Pyrgotidae

Sphecomyiella valida, copyright Carl Barrentine.


Belongs within: Tephritoidea.

The Pyrgotidae are a group of flies whose larvae are endoparasites of adult scarabaeid beetles (Hernández-Ortiz 2010).

Characters (from Hernández-Ortiz 2010): Medium to large-sized flies (body length 5–30 mm), body usually robust with long legs, wing pattern often spotted, banded or reticulated, rarely hyaline. Head spherical with frons large, antennal bases projecting anteriorly, pedicel elongate; arista setiform; ocelli often absent; frons weakly depressed, frontal bristles and vibrissa absent, face broad with deep antennal grooves; mouthparts of moderate size. Scutum mostly bare, setulose, or distinctly haired, sometimes black spotted, major bristles relatively short; scutellum short and broad with one to four pairs of marginal bristles. Legs long in both sexes, frequently without strong bristles, but covered in short setulae; hind tibia tapering to base or with strong constriction in basal third or near midlength. Wing with or without subcostal break, C weakening after vein R4+5; Sc complete to C or disappearing close to C; R1 dorsally setulose; R2+3 sometimes with stump vein at apical third; cell cup closed by CuA2 usually forming acute angle in lower apical corner, sometimes short. Preabdomen elongate, robust, more or less petiolate; lateral membrane broad; syntergite 1+2 elongate, usually with several long hairs on lateral sides near base. Female terminalia with syntergosternite 7 robust and strong basally forming conical tube turned ventrally, sometimes longer than rest of abdominal length. Apical margin of syntergosternite 7 sometimes provided with ventral hook-like sclerite or two apicolateral claw-like sclerites; aculeus usually shorter than length of syntergosternite 7. Male terminalia with epandrium globose, curved ventrally; lateral surstyli usually short and robust basally, medial surstyli without prensisetae developed; aedeagus coiled and elongated; glans more or less developed, sometimes partially sclerotised.

<==Pyrgotidae
    |--Teretrurinae [Teretrurini] H-O10
    |    |--Pyrgotosoma H-O10
    |    `--Teretrura H-O10
    `--Pyrgotinae H-O10
         |--Descoleia [Toxurinae, Toxurini] H-O10
         |    `--D. teretrura H-O10
         `--Pyrgotini H-O10
              |--Anapyrgota personta H-O10
              |--Idiopyrgota setiventris H-O10
              |--Carrerapyrgota H-O10
              |--Pyrgotella chagnoni S87
              |--Boreothrinax H-O10
              |    |--B. dichaetus H-O10
              |    `--B. shewelli S87
              |--Neopyrgota H-O10
              |    |--N. major H-O10
              |    `--N. picea H-O10
              |--Stenopyrgota H-O10
              |    |--S. crassitibia H-O10
              |    `--S. mexicana H-O10
              |--Pyrgota H-O10
              |    |--P. longipes H-O10
              |    |--P. lugens H-O10
              |    `--P. undata H-O10
              |--Leptopyrgota H-O10
              |    |--L. albitarsis H-O10
              |    |--L. amplipennis H-O10
              |    `--L. undulata H-O10
              `--Sphecomyiella [incl. Stirothrinax] H-O10
                   |--S. cribrata [=Stirothrinax cribratus] H-O10
                   |--S. knudseni [=Stirothrinax knudseni] H-O10
                   `--S. valida H-O10

Pyrgotidae incertae sedis:
  Cardiacera miliacea WT11
  Maenomenus ensifer H-O10
  Peltodasia flaviseta H-O10
  Lopadops H-O10
  Tropidothrinax H-O10
  Toxopyrgota K10
  Adapsilia O98
    |--A. aequalis O98
    `--A. illingworthana CM91
  Eumorphomyia CM74
  Campylocera robusta O98
  Tephritopyrgota O98
  Apyrgota O98
  Plectrobrachis O98
  Taeniomastix O98
  Trichopeltia O98
  Epice setifemur O98
  Euphya O98

*Type species of generic name indicated

REFERENCES

[CM74] Colless, D. H., & D. K. McAlpine. 1974. Diptera (flies). In: CSIRO. The Insects of Australia: A textbook for students and research workers. Supplement 1974 pp. 91–96. Melbourne University Press.

