Scaritinae

Dyschirius aeneus, copyright Lech Borowiec.


Belongs within: Geadephaga.
Contains: Laccopterum, Scaraphites, Conopterum, Neocarenum, Monocentrum, Euryscaphus, Carenum.

The Scaritinae are a diverse group of ground beetles with body forms commonly modified for digging. Members of the group have the fore coxal cavities closed behind and the mid coxal cavities laterally open.

<==Scaritinae [Scarititae, Scrobifera] R-JC01
    |--Migadopini [Migadopinae] R-JC01
    |    |--Migadops latus R-JC01
    |    `--Stichonotus Sloane 1910 LB91, LM87
    |         |--*S. leai Sloane 1910 LM87
    |         |--S. limbatus Sloane 1915 LM87
    |         `--S. piceus Sloane 1915 LM87
    `--Scaritini B90
         |--Schizogenius B90
         |--Aspidoglossa subangulata B90
         |--Oxydrepanus rufus B90
         |--Euryscaphus LB91
         |--Carenum LB91
         |--Pasimachus B90
         |    |--P. elongatus BB68
         |    `--P. subangulatus (Chaudoir 1862) B14
         |--Scarites B90
         |    |--S. eurytus G89
         |    |--S. gigas C74
         |    |--S. guineensis [incl. S. guineensis var. asphaltinus] G89
         |    |--S. pyracmon B35
         |    |--S. rugosus B35
         |    |--S. senegalensis B55
         |    `--S. subterraneus BB68
         `--Dyschirius Bonelli 1810 B90, LM87
              |--‘Carabus’ globosus Herbst 1783 [incl. Scarites gibbus Fabricius 1792, *Dyschirius gibbus] LM87
              |--D. aeneus C01
              |--D. macleayi Sloane 1896 LM87
              |--D. mastersii Macleay 1866 LM87
              |--D. ovensensis Blackburn 1891 LM87
              |--D. stephensii Macleay 1865 LM87
              |--D. torrensensis Blackburn 1890 LM87
              |--D. wilsoni Sloane 1923 LM87
              `--D. zonatus Putzeys 1868 LM87

Scaritinae incertae sedis:
  Ochyropus gigas B55
  Taeniolobus B55
    |--T. laevipennis B55
    |--T. lucidus B55
    |    |--T. l. lucidus B55
    |    `--T. l. strigiceps B55
    `--T. vilhenai Basilewsky 1955 B55
  Trilophus congoanus B55
  Syleter Andrewes 1941 [=Psilus Putzeys 1877 nec Panzer 1806 nec Fischer de Waldheim 1813] LM87
    |--*S. paradoxus (Putzeys 1868) [=Ardistomis paradoxa] LM87
    |--S. andrewesi (Basilewsky 1931) [=Dyschirius andrewesi, Psilus andrewesi] B55
    |--S. congoensis B55
    `--S. papua Darlington 1962 LM87
  Pseudoclivina Kult 1947 B55
    `--P. grandis B55
  Rhysocara Sloane 1916 LM87
    `--*R. crassa Sloane 1916 LM87
  Platysphyrus Sloane 1905 LM87
    `--*P. tibialis Sloane 1905 LM87
  Clivinarchus Sloane 1896 LM87
    `--*C. perlongus Sloane 1896 LM87
  Laccopterum LM87
  Philoscaphus Macleay 1871 LM87
    |--*P. tuberculatus (Macleay 1863) [=Carenum tuberculatum] LM87
    |--P. barnardi Macleay 1888 LM87
    |--P. bicostatus Sloane 1905 LM87
    |--P. carinatus (Macleay 1864) [=Carenum carinatum] LM87
    |--P. costalis Macleay 1873 (see below for synonymy) LM87
    `--P. mastersii Macleay 1871 [incl. P. duboulayi Blackburn 1890] LM87
  Scaraphites LM87
  Conopterum LM87
  Neocarenum LM87
  Trichocarenum Blackburn 1892 LM87
    |--*T. elderi Blackburn 1892 LM87
    |--T. castelnaui Sloane 1905 LM87
    `--T. cylindricum Sloane 1897 LM87
  Neoscaphus Sloane 1888 LM87
    `--*N. simplex Sloane 1888 LM87
  Epilectus Blackburn 1888 [=Carenarchus Sloane 1905, Eurygnathus Blackburn 1888 non Wollaston 1854] LM87
    |--*E. fortis (Blackburn 1888) [=*Eurygnathus fortis, *Carenarchus fortis] LM87
    `--E. mastersi (Macleay 1869) [=Neocarenum mastersi] LM87
  Monocentrum LM87
  Steganomma Macleay 1887 LM87
    `--*S. porcatum Macleay 1887 LM87
  Geoscaptus Chaudoir 1855 LM87
    |--*G. laevissimus Chaudoir 1855 (see below for synonymy) LM87
    |--G. cacus (Macleay 1863) (see below for synonymy) LM87
    |--G. crassus Sloane 1895 LM87
    |--G. macleayi Chaudoir 1879 LM87
    `--G. plicatulus (Castelnau 1867) [=Scarites plicatulus] LM87
  Akephorus B14
    |--A. marinus B14
    `--A. obesus LeConte 1866 B14
  Omoglymmius B14
    |--O. americanus (Laporte 1836) B14
    `--O. hamatus B14
  Solenogenys funkei Adis 1981 B14

