Field of Science

Trochus

Spotted top shell Trochus maculatus, copyright Ria Tan.


Belongs within: Trochinae.

Trochus, top shells, is a genus of marine gastropods bearing a conical shell with a more or less flattened base, known from the Miocene to the present day (Knight et al. 1960).

<==Trochus Linnaeus 1758 BR05 [=Lamprostoma Swainson 1840 KC60, Polydonta Schumacher 1817 KC60]
    |--T. (Trochus) WG71
    |    |--*T. (T.) maculatus Linnaeus 1758 [=*Lamprostoma maculatum, *Polydonta maculata] KC60
    |    `--T. (T.) lineatus Lamarck 1822 WG71
    |--T. (Belangeria Fischer 1879) KC60
    |    `--T. (*B.) scabrosus Philippi 1850 KC60
    |--T. (Coelotrochus Fischer 1879) [incl. Neozelandia Cossmann 1918] KC60
    |    |--T. (*C.) tiaratus Quoy & Gaimard 1834 F27a
    |    |--T. (C.) avarus Suter 1917 [=T. (Anthora) avarus] F27a
    |    `--T. (C.) huttoni (Cossmann 1916) (see below for synonymy) F27b
    |--T. (Infundibulops Pilsbry 1889) KC60
    |    `--T. (*I.) erythraeus Brocchi 1821 KC60
    |--T. (Infundibulum Montfort 1810) [=Carinidea Swainson 1840] KC60
    |    `--T. (*I.) concavus Gmelin 1791 [=*Carinidea concava] KC60
    |--T. (Praecia Gray 1857) KC60
    |    `--T. (*P.) elegantulus Wood 1828 KC60
    |--T. (Scalaetrochus Etheridge 1890) F71, KC60
    |    `--T. (*S.) lindstromi Etheridge 1890 KC60, F71
    |--T. (Thorista Iredale 1915) [=Anthora Gray 1857 non Doubleday 1844] KC60
    |    |--T. (T.) viridis Gmelin 1791 KC60 (see below for synonymy)
    |    `--T. camelophorus Webster 1906 F27a
    `--T. (Thoristella Iredale 1915) KC60
         |--T. (*T.) chathamensis (Hutton 1873) [=Polydonta chathamensis] KC60
         |    |--T. c. chathamensis F27a
         |    |--‘Thoristella’ c. aucklandica (Smith 1902) F27a
         |    |--‘Thoristella’ c. benthicola Finlay 1927 F27a
         |    |--‘Thoristella’ c. dunedinensis (Suter 1917) F27a
         |    |--‘Thoristella’ c. fossilis Finlay 1927 F27a
         |    `--‘Thoristella’ c. oppressa (Hutton 1878) F27a
         `--‘Thoristella’ carmesina (Webster 1908) [=Monilea carmesina] F27a

Trochus incertae sedis:
  T. antipodum Wilckens 1922 F27a
  T. astraliformis Lindström 1884 LYR78
  T. calcaratus Souverbie 1875 H09
  T. circinatus Hutton 1873 F27a
  T. fenestratus Gmelin 1791 H09
  T. fenneonianus K54
  T. hexagonus Philippi 1846 H09
  T. histrio Reeve 1842 BW09
  T. mutus Finlay 1924 [=T. nodosus Hutton 1885 (preoc.)] F27b
  T. niloticus C60
  T. occidentalis Mighels & Adams 1842 J49
  T. triserialis (Lamarck 1822) BW09

Nomen nudum: Trochus quattuorcarinata Mighels & Adams 1841 J49

Trochus (Coelotrochus) huttoni (Cossmann 1916) [=*Neozelandia huttoni; incl. Anthora conica Hutton 1883, T. conicus (preoc.)] F27b

Trochus (Thorista) viridis Gmelin 1791 KC60 [incl. Polydonta tuberculata Gray 1843 KC60, *Anthora tuberculata KC60, Trochus (*Thorista) tuberculatus F27a]

*Type species of generic name indicated

REFERENCES

[BR05] Bouchet, P., J.-P. Rocroi, J. Frýda, B. Hausdorf, W. Ponder, Á. Valdés & A. Warén. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1-2): 1-397.

[BW09] Bryce, C. & C. Whisson. 2009. The macromolluscs of Mermaid (Rowley Shoals), Scott and Seringapatam Reefs, Western Australia. Records of the Western Australian Museum Supplement 77: 177-208.

[C60] Cox, L. R. 1960. Gastropoda: general characteristics of Gastropoda. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt I. Mollusca 1: Mollusca—General Features, Scaphopoda, Amphineura, Monoplacophora, Gastropoda—General Features, Archaeogastropoda and some (mainly Paleozoic) Caenogastropoda and Opisthobranchia, pp. I84-I169. Geological Society of America, and University of Kansas Press.

[F27a] Finlay, H. J. 1927a. A further commentary on New Zealand molluscan systematics. Transactions and Proceedings of the New Zealand Institute 57: 320-485.

[F27b] Finlay, H. J. 1927b. New specific names for austral Mollusca. Transactions and Proceedings of the New Zealand Institute 57: 488-533.

[F71] Fletcher, H. O. 1971. Catalogue of type specimens of fossils in the Australian Museum, Sydney. Australian Museum Memoir 13: 1-167.

[H09] Hedley, C. 1909. The Marine Fauna of Queensland: Address by the President of Section D. Australasian Association for the Advancement of Science: Brisbane.

[J49] Johnson, R. I. 1949. Jesse Wedgwood Mighels with a bibliography and a catalogue of his species. Occasional Papers on Mollusks 1 (14): 213-231.

[KC60] Knight, J. B., L. R. Cox, A. M. Keen, R. L. Batten, E. L. Yochelson & R. Robertson. 1960. Gastropoda: systematic descriptions. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt I. Mollusca 1: Mollusca—General Features, Scaphopoda, Amphineura, Monoplacophora, Gastropoda—General Features, Archaeogastropoda and some (mainly Paleozoic) Caenogastropoda and Opisthobranchia, pp. I169-I331. Geological Society of America, and University of Kansas Press.

[K54] Kurtén, B. 1954. The type collection of Ictitherium robustum (Gervais, ex Nordmann) and the radiation of the ictitheres. Acta Zoologica Fennica 86: 1-26.

[LYR78] Linsley, R. M., E. L. Yochelson & D. M. Rohr. 1978. A reinterpretation of the mode of life of some Paleozoic frilled gastropods. Lethaia 11: 105-112.

[WG71] Wilson, B. R., & K. Gillett. 1971. Australian Shells: illustrating and describing 600 species of marine gastropods found in Australian waters. A. H. & A. W. Reed: Sydney.

Trochinae

Strawberry top shell Tectus conus, copyright J. C. Schou.


Belongs within: Trochoidea.
Contains: Clanculus, Micrelenchus, Cantharidus, Trochus, Gibbula, Monodonta, Diloma, Thalotia.

The Trochinae are a group of marine gastropods with conical, turbiniform or littoriniform shells in which the aperture commonly has strongly discordant lips, with the outer lip prosocline (Knight et al. 1960, in which Trochinae as used here is divided between Trochinae and Monodontinae).