[CM91] Colless, D. H., & D. K. McAlpine. 1991. Diptera (flies). In: CSIRO. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 2 pp. 717–786. Melbourne University Press: Carlton (Victoria).

[H-O10] Hernández-Ortiz, V. 2010. Pyrgotidae (pyrgotid flies). In: Brown, B. V., A. Borkent, J. M. Cumming, D. M. Wood, N. E. Woodley & M. A. Zumbado (eds) Manual of Central American Diptera vol. 2 pp. 955–961. NRC Research Press: Ottawa.

[O98] Oosterbroek, P. 1998. The Families of Diptera of the Malay Archipelago. Brill: Leiden.

[S87] Steyskal, G. C. 1987. Pyrgotidae. In: McAlpine, J. F. (ed.) Manual of Nearctic Diptera vol. 2 pp. 813–816. Research Branch, Agriculture Canada.

[WT11] Wiegmann, B. M., M. D. Trautwein, I. S. Winkler, N. B. Barr, J.-W. Kim, C. Lambkin, M. A. Bertone, B. K. Cassel, K. M. Bayless, A. M. Heimberg, B. M. Wheeler, K. J. Peterson, T. Pape, B. J. Sinclair, J. H. Skevington, V. Blagoderov, J. Caravas, S. N. Kutty, U. Schmidt-Ott, G. E. Kampmeier, F. C. Thompson, D. A. Grimaldi, A. T. Beckenbach, G. W. Courtney, M. Friedrich, R. Meier & D. K. Yeates. 2011. Episodic radiations in the fly tree of life. Proceedings of the National Academy of Sciences of the USA 108 (14): 5690–5695.

Bactrocera

Oriental fruit fly Bactrocera dorsalis, from the United States Department of Agriculture.


Belongs within: Tephritidae.

Bactrocera is a widespread genus of fruit flies, a number of species of which are significant horticultural pests. Members have wings with a broad cell bm, about twice as wide as cell cup, and the first antennal flagellomere is long, usually reaching the ventral margin of the face.

<==Bactrocera
    |--B. atrisetosa MHG04
    |--B. carambolae MHG04
    |--B. correcta MHG04
    |--B. cucurbitae (Coquillett 1899) [=Dacus cucurbitae, Zeugodacus cucurbitae] DK08
    |--B. decipiens MHG04
    |--B. dorsalis KP10
    |--B. kirki MHG04
    |--B. occipitalis MHG04
    |--B. oleae KP10
    |--B. papayae MHG04
    |--B. passiflorae MHG04
    |--B. philippinensis MHG04
    |--B. tau MHG04
    |--B. trilineola MHG04
    |--B. trivialis MHG04
    |--B. tryoni CM91 [=Dacus (Bactrocera) tryoni CM91, D. (Strumeta) tryoni CM70]
    |--B. umbrosa MHG04
    |--B. xanthodes MHG04
    `--B. zonata MHG04

*Type species of generic name indicated

REFERENCES

[CM70] Colless, D. H., & D. K. McAlpine. 1970. Diptera (flies). In: CSIRO. The Insects of Australia: A textbook for students and research workers pp. 656–740. Melbourne University Press.

[CM91] Colless, D. H., & D. K. McAlpine. 1991. Diptera (flies). In: CSIRO. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 2 pp. 717–786. Melbourne University Press: Carlton (Victoria).

[DK08] Dohm, P., D. Kovač, A. Freidberg & R. Bin Hashim. 2008. Biology of the Oriental bamboo-inhabiting fly Felderimyia gombakensis and observations on mating trophallaxis in Felderimyia (Insecta, Diptera, Tephritidae, Phytalmiinae, Acanthonevrini). Senckenbergiana Biologica 88 (2): 311–318.

[KP10] Kutty, S. N., T. Pape, B. M. Wiegmann & R. Meier. 2010. Molecular phylogeny of the Calyptratae (Diptera: Cyclorrhapha) with an emphasis on the superfamily Oestroidea and the position of Mystacinobiidae and McAlpine's fly. Systematic Entomology 35: 614–635.

[MHG04] Maynard, G. V., J. G. Hamilton & J. F. Grimshaw. 2004. Quarantine—phytosanitary, sanitary and incursion management: an Australian entomological perspective. Australian Journal of Entomology 43 (3): 318–328.

