Fossil of Dasyleptus sp. from the Mazon Creek, from RCFossils.
Belongs within: Panhexapoda.
Contains: Zygentoma,
Pterygota, Meinertellidae, Machilidae.
One of the most speciose of all groups of organisms, the insects dominate the terrestrial environment. The majority of insects belong to the winged clade Pterygota, but some basal clades retain the primitive wingless morphology. These latter groups, the Archaeognatha and Zygentoma, are superficially similar in having a more or less elongate body form with three long terminal filaments (the cerci and median filament), and so have been combined in the past under the name Thysanura. However, the Zygentoma share a number of synapomorphies with the Pterygota, including the presence of mandibles with a secondary articulation in addition to the single articulation found in Archaeognatha, and are now recognised a forming a clade with the winged insects (Grimaldi & Engel 2005). The Archaeognatha, bristletails, are characterised by large eyes that meet on the top of the head and a distinctly humped thorax. They jump by rapidly flexing the abdomen to propel themselves into the aire (Grimaldi & Engel 2005).
Synapomorphies (from Grimaldi & Engel 2005, many altered in derived subgroups): Antenna lacking musculature beyond the scape; chordotonal organ (Johnston's organ) present in antennal pedicel; posterior tentorium developed into transverse bar; articulations absent between coxae and sterna; tarsus subsegmented into tarsomeres; pretarsal claws articulating with apicalmost tarsomere rather than with tarsal base; ovipositor present in females formed from gonapophyses on eighth and ninth abdominal segments; long terminal filament present on dorsum of eleventh abdominal segment.
<==Insecta [Ectognatha, Ectotropha, Lepismatona, Scarabaeoda, Thysanura, Thysanuroidea]
|--Dicondylia [Cercofilata] GE05
| |--Tricholepidion gertschi Wygodzinsky 1961 TW05, KZ02
| `--+--Zygentoma TW05
| `--Pterygota TW05
`--Archaeognatha [Machilida, Machiloidea, Microcoryphia, Monocondylia] TW05
| i. s.: Triassomachilis Sharov 1948 KS02 [Triassomachilidae GE05]
| `--T. uralensis GE05
|--Dasyleptus Brongniart 1885 FT05 [Cercopodata, Dasyleptidae R02, Monura]
| |--D. brongniarti Sharov 1957 K-P83
| `--D. sharovi GE05
`--+--Meinertellidae R02
`--Machilidae R02
Insecta incertae sedis:
Drosicha mangiferae G01
Clavigralla gibbosa G01
Ramsdelepidion Kukalová-Peck 1987 K-P87
`--*R. schusteri Kukalová-Peck 1987 K-P87
Gaspea palaeoentognathae Z02
Biorhiza pallida A71
Neocolochelyna itoi Takeuchi 1951 I92
Paratlanticus tsushimensis Yamasaki 1986 I92
Cercomantispa shirozui Nakahara 1961 I92
Armatocillenus sumaoi (Morita 1981) I92
Heplochlaenius insularis Uéno 1964 I92
Hydrochus chubu Balfour-Browne & Satô 1962 I92
Georissus granulosus Satô 1972 I92
Liparocephalus tokunagai Sakaguchi 1944 I92
Schwarzerium viridicyaneum Hayashi 1956 I92
Baryrhynchus tokarensis Ohbayashi & Satô 1966 I92
Cyphagogus iwatensis Morimoto 1976 I92
Ogasawarazo daitoensis Voss 1971 I92
Xyelecia japonica Tagashi 1972 I92
Pleroneura hikosana Togashi 1972 I92
Nipponorhynchus I92
|--N. bimaculatus Naito 1973 I92
`--N. mirabilis Takeuchi 1941 I92
Caenosclerogibba japonica Yasumatsu 1958 I92
Pararrhynchium tsunekii Tano & Yamane 1983 I92
Liacos melanogaster Tsuneki 1982 I92
Strymonidia iyonis Ota & Kusunoki 1957 I92
Pithecops I92
|--P. corvus I92
| |--P. c. corvus I92
| `--P. c. ryukyuensis Shirôzu 1964 I92
`--P. fulgens I92
|--P. f. fulgens I92
`--P. f. tsushimanus Shirôzu & Urata 1957 I92
Tongeia fischeri (Eversmann 1843) I92
Archiearis notha (Hübner 1808) I92
Heliophobus texturatus (Alpheraky 1892) I92
Cratosomus flavofasciatus H91
Eldana saccharina H91
Maruca testulalis H91
Tricopria drosophilae GJR97
Asobara tabida GJR97
Aramigus tesselatus GJR97
Leptopilina GJR97
|--L. australis GJR97
`--L. clavipes GJR97
Schizaphis graminum SF98
Leptophyllus Hope 1842 FT93
Amaurocoris laticeps G89
Aspongopus viduatus G89
Nariscus spinosus G89
Spilostethus G89
|--S. macilentus G89
`--S. militaris G89
Anepsius encaustus G89
Ischnopeza hirticornis G89
Piezostethus galactinus G89
Samea multiplicalis B04
Neohydronomus affinus MHG04
Araecerus fasciculatus G17a
Lecanium G25
|--L. baccatum G25
|--L. capraea G17d
`--L. viride G17a
Stictococcus gowdeyi G17b
Tachardia decorella G17b
Physokermes insignicola G17b
Eutettix tenella G17b
Dibolia borealis G17b
Tephrocerus gracilis G17b