<==Trochinae [Trochidia, Trochininae]
    |  i. s.: Clanculus F27
    |         Micrelenchus F27
    |--Cantharidini [Cantharidinae, Canthiridina] BR05
    |    |--Cantharidus BR05
    |    `--Phasianotrochus Fischer 1885 KC60
    |         |--P. eximius (Perry 1811) WG71 (see below for synonymy)
    |         |--P. apicinus M54
    |         |--P. bellulus M54
    |         `--P. irisodontes M54
    |--Trochini BR05
    |    |--Trochus BR05
    |    `--Tectus Montfort 1810 WG71, F27 (see below for synonymy)
    |         |  i. s.: T. niloticus Linnaeus 1767 BW09
    |         |         T. pyramis (Born 1778) BW09 (see below for synonymy)
    |         |         T. triserialis (Lamarck 1822) BW09
    |         |--*T. (Tectus) mauritianus (Gmelin 1791) [=Trochus mauritianus] KC60
    |         |--T. (Cardinalia Gray 1847) KC60
    |         |    `--T. (*C.) virgatus (Gmelin 1791) [=Trochus virgatus] KC60
    |         `--T. (Rochia Gray 1857) KC60
    |              `--T. (R.) conus (Gmelin 1791) (see below for synonymy) KC60
    `--Monodontini [Gibbulinae, Monodontinae] BR05
         |--Gibbula BR05
         |--Monodonta BR05
         |--Diloma KC60
         |--Thalotia KC60
         |--Praelittorina Kutassy 1937 KC60
         |    `--*P. triadica Kutassy 1937 KC60
         |--Ozodochilus Cossmann 1918 KC60
         |    `--*O. subfilosus (Buvignier 1852) [=Trochus subfilosus] KC60
         |--Cochleochilus Cossmann 1918 KC60
         |    `--*C. cottaldinus (d’Orbigny 1853) [=Trochus cottaldinus] KC60
         |--Islipia Cox & Arkell 1950 KC60
         |    `--*I. lycetti (Lycett 1863) [=Monodonta lycetti] KC60
         |--Michaletia Cossmann 1904 KC60
         |    `--*M. semigranulata Cossmann 1904 KC60
         |--Chrysostoma Swainson 1840 KC60
         |    `--C. paradoxum (Born 1780) (see below for synonymy) KC60
         |--Jujubinus Monterosato 1884 [incl. Manotrochus Fischer 1885, Mirulinus Monterosato 1917] KC60
         |    `--J. exasperatus (Pennant 1777) (see below for synonymy) KC60
         |--Lesperonia Tournouër 1874 KC60
         |    `--*L. princeps Tournouër 1874 KC60
         |--Pachydontella Marwick 1948 KC60
         |    `--*P. etiampicta Marwick 1948 KC60
         |--Pictiformes Kolesnikov 1939 KC60
         |    `--*P. mamilla (Andrzejowski 1830) [=Monodonta mamilla] KC60
         |--Turcicula Dall 1881 KC60
         |    `--*T. imperialis (Dall 1881) [=Margarita imperialis] KC60
         |--Buckmanina Cossmann 1920 [=Brasilia Cossmann 1918 non Buckman 1898 nec Clarke 1913] KC60
         |    `--*B. erinus (d’Orbigny 1853) [=Turbo erinus] KC60
         |--Calliovarica Vokes 1939 KC60
         |    `--*C. eocensis Vokes 1939 KC60
         |--Cittarium Philippi 1847 [=Livona Gray 1847, Melagris Montfort 1810 non Linnaeus 1758] KC60
         |    `--*C. pica (Linné 1758) [=Turbo pica, *Livona pica, *Meleagris pica] KC60
         |--Houdasia Cossmann 1902 KC60
         |    `--*H. splendens Cossmann 1902 KC60
         |--Nanula Thiele 1924 KC60
         |    `--*N. tasmanica (Petterd 1879) KC60 [=Gibbula tasmanica KC60, Margarita tasmanica F27]
         |--Norrisia Bayle 1880 [=Trochiscus Sowerby 1838 non Heyden 1826] KC60
         |    `--*N. norrisi (Sowerby 1838) KC60 [=*Trochiscus norrisi KC60; incl. Turbo rotelliformis Jay 1839 BC01]
         |--Phorculus Cossmann 1888 KC60
         |    `--*P. fraterculus (Deshayes 1863) [=Turbo fraterculus] KC60
         |--Trochinella Iredale 1937 KC60
         |    `--*T. perconfusa Iredale 1937 KC60
         |--Bathybembix Crosse 1892 KC60, BC01 [=Bembix Watson 1879 non Fabricius 1775 KC60]
         |    |  i. s.: B. bairdii (Dall 1889) [=Turcicula bairdii] BC01
         |    |--*B. (Bathybembix) aeola (Watson 1879) [=*Bembix aeola] KC60
         |    `--B. (Ginebis Otuka 1942) KC60
         |         `--B. (*G.) argenteonitens (Lischke 1872) [=Trochus argenteonitens] KC60
         |--Turcica Adams & Adams 1854 KC60, BR05 [incl. Ptychostylis Gabb 1866 KC60; Turcicinae]
         |    |  i. s.: T. caffea Gabb 1865 O27
         |    |--*T. (Turcica) monilifera Adams 1854 KC60
         |    `--T. (Perrinia Adams & Adams 1854) KC60
         |         `--T. (*P.) angulifera (Adams 1853) [=Monodonta angulifera] KC60
         |--Clelandella Winckworth 1932 KC60
         |    |--C. azorica Gofas 2005 F05
         |    |--C. dautzenbergeri Gofas 2005 F05
         |    |--C. infuscata (Gould 1861) [=Ziziphinus infuscatus, Cantharidus infuscatus] BD86
         |    |--C. madeirensis Gofas 2005 F05
         |    `--C. perforata Gofas 2005 F05
         |--Austrocochlea Fischer 1885 [=Austrochlea Pilsbry 1889] KC60
         |    |--*A. constricta Lamarck 1822 F27, WG71 [=Monodonta (Austrocochlea) constricta F27]
         |    |--A. adelaidea M54
         |    |--A. concamerata Wood 1828 WG71
         |    |--A. rudis Gray 1826 WG71
         |    `--A. zebra F27
         `--Fossarina Adams & Angas 1864 [incl. Minos Hutton 1884] KC60
              |  i. s.: F. rimata (Hutton 1884) F27
              |--*F. (Fossarina) patula Adams & Angas 1864 KC60
              |--F. (Clydonochilus Fischer 1890) KC60
              |    `--F. (*C.) mariei (Fischer 1890) [=*Clydonochilus mariei] KC60
              `--F. (Minopa Iredale 1924) KC60
                   `--F. (*M.) legrandi Petterd 1879 KC60

Chrysostoma paradoxum (Born 1780) [=Helix paradoxa; incl. Turbo nicobaricus Chemnitz 1781 (non-binomial), *Chrysostoma nicobaricum] KC60

Jujubinus exasperatus (Pennant 1777) [=Trochus exasperatus; incl. T. matoni Payraudeau 1827, *Jujubinus matoni] KC60

Phasianotrochus eximius (Perry 1811) WG71 [=Bulimus eximius KC60, Cantharidus (Phasianotrochus) eximius KC60; incl. Trochus badius Wood 1828 KC60, *Phasianotrochus badius KC60]

Tectus Montfort 1810 WG71, F27 [incl. Pyramidea Swainson 1840 KC60, Pyramis Schumacher 1817 non Röding 1798 KC60; Pyramidinae]

Tectus pyramis (Born 1778) BW09 [=Trochus (Tectus) pyramis WG71; incl. Tr. obeliscus Gmelin 1788 WG71]

Tectus (Rochia) conus (Gmelin 1791) [=Trochus conus; incl. Tr. acutangulus (non-binomial), *Rochia acutangula] KC60

*Type species of generic name indicated

REFERENCES

[BD86] Barash, A., & Z. Danin. 1986. Further additions to the knowledge of Indo-Pacific Mollusca in the Mediterranean Sea (Lessepsian migrants). Spixiana 9 (2): 117-141.

[BR05] Bouchet, P., J.-P. Rocroi, J. Frýda, B. Hausdorf, W. Ponder, Á. Valdés & A. Warén. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1-2): 1-397.