Psychodinae

Psychoda alternata, copyright Ivan Stankovic.


Belongs within: Psychodoidea.

The Psychodinae, moth flies, are a group of densely hairy flies whose larvae are most commonly found in moist or semi-aquatic habitats (Wagner & Ibáñez-Bernal 2009).

Characters (from Wagner & Ibáñez-Bernal 2009): Eye reniform with extension towards midline forming eye bridge above antennal insertions; eyes continuous or separated, rarely with eye bridge short and acutely pointed; flagellomeres fusiform, pyriform or nodiform.

<==Psychodinae WI-B09
    |--Maruina [Maruinini] WI-B09
    |    `--M. hirta WM96
    |--Pericomini WI-B09
    |    |--Lepidiella WI-B09
    |    `--Pericoma (Clytocerus) americana WI-B09, QV81
    |--Paramormiini WI-B09
    |    |--Clogmia albipunctata WI-B09, WT11
    |    |--Telmatoscopus mergacolis WI-B09
    |    `--Paramormia [incl. Duckhousiella] WI-B09
    |         |--P. furcata WI-B09
    |         `--P. ustulata WI-B09
    |--Mormiini WI-B09
    |    |--Atrichobrunettia alternata WI-B09, CM91
    |    |--Bryopharsos WI-B09
    |    |--Eugenys clavellata WI-B09
    |    |--Mormia O98
    |    `--Brunettia [incl. Neobrunettia Bravo & Amorim 1995] WI-B09
    |         `--B. dominicana WI-B09
    |--Psychodini WI-B09
    |    |--Trichopsychoda insulicola WI-B09, QV81
    |    |--Threticus WI-B09
    |    |--Feuerborniella WI-B09
    |    |--Philosepedon WI-B09
    |    |    |--P. helicis WM96
    |    |    |--P. mauroae Wagner & Masteller 1996 WM96
    |    |    |--P. opposita WM96
    |    |    `--P. quatei WM96
    |    `--Psychoda WI-B09
    |         |--P. alternata CM91
    |         |--P. cinerea WI-B09
    |         |--P. harrisi M83
    |         |--P. juliae Wagner & Masteller 1996 WM96
    |         |--P. ocellaris G20
    |         |--P. phalaenoides WI-B09
    |         |--P. puertoricana Wagner & Masteller 1996 WM96
    |         |--P. satchelli WM96
    |         `--P. spatulata M83
    `--Setomimini WI-B09
         |--Nemoneura liparotes WI-B09
         |--Didicrum pyramidon WI-B09
         |--Desmioza WI-B09
         |--Didimioza symphilia WI-B09
         |--Thrysocanthus WI-B09
         |--Valerianna WI-B09
         |--Neoarisemus maesi WI-B09
         |--Neurosystasis WI-B09
         |--Armillipora selvica Quate 1996 WI-B09
         |--*Australopericoma caudata [incl. A. wirthi] WI-B09
         |--Balbagathis discuspis WI-B09
         |--Setomima WI-B09
         |--Platyplastinx WI-B09
         |--Micrommatos WI-B09
         |    |--M. simplex WI-B09
         |    `--M. sylvaticum WI-B09
         |--Caenobrunettia WI-B09
         |    |--C. sarculosa WI-B09
         |    `--C. subditicia WI-B09
         |--Tonnoira WI-B09
         |    |--T. cavernicola WI-B09
         |    `--T. rapiformis WI-B09
         |--Alepia WI-B09
         |    |--A. diocula WI-B09
         |    |--A. hirtiventris WI-B09
         |    `--A. longinoi WI-B09
         `--Arisemus WI-B09
              |--*A. boxi WM96
              |--A. amydrus WI-B09
              |--A. buzbyae Wagner & Masteller 1996 WM96
              |--A. mariannae Wagner & Masteller 1996 WM96
              `--A. waidei Wagner & Masteller 1996 WM96

Psychodinae incertae sedis:
  Eurygarka helicis QV81
  Breviscapus QV81
  Neotelmatoscopus O98
  Paratelmatoscopus O98
  Epacretron O98
  Gerobrunettia O98
  Notiocharis O98
  Peripsychoda O98
  Ulomyia RD77
  Termitomastus RD77
  Termitodipteron RD77
  Termitadelphos RD77
  Thornburghiella marginalis BW09 [=Pericoma marginalis QV81]

