosoma: Length 180–700 μm. Dorsal and ventral plates generally large (or even fused). Plates smooth or heavily ornamented, panelled and with raised circular or subquadrangular areolae and longitudinal costae; areolae and costae with rosette pores or uniform porosity. With 4 pairs of gland pores which may be small, almost inconspicuous, or large and opening on raised cones. Dorsum with 6 pairs of dorsal setae; adanal setae on anal plate. Anterior epimeral plate with 3 pairs of ventral setae and pair of epimeral pores (rarely with small internal vesicles); posterior epimeral plate generally with 1 dorsal and 3 ventral setae. Female genitoanal plate with 3(–5) pairs of perigenital setae and 1 pair of subgenital setae. Male genitoanla plate with 4–40 pairs of perigenital setae; genital sclerites with 3–4 (rarely 2) pairs of subgenital setae. Gnathosoma: Longer than wide, rarely about as long as wide. Shape of rostrum varying from short (half length of gnathosomal base) and triangular to long, slender and parallel-sided (longer than gnathosomal base). Tectum truncate, with median process or huge crest. Normally with 2 pairs of maxillary setae, 1 pair inserted on gnathosomal base, 1 pair on rostrum; a few species with 2–7 pairs of maxillary setae on gnathosomal base. Palps 4-segmented, attached laterally and at least slightly surpassing rostrum. Second palpal segment with 1 distal seta. No seta on third palpal segment (there may be a small spiniform cuticular process which is no seta because an alveolus is lacking). Fourth palpal segment with 3 setae in basal whorl; apically with setula and 2 spurs. Legs: From short and wide to long and slender in shape; a few species with large lamellae. Genua shorter than telofemora and tibiae. Basifemora I to IV with 2,2,(1–)2,(1–)2 setae. Genua I–IV with 4(–3),4(–3),3,3–4(–5) setae. Tibia I with 3 ventral setae, often 1 of these setae bristle-like and smooth and 2 setae pectinate or spiniform. Tibia II with 3 ventral setae (1 species with 2 setae), in general 1 seta slender and smooth, 2 setae bipectinate. Tibia III with 2 ventral setae, often 1 slender and smooth and 1 bipectinate. Tibia IV either with 1 smooth and 1 bipectinate or with 2 smooth setae. In general, tibiae I and II with 4 dorsal setae and tibiae III and IV with 3 such setae. Tarsus I with 3 dorsal setae, dorsolateral seti- or baculiform solenidion, lamellar, rarely digitiform famulus, 3 ventral setae, and pair of parambulacral setae. Tarsus II with 3 dorsal setae, setiform dorsolateral solenidion, no ventral seta but pair of parambulacral setae. Tarsi III and IV with 3–4 dorsal setae, no ventral setae, and pair of parambulacral setae (rarely 1 of parambulacral setae moved to ventral position or lacking); parambulacral setae setiform or short and spur-like. Paired claws large; median claw present, though small. Claws of leg I may be shorter than those of following tarsi. Claws smooth or with accessory process, with or without pecten.

<==Copidognathus Trouessart 1888 B86
    |--C. fabriciusi M62
    |--C. gibboides Bartsch 1977 B86
    |--C. hartmanni Bartsch 1972 B86
    |--C. humerosus M62
    |--C. lepidus Bartsch 1977 B86
    |--C. lineatus Bartsch 1977 B86
    |--C. paucisporus Bartsch 1977 B86
    |--C. pygmaeus Bartsch 1980 B86
    |--C. rhodostigma M62
    `--C. tabellio M62

*Type species of generic name indicated


[B86] Bartsch, I. 1986. Acari: Halacaridae. In Stygofauna Mundi: A Faunistic, Distributional, and Ecological Synthesis of the World Fauna inhabiting Subterranean Waters (including the Marine Interstitial) (L. Botosaneanu, ed.) pp. 638-642. E. J. Brill / Dr. W. Backhuys: Leiden.

Bartsch, I. 2006. Halacaroidea (Acari): a guide to marine genera. Organisms Diversity and Evolution 6 (Electronic Suppl. 6): 1-104.

[M62] Monniot, F. 1962. Recherches sur les graviers a Amphioxus de la région de Banyuls-sur-Mer. Vie et Milieu 13: 231-322.

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