[BC01] Boyko, C. B. & J. R. Cordeiro. 2001. Catalog of Recent type specimens in the Division of Invertebrate Zoology, American Museum of Natural History. V. Mollusca, part 2 (class Gastropoda [exclusive of Opisthobranchia and Pulmonata], with supplements to Gastropoda [Opisthobranchia], and Bivalvia). Bulletin of the American Museum of Natural History 262: 1-170.

[BW09] Bryce, C. & C. Whisson. 2009. The macromolluscs of Mermaid (Rowley Shoals), Scott and Seringapatam Reefs, Western Australia. Records of the Western Australian Museum Supplement 77: 177-208.

[F27] Finlay, H. J. 1927. A further commentary on New Zealand molluscan systematics. Transactions and Proceedings of the New Zealand Institute 57: 320-485.

[KC60] Knight, J. B., L. R. Cox, A. M. Keen, R. L. Batten, E. L. Yochelson & R. Robertson. 1960. Gastropoda: systematic descriptions. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt I. Mollusca 1: Mollusca—General Features, Scaphopoda, Amphineura, Monoplacophora, Gastropoda—General Features, Archaeogastropoda and some (mainly Paleozoic) Caenogastropoda and Opisthobranchia, pp. I169-I331. Geological Society of America, and University of Kansas Press.

[M54] Macpherson, J. H. 1954. Molluscs (sea shells and snails). In The Archipelago of the Recherche (J. M. Willis, ed.) pp. 55-63. Australian Geographical Society: Melbourne.

[O27] Oldroyd, I. S. 1927. The Marine Shells of the West Coast of North America vol. II, pt. III. Stanford University Press: Stanford University (California).

[WG71] Wilson, B. R., & K. Gillett. 1971. Australian Shells: illustrating and describing 600 species of marine gastropods found in Australian waters. A. H. & A. W. Reed: Sydney.

Trochoidea

Stomatia phymotis, copyright Femorale.


Belongs within: Vetigastropoda.
Contains: Calliostomatinae, Anomphalinae, Pycnomphalinae, Solariella, Margaritinae, Trochinae, Umboniinae, Stomatella.

The Trochoidea is a diverse group of vetigastropods lacking either a slit or selenizone (Bouchet et al. 2005).