*Type species of generic name indicated

REFERENCES

[BW09] Buck, M., N. E. Woodley, A. Borkent, D. M. Wood, T. Pape, J. R. Vockeroth, V. Michelsen & S. A. Marshall. 2009. Key to Diptera families—adults. In: Brown, B. V., A. Borkent, J. M. Cumming, D. M. Wood, N. E. Woodley & M. A. Zumbado (eds) Manual of Central American Diptera vol. 1 pp. 95–156. NRC Research Press: Ottawa.

[CM91] Colless, D. H., & D. K. McAlpine. 1991. Diptera (flies). In: CSIRO. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 2 pp. 717–786. Melbourne University Press: Carlton (Victoria).

[G20] Goldfuss, G. A. 1820. Handbuch der Naturgeschichte vol. 3. Handbuch der Zoologie pt 1. Johann Leonhard Schrag: Nürnberg.

[M83] Martin, N. A. 1983. Miscellaneous observations on a pasture fauna: an annotated species list. DSIR Entomology Division Report 3: 1–98.

[O98] Oosterbroek, P. 1998. The Families of Diptera of the Malay Archipelago. Brill: Leiden.

[QV81] Quate, L. W., & J. R. Vockeroth. 1981. Psychodidae. In: McAlpine, J. F., B. V. Peterson, G. E. Shewell, H. J. Teskey, J. R. Vockeroth & D. S. Wood (eds) Manual of Nearctic Diptera vol. 1 pp. 293–300. Research Branch, Agriculture Canada.

[RD77] Richards, O. W., & R. G. Davies. 1977. Imms' General Textbook of Entomology 10th ed. vol. 2. Classification and Biology. Chapman and Hall: London.

[WI-B09] Wagner, R., & S. Ibáñez-Bernal. 2009. Psychodidae (sand flies, and moth flies or owl flies). In: Brown, B. V., A. Borkent, J. M. Cumming, D. M. Wood, N. E. Woodley & M. A. Zumbado (eds) Manual of Central American Diptera vol. 1 pp. 319–335. NRC Research Press: Ottawa.

[WM96] Wagner, R., & E. C. Masteller. 1996. New moth flies (Diptera: Psychodidae) and a key to species from Puerto Rico. Proceedings of the Entomological Society of Washington 98 (3): 450–464.

[WT11] Wiegmann, B. M., M. D. Trautwein, I. S. Winkler, N. B. Barr, J.-W. Kim, C. Lambkin, M. A. Bertone, B. K. Cassel, K. M. Bayless, A. M. Heimberg, B. M. Wheeler, K. J. Peterson, T. Pape, B. J. Sinclair, J. H. Skevington, V. Blagoderov, J. Caravas, S. N. Kutty, U. Schmidt-Ott, G. E. Kampmeier, F. C. Thompson, D. A. Grimaldi, A. T. Beckenbach, G. W. Courtney, M. Friedrich, R. Meier & D. K. Yeates. 2011. Episodic radiations in the fly tree of life. Proceedings of the National Academy of Sciences of the USA 108 (14): 5690–5695.

Strepsiptera

Female Mengenilla moldryzki, from Pohl et al. (2012).


Belongs within: Holometabola.
Contains: Stylopiformia.

The Strepsiptera are a group of insects that develop as endoparasites of other insects. Adult males are free-living and non-feeding; mature females are larviform and those of Stylopidia remain within the original host.

See also: A queenage of Strepsiptera.

Characters (from Kathirithamby 1991): Endopterygote Neoptera with reduced mandibulate mouth-parts, extreme development of metathorax, reduced prothorax, and without differentiated trochanters in fore and mid legs. Adult males free living, with functional hind wings and small haltere-like fore wings. Females larviform, viviparous; usually parasitic, in puparium, and then with secondary progoneate genital apertures. Heteromorphosis during larval growth, 1st instar larvae free living and active, later instars parasitic.