Evetria taxifoliella G17c
Dioryctria xanthaenobares G17c
Chelymorpha argus G17c
Eudamus proteus G18
Melitara junctolineella B88
Moneilema ulkei G25
Mytilaspis cordylinidis G29
Cylindrococcus B88
|--C. amplior G34
`--C. casuarinae B88
Proceras sacchariphagum B88
Hypolycaena phorbas B88
Stomopteryx simplexella B88
Diplosis sorghicola B88
Tryporyza VHK02
|--T. incertulas B88
|--T. monostigma VHK02
|--T. nivella VHK02
`--T. rhodoproctalis VHK02
Actinoscytina belmontensis Tillyard 1926 F71
Adelidium cordatum Tillyard 1917 F71
Agetochoristella adscita Riek 1952 F71
Anomaloscytina F71
|--A. incompleta Evans 1943 F71
`--A. metapteryx Davis 1942 F71
Aphryganoneura anomala Tillyard 1926 F71
Archaeosmylus pectinatus Riek 1952 F71
Atelophlebis culleni (Etheridge & Olliff 1890) [=Ephemera culleni] F71
Austroidelia perplexa Riek 1954 F71
Austroprosbole maculata Evans 1943 F71
Austroscytina imperfecta Evans 1943 F71
Beaconiella F71
|--B. fennahi Evans 1963 F71
`--B. multivenata Evans 1963 F71
Belpsylla reticulata Evans 1943 F71
Clavopsyllidium minutum Davis 1942 F71
Elateridium angustius Tillyard 1917 F71
Eochiliocycla angusta Davis 1942 F71
Eopsyllidium delicatulum Davis 1942 F71
Eupincombea postiva Davis 1942 F71
Homaloscytina plana Tillyard 1926 F71
Mesonotoperla sinuata Riek 1954 F71
Mesophlebia antinodalis Tillyard 1916 F71
Mesothyris westraliensis Riek 1968 F71
Metrorhynchites grandis Tillyard 1917 F71
Mitchelloneura permiana Tillyard 1921 F71
Nannochoristella reducta Riek 1953 F71
Neoageta elongata Riek 1953 F71
Neochoristella aptata Riek 1953 F71
Neopetromantis australis Riek 1953 F71
Notoblattites F71
|--N. mitchelli Tillyard 1917 F71
`--N. wianamattensis Tillyard 1917 F71
Orthoscytina F71
|--O. belmontensis Tillyard 1926 F71
|--O. indistincta Tillyard 1926 F71
|--O. mitchelli Tillyard 1926 F71
|--O. obliqua Tillyard 1926 F71
|--O. pincombei Tillyard 1926 F71
|--O. quinquemedia Tillyard 1926 F71
|--O. subcostalis Tillyard 1926 F71
`--O. tetraneura Tillyard 1926 F71
Palaeovicia incerta Evans 1943 F71
Permagra distincta Evans 1943 F71
Permopsychops belmontensis Tillyard 1926 [incl. Permithone venosa Davis 1943] F71
Permoberothella perplexa Riek 1953 F71
Permobrachus F71
|--P. dubia (Tillyard 1926) [=Permodiphthera dubia] F71
`--P. magnus Evans 1943 F71
Permocapitus globulus Evans 1943 F71
Permocephalus knighti Evans 1943 F71
Permodiphthera robusta Tillyard 1926 F71
Permoglyphis belmontensis Tillyard 1926 F71
Permojassus F71
|--P. australis Tillyard 1926 F71
`--P. dubius Tillyard 1926 F71
Permopsyllidium F71
|--P. affine Tillyard 1926 F71
`--P. mitchelli Tillyard 1926 F71
Permopsyllidops stanleyi Davis 1942 F71
Permopsylloides insolita Evans 1943 F71
Permorapisma biserialis Tillyard 1926 [incl. P. triserialis Tillyard 1926] F71
Permoscarta F71
|--P. mitchelli Tillyard 1917 F71
`--P. triventulata Tillyard 1926 F71
Permosyne F71
|--P. affinis Tillyard 1924 F71
|--P. belmontensis Tillyard 1924 F71
|--P. mitchelli Tillyard 1924 F71
`--P. pincombeae Tillyard 1924 F71
Permothea latipennis Tillyard 1926 F71
Permotheela scytinopteroides Davis 1942 F71
Permovicia obscura Evans 1943 F71
Phipoides elegans Riek 1953 F71
Prohagla superba Riek 1954 F71
Protopincombea obgscura Evans 1943 F71
Protopsyllops minuta Evans 1943 F71
Psocopsyllidium media Davis 1942 F71
Psocoscytina bifida Davis 1942 F71
Psyllidella magna Evans 1943 F71
Psyllidiana davisia Evans 1943 F71
Stanleyana pulchra Evans 1943 F71
Stenoglyphis kimblensis Evans 1947 F71
Stenoscytina australiensis Tillyard 1926 F71
Triassagrion australiensis Tillyard 1922 F71
Triassodoecus chinai Evans 1963 F71
Triassophlebis stigmata Tillyard 1922 F71
Triassopsylla plecioides Tillyard 1917 F71
Triassocytinopsis paranotalis Evans 1956 F71
Tripsyllidium wadei Evans 1956 F71
Tychticoloides belmontensis Evans 1963 F71
Tenebroides mauritanicus BC91
Fumea casta BC91
Paleacrita vernata Z91
Itame argillacearia DC10
Biloba subsecivela VHK02
Diaphania indica [=Margaronia indica] VHK02
Lygropia quaternalis VHK02
Marasma suspicalis VHK02
Psara stultalis VHK02
Holcocera pulverea VHK02
Noorda moringa VHK02
Nephanteryx rhodobasalis VHK02
Anomalococcus indicus VHK02
Phidodonta modesta VHK02
Bissetia steniella VHK02
‘Cheyletus’ nigripes Mola 1907 B49
Stephanostoma Danielsen 1880 D56
Phrynomorphus Curtis 1831 FG06
*Type species of generic name indicated
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