Trochoidea [Microdomatacea, Microdomatoidea, Trochacea]
    |--Tychobrahea Horný 1992 [Tychobraheidae] BR05
    |--Eucochlis Knight 1933 [Eucochlidae] BR05
    |    `--*E. perminuta Knight 1933 KC60
    |--Velainella Vasseur 1880 BR05 [=Velainiella (l. c.) KC60; Velainellidae, Velainellinae, Vellainelloidea]
    |    `--*V. columnaris Vasseur 1880 KC60
    |--Calliostomatidae BR05
    |    |--Calliostomatinae BR05
    |    `--Thysanodonta Marshall 1988 [Thysanodontidae, Thysanodontinae] BR05
    |--Elasmonematidae BR05
    |    |--Elasmonema Fischer 1885 BR05 [=Callonema Hall 1879 non Conrad 1875 KC60]
    |    |    `--*E. bellatulum (Hall 1861) [=Loxonema bellatulum, *Callonema bellatulum] KC60
    |    |--Anematina Knight 1933 B79
    |    |    `--*A. proutana (Hall 1858) B79 [=Holopea proutana KC60]
    |    |--Streptotrochus Perner 1907 KC60
    |    |    `--*S. rugulosus Perner 1907 KC60
    |    `--Discordichilus Cossmann 1918 KC60
    |         `--*D. mollis (Lindström 1884) [=Trochus mollis] KC60
    |--Anomphalidae [Anomphalacea] P84
    |    |  i. s.: Cycloscena Fletcher 1958 W63
    |    |           |--*C. anomphala Fletcher 1958 W63
    |    |           |--C. costata Waterhouse 1963 W63
    |    |           |--C. glabra Waterhouse 1963 W63
    |    |           `--‘Pleurotomaria’ rotunda Etheridge 1872 [=Platyschisma rotunda] W63
    |    |         Eiselia Dietz 1911 P84
    |    |           `--*E. dyadica Dietz 1911 KC60
    |    |--Anomphalinae P84
    |    `--Pycnomphalinae P84
    |--Solariellidae [Solariellinae] BR05
    |    |--Solariella BR05
    |    |--Cidarina Dall 1909 KC60
    |    |    `--*C. cidaris (Carpenter 1864) KC60 [=Margarita cidaris KC60, Margarites cidaris O27]
    |    |--Zeminolia Finlay 1927 KC60
    |    |    |--*Z. plicatula (Murdoch & Suter 1906) [=Minolia plicatula] F27
    |    |    `--Z. semireticulata (Suter 1908) [=Minolia semireticulata, Spectamen semireticulatum] F27
    |    `--Minolia Adams 1860 H92 [=Minosia Dunker 1882 KC60; incl. Minolops Iredale 1929 KC60; Minoliinae]
    |         |--*M. punctata Adams 1860 H92
    |         |--M. casta (Nevill & Nevill 1874) H92
    |         |--M. gradata Sowerby 1895 H92
    |         |--M. holdsworthana Nevill & Nevill 1871 H92
    |         |--M. nedyma Melvill 1897 BD86
    |         |--M. singaporensis (Pilsbry 1889) [=Monilea (Minolia) singaporensis] BC01
    |         |--M. strigata Tenison-Woods 1878 F71
    |         `--M. vitiliginea (Menke 1843) H92
    |--Holopeidae [Holopeinae] PE02
    |    |--Raphispira Perner 1903 KC60
    |    |    `--*R. plena Perner 1903 KC60
    |    |--Cyclora Hall 1845 [Cycloridae] BR05
    |    |    `--*C. minuta Hall 1845 KC60
    |    |--Straparollina Billings 1865 [Straparollinidae, Straparollinoidea] KC60
    |    |    `--*S. pelagica Billings 1865 KC60
    |    |--Globonema Wenz 1938 KC60
    |    |    `--*G. kokeni (Wenz 1938) (see below for synonymy) KC60
    |    |--Protospirialis Clarke 1904 KC60
    |    |    `--*P. minutissima (Clarke 1885) [=Platyostoma minutissima] KC60
    |    `--Holopea Hall 1847 BR05 (see below for synonymy)
    |         |--*H. symmetrica Hall 1847 PE02 [=*Litiopsis symmetrica KC60]
    |         |--H. baltica YN83
    |         |--H. rotunda YN83
    |         |--H. ruoqiangensis Yu & Ning 1983 YN83
    |         |--H. teres Pan & Erwin 2002 PE02
    |         `--H. wellingtonensis Etheridge 1898 F71
    |--Proconulidae [Proconulinae] BR05
    |    |--Dimorphotectus Cossmann 1918 KC60
    |    |    `--D. fasciatus (Hoernes 1856) (see below for synonymy) KC60
    |    |--Callotrochus Kutassy in Wenz 1938 [=Mesotrochus Kutassy 1927 non Wasmann 1890] KC60
    |    |    `--*C. triadicus (Kutassy 1927) [=*Mesotrochus triadicus] KC60
    |    |--Eocalliostoma Haas 1953 KC60
    |    |    `--*E. interruptum (Cox 1947) [=Calliostoma interruptum] KC60
    |    |--Epulotrochus Cossmann 1918 KC60
    |    |    `--*E. epulus (d’Orbigny 1850) [=Trochus epulus] KC60
    |    |--Lithotrochus Conrad 1855 KC60
    |    |    `--L. humboldtii (von Buch 1839) (see below for synonymy) KC60
    |    |--Trypanotrochus Cossmann 1918 KC60
    |    |    `--*T. normanianus (d’Orbigny 1850) [=Trochus normanianus] KC60
    |    |--Proconulus Cossmann 1918 BR05
    |    |    |--*P. guillieri (Cossmann 1885) [=Trochus guillieri] KC60
    |    |    `--P. raulineus KC60
    |    |--Diplochilus Wöhrmann 1894 [=Raiblia Cossmann 1916, Woehrmannia Cossmann 1916] KC60
    |    |    |--*D. gracilis Wöhrmann 1894 [=*Raiblia gracilis, *Woehrmannia gracilis] KC60
    |    |    `--D. bistriatus KC60
    |    |--Tylotrochus Koken 1896 KC60
    |    |    |--*T. konincki (Hörnes 1856) [=Trochus konincki] TE01
    |    |    `--T. elongatus Bandel 1993 [=Ziziphinus semipunctatus Zardini 1978 (n. n.)] TE01
    |    |--Anticonulus Cossmann 1918 KC60
    |    |    |--*A. mariae (d’Orbigny 1853) [=Trochus mariae] KC60
    |    |    `--A. nisus KC60
    |    |--Muricotrochus Cossmann 1918 KC60
    |    |    |--*M. hudlestoni Cossmann 1918 KC60
    |    |    `--M. subluciensis KC60
    |    `--Discotectus Favre 1913 KC60
    |         |--*D. massalongoi [=Trochus massalongoi] KC60
    |         `--D. crassiplicatus KC60
    |--Microdomatidae BR05
    |    |  i. s.: Episfaxis Perner 1903 FB04 [=Cosmina Perner 1903 non Robineau-Desvoidy 1830 KC60]
    |    |           `--*E. complacens (Perner 1903) [=*Cosmina complacens] KC60
    |    |         Glyptospira Chronic 1952 FB04
    |    |           |--*G. cristulata Chronic 1952 PE02
    |    |           `--G. pinna Pan & Erwin 2002 PE02
    |    |         Daidia Wilson 1951 FB04
    |    |           `--*D. cerithioides (Salter 1859) [=Eunema cerithioides] KC60
    |    |         Pagodea Perner 1903 FB04
    |    |           `--*P. concomitans Perner 1903 KC60
    |    |         Dongiovannia Horný 1992 FB04
    |    |         Petrochus Horný 1992 FB04
    |    |         Threavia Lamont 1946 FB04
    |    |           `--*T. gulosa Lamont 1946 KC60
    |    |         Eopagodea Frýda & Rohr 2001 FB04
    |    |--Microdoma Meek & Worthen 1866 BR05 (see below for synonymy)
    |    |    |--*M. conicum Meek & Worthen 1867 FB04
    |    |    |--M. bicrenulata (De Koninck 1883) B79
    |    |    |--M. biserratum (Phillips 1836) B79
    |    |    |--M. nodosum Batten 1979 B79
    |    |    |--M. serrilimba (Phillips 1836) B79
    |    |    `--M. variegata Batten 1979 B79
    |    `--Decorospirinae BR05
    |         |--Dutrochus Blodgett 1993 FB04
    |         |--Roemeriella Blodgett & Frýda 1999 FB04
    |         `--Decorospira Blodgett & Johnson 1992 BR05
    |              |--*D. tasselli Blodgett & Johnson 1992 FB04
    |              |--D. gilberti Blodgett & Johnson 1992 FB04
    |              |--D. lepaini Frýda & Blodgett 2004 FB04
    |              |--D. minutula Frýda & Blodgett 2004 FB04
    |              |--D. pragensis (Horný 1992) FB04
    |              `--D. rigbyi Blodgett & Johnson 1992 FB04
    `--Trochidae [Trochinia] BR05
         |  i. s.: Cirsostylus Cossmann 1918 TE01, KC60
         |           `--*C. glandulus (Laube 1868) [=Trochus glandulus] TE01
         |         Kittlitrochus Cossmann 1909 (see below for synonymy) KC60
         |           `--*K. marginenodosus (Böhm 1895) (see below for synonymy) KC60
         |         Trochodon Seeley 1861 KC60
         |           `--*T. cancellatus (Seeley 1861) [=Trochus (*Trochodon) cancellatus] KC60
         |--Margaritinae H92
         |--Trochinae H92
         |--Umboniinae H92
         |--Halistylus Dall 1890 [Halistylinae] BR05
         |    |--*H. columna (Dall 1890) [=Cantharidus (*Halistylus) columna] KC60
         |    `--H. subpupoideus Tryon 1887 [=Fenella pupoidea (preoc.)] O27
         `--Stomatellinae [Macrostoma, Stomatellidae, Stomatidae, Stomatiidae, Stomatiinae] BR05
              |--Stomatella BW09
              |--Stylobates F27
              |--Phaneta F27
              |--Broderipia Gray 1847 KC60
              |    `--*B. rosea (Broderip 1834) [=Scutella rosea] KC60
              |--Mesogena Kutassy ex Cox in Knight, Cox et al. 1960 KC60
              |    `--*M. arcta (Hoernes 1855) [=Inoceramus arctus] KC60
              |--Praestomatia Kutassy ex Cox in Knight, Cox et al. 1060 KC60
              |    `--*P. acutangula (Koken 1897) [=Stomatia acutangula] KC60
              |--Roya Iredale 1912 KC60
              |    `--*R. kermadecensis Iredale 1912 KC60
              |--Synaptocochlea Pilsbry 1890 KC60
              |    |--S. stellata (Souverbie 1863) (see below for synonymy) KC60
              |    `--S. picta (d’Orbigny 1842) [incl. S. picta var. lactea Usticke 1959] BC01
              |--Pseudostomatella Thiele 1924 KC60
              |    |  i. s.: P. erythrocoma BC01
              |    |           |--P. e. erythrocoma BC01
              |    |           `--P. e. cycloradiata Usticke 1959 BC01
              |    |--*P. (Pseudostomatella) papyracea (Gmelin 1791) KC60 (see below for synonymy)
              |    `--P. (Stomatolina Iredale 1937) KC60
              |         `--P. (*S.) rufescens (Gray 1847) [=Stomatella rufescens] KC60
              `--Stomatia Helbling 1779 BR05 [=Stomax Montfort 1810 KC60; incl. Niphonia F27]
                   |  i. s.: S. angulata Adams 1850 H09
                   |         S. australis Adams 1850 H09
                   |         S. decussata Adams 1850 H09
                   |--*S. (Stomatia) phymotis Helbling 1779 [=*Stomax phymotis] KC60
                   |--S. (Microtis Adams & Angas 1850 non Br. 1810 [ICBN]) KC60
                   |    `--S. (*M.) tuberculata (Adams 1850) [=*Microtis tuberculata] KC60
                   `--S. (Miraconcha Bergh in Schepman 1908) KC60
                        `--S. (*M.) obscura (Bergh in Schepman 1908) [=*Miraconcha obscura] KC60

Dimorphotectus fasciatus (Hoernes 1856) [=Scoliostoma fasciatum; incl. Tectus hoernesi Koken 1896, *Dimorphotectus hoernesi] KC60

*Globonema kokeni (Wenz 1938) [=Trochonema (*Globonema) kokeni, Nematotrochus bicarinatus Koken 1925 non (Wahlenberg 1821)] KC60

Holopea Hall 1847 BR05 [=Litiopsis Salter 1866 KC60; incl. Anastrophina Knight 1937 KC60, Cirropsis Perner 1903 KC60, Haplospira Koken 1897 KC60, Staurospira Perner 1907 non Haeckel 1887 KC60, Tortilla Perner 1903 KC60]

Kittlitrochus Cossmann 1909 [=Paratrochus Kittl 1899 non Pilsbry 1893, Paratrochoides Tomlin 1929] KC60

*Kittlitrochus marginenodosus (Böhm 1895) [=Tectus marginenodosus, *Paratrochoides marginenodosus, *Paratrochus marginenodosus] KC60

Lithotrochus humboldtii (von Buch 1839) [=Pleurotomaria humboldtii; incl. Turritella andii d’Orbigny 1842, *Lithotrochus andii] KC60

Microdoma Meek & Worthen 1866 BR05 [=Microdomus Cossmann 1915 KC60; incl. Pleurotrochus Sherzer & Grabau 1909 KC60, Tuberculopleura Jakowlew 1899 KC60; Microdomatinae, Microdominae]

*Pseudostomatella (Pseudostomatella) papyracea (Gmelin 1791) KC60 [=Turbo papyraceus KC60, Stomatella papyracea KC60, Stomatia (*Pseudostomatella) papyracea F27]

Synaptocochlea stellata (Souverbie 1863) [=Stomatella stellata; incl. St. montrouzieri Pilsbry 1890, *Synaptocochlea montrouzieri] KC60

*Type species of generic name indicated

REFERENCES

[BD86] Barash, A., & Z. Danin. 1986. Further additions to the knowledge of Indo-Pacific Mollusca in the Mediterranean Sea (Lessepsian migrants). Spixiana 9 (2): 117-141.