Strepsiptera [Mengenillidia, Rhipiptera, Stylopida, Stylopoidea]
    |  i. s.: Neostylops aterrima A71
    |         Austrostylops gracilipes R35
    |         Bahiaxenos Bravo, Pohl et al. 2009 [Bahiaxenidae] BP09
    |           `--*B. relictus Bravo, Pohl et al. 2009 BP09
    |--Protoxenos [Protoxenidae] BP09
    |    `--P. janzeni Pohl, Beutel & Kinzelbach 2005 BP09
    `--+--Cretostylops engeli Grimaldi & Kathirithamby 2005 BP09
       `--+--Mengea [Mengeidae, Mengeiformia, Mengeina, Mengeoidea] BP09
          |    |--M. mengei GE05
          |    `--M. tertiaria K91
          `--+--Mengenillidae [Mengenilloidea] BP09
             |    |  i. s.: Eoxenos laboulbenei K91
             |    |--Iberoxeninae K91
             |    |--Congoxenos [Congoxeninae] K91
             |    `--Mengenilla [Mengenillinae] K91
             |         |--M. australiensis K91
             |         |--M. chobauti MHK11
             |         |--M. gracilipes K91
             |         `--M. moldryzki MW15
             `--Stylopidia [Stylopina] BP09
                  |  i. s.: Dundoxenos vilhenai A99
                  |         Bohartilla GE05 [Bohartillidae K91, Bohartilloidea]
                  |           |--B. kinzelbachi GE05
                  |           `--B. megalognatha GE05
                  |--Corioxenidae [Corioxenoidea] MHK11
                  |    |  i. s.: Uniclavinae K91
                  |    |--Corioxenos [Corioxeninae] MHK11
                  |    |    |--C. acucyrtophallus MHK11
                  |    |    `--C. antestiae K91
                  |    `--+--Blisseoxenos esaki MHK11
                  |       `--Triozocera [Triozocerinae] MHK11
                  |            `--T. papuana K91
                  `--Stylopiformia MHK11

*Type species of generic name indicated

REFERENCES

[A71] Askew, R. R. 1971. Parasitic Insects. Heinemann Educational Books: London.

[A99] Ax, P. 1999. Das System der Metazoa II. Ein Lehrbuch der phylogenetischen Systematik. Gustav Fisher Verlag: Stuttgart (transl. 2000. Multicellular Animals: The phylogenetic system of the Metazoa vol. 2. Springer).

[BP09] Bravo, F., H. Pohl, A. Silva-Neto & R. G. Beutel. 2009. Bahiaxenidae, a "living fossil" and a new family of Strepsiptera (Hexapoda) discovered in Brazil. Cladistics 25 (6): 614–623.

[GE05] Grimaldi, D., & M. S. Engel. 2005. Evolution of the Insects. Cambridge University Press: New York.

[K91] Kathirithamby, J. 1991. Strepsiptera. In: CSIRO. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 2 pp. 684–695. Melbourne University Press: Carlton (Victoria).

[MW15] McKenna, D. D., A. L. Wild, K. Kanda, C. L. Bellamy, R. G. Beutel, M. S. Caterino, C. W. Farnum, D. C. Hawks, M. A. Ivie, M. L. Jameson, R. A. B. Leschen, A. E. Marvaldi, J. V. McHugh, A. F. Newton, J. A. Robertson, M. K. Thayer, M. F. Whiting, J. F. Lawrence, A. Ślipiński, D. R. Maddison & B. D. Farrell. 2015. The beetle tree of life reveals that Coleoptera survived end-Permian mass extinction to diversify during the Cretaceous terrestrial revolution. Systematic Entomology 40 (4): 835–880.

[MHK11] McMahon, D. P., A. Hayward & J. Kathirithamby. 2011. The first molecular phylogeny of Strepsiptera reveals an early burst of molecular evolution correlated with the transition to endoparasitism. PLoS One 6 (6): e21206.

[R35] Rayment, T. 1935. A Cluster of Bees: Sixty essays on the life-histories of Australian bees, with specific descriptions of over 100 new species. Endeavour Press: Sydney.

Merizodinae

Merizodus soledadinus, from Lebouvier et al. (2020).


Belongs within: Conjunctae.

The Merizodinae are a group of small ground beetles with a disjunct distribution in southern South America, southern Australasia and northeastern Asia (Roig-Juñent & Cicchino 2001).