[B79] Batten, R. L. 1979. Gastropods from Perak, Malaysia. Part 2. The trochids, patellids and neritids. American Museum Novitates 2685: 1-26.

[BR05] Bouchet, P., J.-P. Rocroi, J. Frýda, B. Hausdorf, W. Ponder, Á. Valdés & A. Warén. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1-2): 1-397.

[BC01] Boyko, C. B. & J. R. Cordeiro. 2001. Catalog of Recent type specimens in the Division of Invertebrate Zoology, American Museum of Natural History. V. Mollusca, part 2 (class Gastropoda [exclusive of Opisthobranchia and Pulmonata], with supplements to Gastropoda [Opisthobranchia], and Bivalvia). Bulletin of the American Museum of Natural History 262: 1-170.

[BW09] Bryce, C. & C. Whisson. 2009. The macromolluscs of Mermaid (Rowley Shoals), Scott and Seringapatam Reefs, Western Australia. Records of the Western Australian Museum Supplement 77: 177-208.

[F27] Finlay, H. J. 1927. A further commentary on New Zealand molluscan systematics. Transactions and Proceedings of the New Zealand Institute 57: 320-485.

[F71] Fletcher, H. O. 1971. Catalogue of type specimens of fossils in the Australian Museum, Sydney. Australian Museum Memoir 13: 1-167.

[FB04] Frýda, J., & R. B. Blodgett. 2004. New Emsian (Late Early Devonian) gastropods from Limestone Mountain, Medfra B-4 quadrangle, west-central Alaska (Farewell Terrane), and their paleobiogeographic affinities and evolutionary significance. Journal of Paleontology 78 (1): 111-132.

[H09] Hedley, C. 1909. The Marine Fauna of Queensland: Address by the President of Section D. Australasian Association for the Advancement of Science: Brisbane.

[H92] Herbert, D. G. 1992. Revision of the Umboniinae in southern Africa and Mozambique (Mollusca: Prosobranchia: Trochidae). Annals of the Natal Museum 33 (2): 379-459.

[KC60] Knight, J. B., L. R. Cox, A. M. Keen, R. L. Batten, E. L. Yochelson & R. Robertson. 1960. Gastropoda: systematic descriptions. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt I. Mollusca 1: Mollusca—General Features, Scaphopoda, Amphineura, Monoplacophora, Gastropoda—General Features, Archaeogastropoda and some (mainly Paleozoic) Caenogastropoda and Opisthobranchia, pp. I169-I331. Geological Society of America, and University of Kansas Press.

[O27] Oldroyd, I. S. 1927. The Marine Shells of the West Coast of North America vol. II, pt. III. Stanford University Press: Stanford University (California).

[PE02] Pan, H.-Z., & D. H. Erwin. 2002. Gastropods from the Permian of Guangxi and Yunnan Provinces, south China. Journal of Paleontology 76 (Memoir 56): 1-49.

[P84] Peel, J. S. 1984. Autecology and systematics of a new Silurian anomphalid gastropod from western North Greenland. Rapp. Grønlands geol. Unders. 121: 77-87.

[TE01] Tong, J., & D. H. Erwin. 2001. Triassic gastropods of the southern Qinling Mountains, China. Smithsonian Contributions to Paleobiology 92: 1-47.

[W63] Waterhouse, J. B. 1963. Permian gastropods of New Zealand. Part 4—Platyceratacea, Anomphalacea, Neritacea, and correlations. New Zealand Journal of Geology and Geophysics 6 (5): 817-842.

[YN83] Yu W. & Ning H. 1983. Middle Ordovician gastropods from Ruoqiang, Xinjiang. Acta Palaeontologica Sinica 22 (2): 199-203.

Natica

Natica canrena, copyright Laura Milano.


Belongs within: Naticidae.

Natica is a genus of predatory gastropods found in warm seas around the world, feeding on other molluscs.

Characters (from Oldroyd 1927): Shell oval globular, porcellanous, solid, generally smooth, covered by fine transparent epidermis that is generally not very persistent; umbilicated, or umbilicus more or less filled with callus; aperture semilunar, vertical, outer lip simple. Operculum large, semilunar, paucispiral, corneous or calcareous.

<==Natica Scopoli 1777 BR05
    |--*N. (Natica) canrena O27
    `--N. (Cryptonatica Dall 1892) O27
         |--N. (C.) affinis Gmelin 1792 O27
         |--N. (C.) aleutica Dall 1919 O27
         |--N. (C.) clausa Broderip & Sowerby 1829 O27
         |--‘Cryptonatica’ coosensis Finlay 1927 (see below for synonymy) F27b
         |--N. (C.) janthostoma Deshayes 1841 O27
         |--N. (C.) russa Gould 1859 O27
         `--N. (C.) salimba Dall 1919 O27

Natica incertae sedis:
  N. acinonyx Marche-Marchad 1957 BC01
  N. alapapilionis (Bolten 1798) [=Cochlis alapapilionis] H09
  N. alderi D56
  N. ambigua Morris & Lycett 1854 F27b
  N. arata Morris & Lycett 1854 F27b
  N. araucana [incl. N. australis d’Orbigny 1842 non Bosc 1801] F27b
  N. areolata Recluz 1844 H09
  N. australis Bosc 1801 F27b
  N. bacca A27
  N. bougei Sowerby 1908 BW09
  N. brunneolinea McLean 1970 BC01
  N. buriasensis Recluz 1844 H09
  N. callosa Cristofori & Jan 1832 F27b
  N. callosior F27b
  N. cayennensis Recluz 1850 [incl. N. hayesae Usticke 1959] BC01
  N. chinensis Lamarck 1822 H09
  N. colliei Recluz 1844 H09
  N. cunninghamensis Harris 1897 [=N. (Neverita) varians Tate 1893 (preoc.)] F27b
  N. deidosa Reeve 1855 H09
  N. denticulifera Marwick 1924 F27a
  N. dixoni Fernandes & Rolán 1992 BC01
  N. floridana C60
  N. groenlandica C60
  N. gualteriana Recluz 1844 H09
  N. guesti Harasewych & Jensen 1984 BC01
  N. harrisensis A27
  N. haydeni Cossmann 1899 [=N. ambigua Meek & Hayden 1855 non Morris & Lycett 1854] F27b
  N. helvacea Lamarck 1822 H09
  N. incerta Smith 1906 F27b
  N. japovata Finlay 1927 [=N. ovata Sowerby 1914 nec Klipstein 1843 nec Hutton 1873] F27b
  N. koatensis Krumbeck 1924 NE02
  N. laevis Weerth 1884 F27b
  N. ‘laevis’ Kaunhowen 1898 nec Weerth 1884 nec N. (Ampullina) laevis Hutton 1885 nec N. levis Smith 1896 F27b
  N. limpida Smith 1884 H09
  N. lineata (Bolten 1798) F27b [=Cochlis lineatus H09]
  N. livida BC01
    |--N. l. livida BC01
    `--N. l. oceani Usticke 1959 BC01
  N. maroccana F66
  N. migratoria (Powell 1927) HS01
  N. multipunctata [incl. N. multipunctata var. consors Wood 1848] F27b
  N. oolithica Auerbach 1871 NE02
  N. ovata Klipstein 1843 F27b
  N. percostata (Schauroth 1859) NE02
  N. phytelephas Reeve 1855 MG-H11
  N. planispira Phillips 1836 F27b
  N. polita Q72
  N. pusilla HC79
  N. raynoldiana Recluz 1844 H09
  N. robillardi Sowerby 1843 BW09
  N. ryalli Fernandes & Rolán 1992 BC01
  N. semicostata Lepsius 1878 NE02
  N. seychellium Watson 1886 H09
  N. sigillata McLean 1970 BC01
  N. solida Blainville 1825 WG71
  N. stellata (Martyn 1786) [=Nerita stellata] H09
  N. subcostata Ten.-Woods 1878 H09
  N. subinfundibulum [incl. N. subinfundibulum var. crassa Tate 1893] F27b
  N. suppleta Finlay 1927 [=N. incerta Harmer 1919 non Smith 1906] F27b
  N. variabilis Reeve 1855 F27b
  N. vixumbilicata Q72
  N. wintlei [incl. N. wintlei var. hamiltonensis Tenison-Woods 1878] F71

‘Cryptonatica’ coosensis Finlay 1927 [=Natica consors Dall 1909 non N. multipunctata var. consors Wood 1848] F27b

*Type species of generic name indicated

REFERENCES

[A27] Allan, R. S. 1927. The geology and palaeontology of the Lower Waihao Basin, south Canterbury, New Zealand. Transactions and Proceedings of the New Zealand Institute 57: 265-309.

[BR05] Bouchet, P., J.-P. Rocroi, J. Frýda, B. Hausdorf, W. Ponder, Á. Valdés & A. Warén. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1-2): 1-397.

[BC01] Boyko, C. B. & J. R. Cordeiro. 2001. Catalog of Recent type specimens in the Division of Invertebrate Zoology, American Museum of Natural History. V. Mollusca, part 2 (class Gastropoda [exclusive of Opisthobranchia and Pulmonata], with supplements to Gastropoda [Opisthobranchia], and Bivalvia). Bulletin of the American Museum of Natural History 262: 1-170.

[BW09] Bryce, C. & C. Whisson. 2009. The macromolluscs of Mermaid (Rowley Shoals), Scott and Seringapatam Reefs, Western Australia. Records of the Western Australian Museum Supplement 77: 177-208.

[C60] Cox, L. R. 1960. Gastropoda: general characteristics of Gastropoda. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt I. Mollusca 1: Mollusca—General Features, Scaphopoda, Amphineura, Monoplacophora, Gastropoda—General Features, Archaeogastropoda and some (mainly Paleozoic) Caenogastropoda and Opisthobranchia, pp. I84-I169. Geological Society of America, and University of Kansas Press.

[D56] Dawes, B. 1956. The Trematoda with special reference to British and other European forms. University Press: Cambridge.

[F27a] Finlay, H. J. 1927a. A further commentary on New Zealand molluscan systematics. Transactions and Proceedings of the New Zealand Institute 57: 320-485.

[F27b] Finlay, H. J. 1927b. New specific names for austral Mollusca. Transactions and Proceedings of the New Zealand Institute 57: 488-533.

[F71] Fletcher, H. O. 1971. Catalogue of type specimens of fossils in the Australian Museum, Sydney. Australian Museum Memoir 13: 1-167.

[F66] Fraser, L. 1866. Communication of a list of mollusks collected by R. Swinhoe, Esq., in Formosa. Proceedings of the Zoological Society of London 1866: 146.

[HS01] Hayward, B. W., A. B. Stephenson, M. S. Morley, W. M. Blom, H. R. Grenfell, F. J. Brook, J. L. Riley, F. Thompson & J. J. Hayward. 2001. Marine biota of Parengarenga Harbour, Northland, New Zealand. Records of the Auckland Museum 37: 45-80.

[H09] Hedley, C. 1909. The Marine Fauna of Queensland: Address by the President of Section D. Australasian Association for the Advancement of Science: Brisbane.

[HC79] Huff, J. A., & S. P. Cobb. 1979. Penaeoid and sergestoid shrimps (Crustacea: Decapoda). Memoirs of the Hourglass Cruises 5 (4): 1-101.

[MG-H11] McEnnulty, F. R., K. L. Gowlett-Holmes, A. Williams, F. Althaus, J. Fromont, G. C. B. Poore, T. D. O’Hara, L. Marsh, P. Kott, S. Slack-Smith, P. Alderslade & M. V. Kitahara. 2011. The deepwater megabenthic invertebrates on the western continental margin of Australia (100–1100 m depths): composition, distribution and novelty. Records of the Western Australian Museum Supplement 80: 1-191.

[NE02] Nützel, A. & D. H. Erwin. 2002. Battenizyga, a new Early Triassic gastropod genus with a discussion of the caenogastropod evolution at the Permian/Triassic boundary. Paläontologische Zeitschrift 76 (1): 21-27.

[O27] Oldroyd, I. S. 1927. The Marine Shells of the West Coast of North America vol. II, pt. III. Stanford University Press: Stanford University (California).

[Q72] Quilty, P. G. 1972. The biostratigraphy of the Tasmanian marine Tertiary. Papers and Proceedings of the Royal Society of Tasmania 106: 25-44.

[WG71] Wilson, B. R., & K. Gillett. 1971. Australian Shells: illustrating and describing 600 species of marine gastropods found in Australian waters. A. H. & A. W. Reed: Sydney.

Campaniloidea

Globularia fluctuata, from here.


Belongs within: Sorbeoconcha.

The Campaniloidea are a group of marine gastropods united by sperm morphology, with members having a euspermatozoon containing seven or eight stright mitochondria, accompanied by a dense segmented sheath. They also share a simple larval shell morphology (Ponder et al. 2008). The Campaniloidea are largely relictual in the modern fauna, with Campanilidae and Ampullinidae each being represented by only a single surviving species, together with the living genus Plesiotrochus.

<==Campaniloidea
    |--Trypanaxis Cossmann 1889 [Trypanaxidae, Trypanaxinae] BR05
    |--Plesiotrochus Fischer 1878 [Plesiotrochidae] BR05
    |    |--‘Tectarius’ luteus Gould 1861 R89
    |    `--P. pagodiformis Hedley 1907 H09
    |--Campanilidae BR05
    |    |--Gymnocerithium Cossmann 1906 [Gymnocerithiidae] BR05
    |    |    `--*G. collegiale (Zittel 1873) [=Cerithium collegiale] S98
    |    |--Campanile Bayle in Fischer 1884 [Campanilinae] BR05
    |    |    `--C. symbolicum Iredale 1917 WG71
    |    `--Diozoptyxis Cossmann 1896 [Diozoptyxidae, Diozoptyxinae, Diozoptyxisinae] BR05
    |         `--*D. monilifera [=Nerinea monilifera] BR05
    `--Ampullinidae [Ampullinoidea, Ampullospiridae] BR05
         |--Amaurellina Bayle in Fischer 1885 [Amaurellinidae] BR05
         |--Ampullospira Harris 1897 [Ampullospirinae] BR05
         |--Gyrodes Conrad 1860 [Gyrodeidae, Gyrodesacea, Gyrodinae] BR05
         |--Ampullina Bowdich 1822 [Ampullininae] BR05
         |    `--A. apora (Watson 1881) F27
         |--Globularia Swainson 1840 [Globulariidae, Globulariinae] BR05
         |    `--G. fluctuata (Sowerby 1825) [incl. Natica imperforata Jay 1836] BC01
         `--Pseudamaura Fischer 1885 [Pseudamauridae, Pseudamaurinidae] BR05
              |--P. globosa (Ahlburg 1906) NE02
              |--P. gregaria (v. Schlotheim 1822) NE02
              |--P. reflecta Etheridge 1902 F71
              `--P. variablis F71

*Type species of generic name indicated

REFERENCES

[BR05] Bouchet, P., J.-P. Rocroi, J. Frýda, B. Hausdorf, W. Ponder, Á. Valdés & A. Warén. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1-2): 1-397.