Characters (from Roig-Juñent & Cicchino 2001, as Zolini): Maxillary palpomere 3 plurisetose; elytron with apical plica, recurrent stria joined with apical portion of stria 5; tarsomeres dorsally sulcate.

<==Merizodinae [Merizodini, Oopterini, Zolini] R-JC01
    |--Sinozolina R-JC01
    |--Chaltenia Roig-Juñent & Cicchino 2001 [Chalteniina] R-JC01
    |    `--*C. patagonica Roig-Juñent & Cicchino 2001 R-JC01
    `--+--Oopterus Guérin-Ménéville 1841 [incl. Zolus Sharp 1886; Oopterina, Zolina] R-JC01
       |    `--‘Zolus’ helmsi H15
       `--Merizodus [Merizodina] R-JC01
            |--M. angustiocollis R-JC01
            `--M. soledadinus R-JC01

Merizodinae incertae sedis:
  Idacarabus Lea 1910 LB91, LM87
    |--I. cordicollis Moore 1967 LM87
    |--I. longicollis Moore 1978 LM87
    `--I. troglodytes Lea 1910 LM87
  Pterocyrtus Sloane 1920 B02, LM87
    |--*P. globosus Sloane 1920 LM87
    |--P. rubescens Sloane 1920 LM87
    |--P. striatulus Sloane 1920 LM87
    |--P. tasmanicus (Castelnau 1867) [=Drimostoma tasmanica] LM87
    `--P. truncaticollis Sloane 1923 LM87
  Sloaneana Csiki 1933 B02, LM87 (see below for synonymy)
    |--S. lamingtonensis Baehr 2002 B02
    |--S. similis Baehr 2002 B02
    `--S. tasmaniae (Sloane 1915) [=Brachydema tasmaniae; incl. B. victoriae Sloane 1915] LM87

Sloaneana Csiki 1933 B02, LM87 [=Brachydema Sloane 1915 non Fairmaire 1881 LM87, Sloanella Csiki 1928 non Jeannel 1927 LM87]

*Type species of generic name indicated

REFERENCES

[B02] Baehr, M. 2002. Two new species of Sloaneana Csiki from southern Queensland (Coleoptera, Carabidae, Merizodinae). Memoirs of the Queensland Museum 48: 9–15.

[H15] Hedley, C. 1915. Presidential address. Journal and Proceedings of the Royal Society of New South Wales 49 (1): 1–77, pls 1–7.

[LB91] Lawrence, J. F., & E. B. Britton. 1991. Coleoptera (beetles). In: CSIRO. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 2 pp. 543–683. Melbourne University Press: Carlton (Victoria).

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

[R-JC01] Roig-Juñent, S., & A. C. Cicchino. 2001. Chaltenia patagonica, new genus and species belonging to Chalteniina, a new subtribe of Zolini (Coleoptera: Carabidae). Canadian Entomologist 133: 651–670.

Megatominae

Variegated carpet beetle Anthrenus verbasci, copyright Olei.


Belongs within: Bostrichoidea.

The Megatominae are a group of dermestid beetles characterised by the presence of an ocellus together with a relatively long basal segment on the hind tarsus. A number of species are notorious storage pests.

<==Megatominae [Anthreninae] B14
    |--Anthrenocerus australis LB91
    |--Myrmeanthrenus frontalis LB91
    |--Thaumaglossa nigricans LB91
    |--Megatoma undata (Linnaeus 1758) G20, L58 [=Dermestes undatus G20]
    |--Trogoderma B74
    |    |--T. adelaidae R35
    |    |--T. granarium MHG04
    |    |--T. variabile MHG04
    |    `--T. versicolor B70
    `--Anthrenus B14
         |--A. caucasicus C01
         |--A. fuscus C01
         |--A. museorum (Linnaeus 1761) B14
         |--A. pulchellus Gestro 1889 G89
         |--A. scrophulariae B70
         |--A. subclaviger G89
         |--A. verbasci B14 [=Nathrenus verbasci OM91]
         `--A. vorax B70

*Type species of generic name indicated

REFERENCES

[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature's gems. Ivy Press: Lewes (United Kingdom).

[B70] Britton, E. B. 1970. Coleoptera (beetles). In: CSIRO. The Insects of Australia: A textbook for students and research workers pp. 495–621. Melbourne University Press.