[BC01] Boyko, C. B. & J. R. Cordeiro. 2001. Catalog of Recent type specimens in the Division of Invertebrate Zoology, American Museum of Natural History. V. Mollusca, part 2 (class Gastropoda [exclusive of Opisthobranchia and Pulmonata], with supplements to Gastropoda [Opisthobranchia], and Bivalvia). Bulletin of the American Museum of Natural History 262: 1-170.

[F27] Finlay, H. J. 1927. A further commentary on New Zealand molluscan systematics. Transactions and Proceedings of the New Zealand Institute 57: 320-485.

[F71] Fletcher, H. O. 1971. Catalogue of type specimens of fossils in the Australian Museum, Sydney. Australian Museum Memoir 13: 1-167.

[H09] Hedley, C. 1909. The Marine Fauna of Queensland: Address by the President of Section D. Australasian Association for the Advancement of Science: Brisbane.

[NE02] Nützel, A. & D. H. Erwin. 2002. Battenizyga, a new Early Triassic gastropod genus with a discussion of the caenogastropod evolution at the Permian/Triassic boundary. Paläontologische Zeitschrift 76 (1): 21-27.

Ponder, W. F., D. J. Colgan, J. M. Healy, A. Nützel, L. R. L. Simone & E. E. Strong (2008). Caenogastropoda. In: Ponder, W. F., & D. L. Lindberg (eds) Phylogeny and Evolution of the Mollusca, pp. 331-383. University of California Press: Berkeley.

[R89] Reid, D. G. 1989. Systematic revision of the Recent species of Peasiella Nevill, 1885 (Gastropoda: Littorinidae), with notes on the fossil species. Nautilus 103 (2): 43-69.

[S98] Stilwell, J. D. 1998. Late Cretaceous Mollusca from the Chatham Islands, New Zealand. Alcheringa 22 (1): 29-85.

[WG71] Wilson, B. R., & K. Gillett. 1971. Australian Shells: illustrating and describing 600 species of marine gastropods found in Australian waters. A. H. & A. W. Reed: Sydney.

Turritellidae

Tachyrhynchus reticulatus, copyright Anders Warén.


Belongs within: Cerithioidea.
Contains: Turritella.

The Turritellidae are a group of high-spired marine gastropods. Fossils have been assigned to the Turritellidae going back as far as the Lower Devonian (Knight et al. 1960).

Characters (from Knight et al. 1960): High-spired, with 8 to 20 or more whorls, spiral ornament commonly conspicuous; shallow labral sinus usually culminating at or above mid-whorl with upper limb extending farther adaperturally than lower; siphonal notch or canal wanting.

<==Turritellidae [Turritellacea, Turritellae, Turritellea]
    |--Orectospira Dall 1925 [Orectospiridae, Orectospirinae] BR05
    |--Pareora Marwick 1931 [Pareoridae, Pareorinae] BR05
    |--Protoma Baird 1870 [Protominae] BR05
    |--Vermiculariinae [Vermiculariidae] BR05
    |    |--‘Pseudomesalia’ Douvillé 1916 non Ganglbauer 1900 [Pseudomesaliidae] BR05
    |    `--Vermicularia Lamarck 1799 BR05
    |         |--V. eburnea Reeve 1842 O27
    |         |--V. hedleyi Finlay 1927 [=V. nodosa Hedley 1907 non Kaunhowen 1897] F27b
    |         |--V. knorri (Deshayes 1843) HC79
    |         |--V. maxima (Sowerby 1825) [=Serpula maxima] H09
    |         |--V. novaehollandiae H09
    |         |--V. ophioides Marshall & Murdoch 1921 F27a
    |         |--‘Siphonium’ planatum Suter 1913 F27a
    |         `--V. quoyi (Adams & Adams 1854) [=Cladopoda quoyi] H09
    `--Turritellinae BR05
         |--Turritella BR05
         |--Zaria Gray 1842 [Zariinae] BR05
         |--Archimediella Sacco 1895 [Archimediellidae, Archimedielloidea] BR05
         |    `--A. dirkhartogensis Garrard 1972 MG-H11
         |--Zeacolpus Finlay 1926 [Zeacolpini] BR05
         |    |--*Z. vittatus (Hutton 1873) [=Turritella vittata; incl. T. carlottae Watson 1881] F27a
         |    |--Z. abscisus [=Turritella abscisa] F27a
         |    |--Z. ahiparana (Powell 1927) [=Turritella (Zeacolpus) ahiparana] BC01
         |    |--Z. albolapis (Finlay 1924) F27a [=Turritella albolapis F27a, T. concava Hutton 1877 (preoc.) F27b]
         |    |--Z. bartoni Darragh & Kendrick 2008 DK08
         |    |--Z. fulminatus [=Turritella fulminata] F27a
         |    |--Z. lornensis (Marwick 1926) [=Turritella lornensis] F27a
         |    |--Z. pagoda [=Turritella pagoda] F27a
         |    `--Z. semiconcavus [=Turritella semiconcava] F27a
         `--Tachyrhynchus Mörch 1868 [Tachyrhynchiinae, Tachyrhynchinae] BR05
              |--*T. acicula [=Turritella acicula] O27
              |--T. erosus Couthouy 1838 O27
              |    |--T. e. erosus O27
              |    `--T. e. major Dall 1919 O27
              |--T. lacteolus (Carpenter 1865) [=Masalia lacteola] O27
              |    |--T. l. lacteolus O27
              |    `--T. l. subplanatus Carpenter 1865 [=Mesalia lacteola subplanata] O27
              |--T. pratomus Dall 1919 O27
              |--T. reticulatus Mighels 1842 O27
              `--T. stearnsii Dall 1919 O27

Turritellidae incertae sedis:
  Glyptozaria Iredale 1924 F27a
    `--*G. opulenta [=Turritella opulenta] F27a
  Platycolpus Donald 1900 F27a
    |--*P. quadratus [=Turritella quadrata, Colpospira (*Platycolpus) quadrata] F27a
    `--P. guillaumei [=Colpospira guillaumei] F27a
  Maoricolpus Finlay 1927 F27a
    |--*M. roseus F27a (see below for synonymy)
    |--M. cavershamensis (Harris 1897) F27a (see below for synonymy)
    `--M. murrayanus [=Turritella murrayana] F27a
  Colpospira Donald 1900 F27a
    |--*C. runcinata [=Turritella runcinata; incl. T. accisa] F27a
    |--C. cordismei [=Turritella cordismei] F27a
    |--C. platyspira (Tenison-Woods 1878) F27a, F71 [=Turritella platyspira F27a]
    `--C. sinuata [=Turritella sinuata] F27a
  Gazameda Iredale 1924 F27a
    |--*G. gunnii [=Turritella gunnii] F27a
    |--G. acricula [=Turritella acricula] F27a
    |--G. iredalei Finlay 1927 [=Turritella clathrata Kiener 1843 non Deshayes 1833] F27b
    `--G. tasmanica [=Turritella tasmanica; incl. T. subsquamosa] F27a
  Ctenocolpus Iredale 1925 F27a
    |--*C. australis [=Turritella australis] F27a
    |    |--C. a. australis F27a
    |    `--C. a. diffidens Iredale 1925 F27a
    |--C. pagodula [=Turritella pagodula] F27a
    `--C. warburtoni [=Turritella warburtoni] F27a
  Spirocolpus Finlay 1927 F27a
    |--*S. waihaoensis (Marwick 1924) [=Turritella waihaoensis] F27a
    |--S. aldingae (Tate 1882) DK08 [=Turritella aldingae F27a]
    |--S. conspicabilis (Tate 1924) [=Turritella conspicabilis] F27a
    |--S. rudis (Marshall 1919) [=Turritella rudis] F27a
    |--S. tophina (Marwick 1926) [=Turritella tophina] F27a
    `--S. tristira (Tate 1924) [=Turritella tristira] F27a
  Stiracolpus Finlay 1927 F27a
    |--*S. symmetricus (Hutton 1873) [=Turritella symmetrica; incl. T. chordata] F27a
    |--S. atkinsoni [=Turritella atkinsoni] F27a
    |--S. godeffroyanus [=Turritella godeffroyana] F27a
    |--S. kanieriensis (Harris 1897) F27a (see below for synonymy)
    |--S. smithianus [=Turritella smithiana] F27a
    `--S. waikopiroensis (Suter 1917) [=Turritella waikopiroensis] F27a
  Callispira Nelson 1947 KC60
    `--*C. quinquecostata Nelson 1947 KC60

Maoricolpus cavershamensis (Harris 1897) F27a [=Turritella cavershamensis F27a, T. gigantea Hutton 1873 (preoc.) F27b]

*Maoricolpus roseus F27a [=Turritella rosea F27a; incl. T. difficilis Suter 1908 nec d’Orbigny 1842 nec Zekeli 1852 F27b]

Stiracolpus kanieriensis (Harris 1897) F27a [=Turritella kanieriensis F27a, T. (Zaria) tricincta Hutton 1873 nec Borson 1820 nec Morris 1845 F27b]

*Type species of generic name indicated

REFERENCES

[BR05] Bouchet, P., J.-P. Rocroi, J. Frýda, B. Hausdorf, W. Ponder, Á. Valdés & A. Warén. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1-2): 1-397.

[BC01] Boyko, C. B. & J. R. Cordeiro. 2001. Catalog of Recent type specimens in the Division of Invertebrate Zoology, American Museum of Natural History. V. Mollusca, part 2 (class Gastropoda [exclusive of Opisthobranchia and Pulmonata], with supplements to Gastropoda [Opisthobranchia], and Bivalvia). Bulletin of the American Museum of Natural History 262: 1-170.

[DK08] Darragh, T. A., & G. W. Kendrick. 2008. Silicified Eocene molluscs from the Lower Murchison district, Southern Carnarvon Basin, Western Australia. Records of the Western Australian Museum 24 (3): 217-246.

[F27a] Finlay, H. J. 1927a. A further commentary on New Zealand molluscan systematics. Transactions and Proceedings of the New Zealand Institute 57: 320-485.

[F27b] Finlay, H. J. 1927b. New specific names for austral Mollusca. Transactions and Proceedings of the New Zealand Institute 57: 488-533.

[F71] Fletcher, H. O. 1971. Catalogue of type specimens of fossils in the Australian Museum, Sydney. Australian Museum Memoir 13: 1-167.

[H09] Hedley, C. 1909. The Marine Fauna of Queensland: Address by the President of Section D. Australasian Association for the Advancement of Science: Brisbane.

[HC79] Huff, J. A., & S. P. Cobb. 1979. Penaeoid and sergestoid shrimps (Crustacea: Decapoda). Memoirs of the Hourglass Cruises 5 (4): 1-101.

[KC60] Knight, J. B., L. R. Cox, A. M. Keen, R. L. Batten, E. L. Yochelson & R. Robertson. 1960. Gastropoda: systematic descriptions. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt I. Mollusca 1: Mollusca—General Features, Scaphopoda, Amphineura, Monoplacophora, Gastropoda—General Features, Archaeogastropoda and some (mainly Paleozoic) Caenogastropoda and Opisthobranchia, pp. I169-I331. Geological Society of America, and University of Kansas Press.

[MG-H11] McEnnulty, F. R., K. L. Gowlett-Holmes, A. Williams, F. Althaus, J. Fromont, G. C. B. Poore, T. D. O’Hara, L. Marsh, P. Kott, S. Slack-Smith, P. Alderslade & M. V. Kitahara. 2011. The deepwater megabenthic invertebrates on the western continental margin of Australia (100–1100 m depths): composition, distribution and novelty. Records of the Western Australian Museum Supplement 80: 1-191.

[O27] Oldroyd, I. S. 1927. The Marine Shells of the West Coast of North America vol. II, pt. III. Stanford University Press: Stanford University (California).

Liotia

Liotia admirabilis, from Smith 1890.


Belongs within: Turbinoidea.

Liotia is a genus of marine gastropods with a latticed ornamentation, known from the Indo-Pacific and western Atlantic.

Characters (from Knight et al. 1960): Axial and spiral ribs well developed, forming latticed surface; umbilicus funnel-shaped, bounded by beaded cord; operculum concave, of many narrow whorls.

Liotia Gray 1842 BR05
    |--*L. cancellata (Gray 1828) [=Delphinula cancellata] KC60
    |--L. acidalia (Melvill & Standen 1899) [=Microtheca acidalia] H09
    |--L. acuticostata Carpenter 1864 O27
    |    |--L. a. acuticostata O27
    |    `--L. a. radiata Dall 1919 O27
    |--L. admirabilis Smith 1890 MASC88
    |--L. calliglypta Melvill 1891 H09
    |--L. corona Hedley 1902 H09
    |--L. devexa Hedley 1901 H09
    |--L. discoidea (Reeve 1843) [=Delphinula discoidea] H09
    |--L. fenestrata Carpenter 1864 O27
    |--L. incidata Hedley 1902 H09
    |--L. lamellosa Tenison-Woods 1878 F71
    |--L. latebrosa Hedley 1907 H09
    |--L. micans (Adams 1850) [=Cyclostrema micans] H09
    |--L. muricata (Reeve 1843) [=Delphinula muricata] H09
    |--L. philtata Hedley 1900 H09
    |--L. rostrata Hedley 1900 H09
    |--L. scalaroides (Reeve 1843) [=Delphinula scalaroides] H09
    |--L. varicosa (Reeve 1843) [=Delphinula varicosa] H09
    `--L. venusta Hedley 1901 H09

*Type species of generic name indicated

REFERENCES

[BR05] Bouchet, P., J.-P. Rocroi, J. Frýda, B. Hausdorf, W. Ponder, Á. Valdés & A. Warén. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1-2): 1-397.

[F71] Fletcher, H. O. 1971. Catalogue of type specimens of fossils in the Australian Museum, Sydney. Australian Museum Memoir 13: 1-167.

[H09] Hedley, C. 1909. The Marine Fauna of Queensland: Address by the President of Section D. Australasian Association for the Advancement of Science: Brisbane.

[KC60] Knight, J. B., L. R. Cox, A. M. Keen, R. L. Batten, E. L. Yochelson & R. Robertson. 1960. Gastropoda: systematic descriptions. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt I. Mollusca 1: Mollusca—General Features, Scaphopoda, Amphineura, Monoplacophora, Gastropoda—General Features, Archaeogastropoda and some (mainly Paleozoic) Caenogastropoda and Opisthobranchia, pp. I169-I331. Geological Society of America, and University of Kansas Press.

[MASC88] McLean, J. H., R. S. Absalão & R. L. dos Santos Cruz. 1988. A new species of Macrarene (Turbinidae: Liotiinae) from Brazil. Nautilus 102 (3): 99-101.

[O27] Oldroyd, I. S. 1927. The Marine Shells of the West Coast of North America vol. II, pt. III. Stanford University Press: Stanford University (California).