[B74] Britton, E. B. 1974. Coleoptera (beetles). In: CSIRO. The Insects of Australia: A textbook for students and research workers. Supplement 1974 pp. 62–89. Melbourne University Press.

[C01] Csiki, E. 1901. Bogarak [Coleopteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 75–120. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[G89] Gestro, R. 1889. Viaggio ab Assab nel Mar Rosso dei signori G. Doria ed O. Beccari con il R. Avviso “Esploratore” dal 16 Novembre 1879 al 26 Febbraio 1880.—IV. Coleotteri. Annali del Museo Civico di Storia Naturale di Genova, Serie 2a, 7: 5–72.

[G20] Goldfuss, G. A. 1820. Handbuch der Naturgeschichte vol. 3. Handbuch der Zoologie pt 1. Johann Leonhard Schrag: Nürnberg.

[LB91] Lawrence, J. F., & E. B. Britton. 1991. Coleoptera (beetles). In: CSIRO. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 2 pp. 543–683. Melbourne University Press: Carlton (Victoria).

[MHG04] Maynard, G. V., J. G. Hamilton & J. F. Grimshaw. 2004. Quarantine—phytosanitary, sanitary and incursion management: an Australian entomological perspective. Australian Journal of Entomology 43 (3): 318–328.

[OM91] Okamoto, M., & K. Matsumoto. 1991. Observations on the behavior of the hypopus of Lardoglyphus konoi. In: Dusbábek, F., & V. Bukva (eds) Modern Acarology: Proceedings of the VIII International Congress of Acarology, held in České Budĕjovice, Czechoslovakia, 6–11 August 1990 vol. 2 pp. 555–559. SPB Academic Publishing: The Hague.

[R35] Rayment, T. 1935. A Cluster of Bees: Sixty essays on the life-histories of Australian bees, with specific descriptions of over 100 new species. Endeavour Press: Sydney.

Ptinus

Whitemarked spider beetle Ptinus fur, copyright Joseph Berger.


Belongs within: Ptinidae.

Ptinus, spider beetles, is a cosmopolitan genus of long-legged beetles with elytra bearing a vestiture of appressed scales or setae. A number of species are widespread as storage pests.

Ptinus
    |--P. adeps Olliff 1886 M86
    |--P. albomaculatus Macleay 1872 M86
    |--P. attritus Olliff 1886 M86
    |--P. bicinctus C01
    |--P. carinatus Lucas 1846 E12
    |--P. clavipes [incl. P. mobilis] B70
    |--P. egenus Olliff 1886 M86
    |--P. eminens Olliff 1886 M86
    |--P. exulans Erichs. 1842 M86
    |--P. fossulatus Lucas 1846 E12
    |--P. fur Linn. 1761 M86
    |--P. gibbicollis Lucas 1846 E12
    |--P. hirticollis Lucas 1846 E12
    |--P. imulus Olliff 1886 M86
    |--P. latro C01
    |--P. longus Olliff 1886 M86
    |--P. mauritanicus Lucas 1846 E12
    |--P. obesus Lucas 1846 E12
    |--P. pusillus LB91
    |--P. rotundicollis Lucas 1846 E12
    |--P. rufus Lucas 1846 E12
    `--P. tectus [incl. P. pilosus White 1846] M86

*Type species of generic name indicated

REFERENCES

[B70] Britton, E. B. 1970. Coleoptera (beetles). In: CSIRO. The Insects of Australia: A textbook for students and research workers pp. 495–621. Melbourne University Press.

[C01] Csiki, E. 1901. Bogarak [Coleopteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy], pp. 75–120. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[E12] Evenhuis, N. L. 2012. Publication and dating of the Exploration Scientifique de l’Algérie: Histoire Naturelle des Animaux Articulés (1846–1849) by Pierre Hippolyte Lucas. Zootaxa 3448: 1–61.

[LB91] Lawrence, J. F., & E. B. Britton. 1991. Coleoptera (beetles). In: CSIRO. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 2 pp. 543–683. Melbourne University Press: Carlton (Victoria).

[M86] Masters, G. 1886. Catalogue of the described Coleoptera of Australia. Part IV. Proceedings of the Linnean Society of New South Wales, series 2, 1 (2): 259–380.