Geoscaptus cacus (Macleay 1863) [=Scarites cacus; incl. S. approximatus Macleay 1865, S. bipunctatus Castelnau 1867, S. mitchellii Castelnau 1867, S. ruficornis Castelnau 1867, S. subporcatulus Macleay 1865, S. substriatus Castelnau 1867, S. waterhousei Macleay 1865] LM87

*Geoscaptus laevissimus Chaudoir 1855 [incl. Scarites bostocki Castelnau 1867, S. damastes Macleay 1863, S. geryon Macleay 1863, S. jacksoniensis Macleay 1865, S. planiusculus Macleay 1865] LM87

Philoscaphus costalis Macleay 1873 [incl. P. crassus Blackburn 1888, P. lateralis Macleay 1873, P. tepperi Blackburn 1888] LM87

*Type species of generic name indicated

REFERENCES

[BB68] Basford, N. L., J. E. Butler, C. A. Leone & F. J. Rohlf. 1968. Immunologic comparisons of selected Coleoptera with analyses of relationships using numerical taxonomic methods. Systematic Zoology 17 (4): 388–406.

[B55] Basilewsky, P. 1955. Carabidae (Coleoptera) de l’Angola (première partie). Diamang Publicações Culturais 27: 91–137.

[B90] Bell, R. T. 1990. Insecta: Coleoptera Carabidae adults and larvae. In: Dindal, D. L. (ed.) Soil Biology Guide pp. 1053–1092. John Wiley & Sones: New York.

[B35] Boisduval, J. B. 1835. Voyage de Découvertes de l’Astrolabe. Exécuté par ordre du Roi, pendant les années 1826–1827–1828–1829, sous le commandement de M. J. Dumont d'Urville. Faune entomologique de l'océan Pacifique, avec l'illustration des insectes nouveaux recueillis pendant le voyage vol. 2. Coléoptères et autres ordres. J. Tastu: Paris.

[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature's gems. Ivy Press: Lewes (United Kingdom).

[C74] Coineau, Y. 1974. Éléments pour une monographie morphologique, écologique et biologique des Caeculidae (Acariens). Mémoires du Muséum National d’Histoire Naturelle, nouvelle série, Série A, Zoologie 81: 1–299, 24 pls.

[C01] Csiki, E. 1901. Bogarak [Coleopteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 75–120. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[G89] Gestro, R. 1889. Viaggio ab Assab nel Mar Rosso dei signori G. Doria ed O. Beccari con il R. Avviso «Esploratore» dal 16 Novembre 1879 al 26 Febbraio 1880.—IV. Coleotteri. Annali del Museo Civico di Storia Naturale di Genova, Serie 2a, 7: 5–72.

[LB91] Lawrence, J. F., & E. B. Britton. 1991. Coleoptera (beetles). In: CSIRO. The Insects of Australia: A textbook for students and research workers vol. 2 pp. 543–683. Melbourne University Press: Carlton (Victoria).

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

[R-JC01] Roig-Juñent, S., & A. C. Cicchino. 2001. Chaltenia patagonica, new genus and species belonging to Chalteniina, a new subtribe of Zolini (Coleoptera: Carabidae). Canadian Entomologist 133: 651–670.

Adelotopus

Adelotopus haemorrhoidalis, copyright Udo Schmidt.


Belongs within: Pseudomorphini.

Adelotopus is a genus of pseudomorphin ground beetles found in Australia and south-east Asia, characterised by a reduced chaetotaxy and foliaceous female stylomeres (Baehr 2008).

<==Adelotopus Hope 1834 B08
    |--*A. gyrinoides Hope 1834 B08
    |--A. adustus Baehr 1997 B08
    |--A. affinis Castelnau 1867 LM87
    |--A. analis Macleay 1871 LM87
    |--A. aphodioides Westwood 1853 LM87
    |--A. apicalis Macleay 1864 LM87
    |--A. basalis Baehr 2008 B08
    |--A. bimaculatus Macleay 1864 [incl. Silphomorpha rufoguttata Blackburn 1893] LM87
    |--A. bolitus OM08
    |--A. brevipennis Macleay 1888 B08
    |--A. brunneus Castelnau 1867 LM87
    |--A. celeripes Lea 1910 LM87
    |--A. coriaceus Baehr 1997 B08
    |--A. cornutus Castelnau 1867 LM87
    |--A. dysticides Newman 1842 [incl. A. fortnumi Hope 1845] LM87
    |--A. elongatulus Macleay 1888 LM87
    |--A. fasciatus Castelnau 1867 LM87
    |--A. filiformis Castelnau 1867 LM87
    |--A. flavus B08
    |--A. haemorrhoidalis Erichson 1842 [incl. A. inquinatus Newman 1842] LM87
    |--A. houstoni Baehr 1997 B08
    |--A. hydrobioides Westwood 1853 LM87
    |--A. insignis Sloane 1910 LM87
    |--A. laevis Macleay 1888 LM87
    |--A. linearis Macleay 1888 LM87
    |--A. longipennis Macleay 1888 LM87
    |--A. maculipennis Macleay 1871 LM87
    |--A. mastersii Macleay 1871 LM87
    |--A. micans Blackburn 1901 LM87
    |--A. minutus Baehr 2008 B08
    |--A. nemosomoides Westwood 1853 LM87
    |--A. niger Notman 1925 LM87
    |--A. occidentalis Castelnau 1867 LM87
    |--A. paroensis Castelnau 1867 B08
    |--A. piceus B08
    |--A. pilbarae Baehr 2008 B08
    |--A. politus Castelnau 1867 LM87
    |--A. punctatus Castelnau 1867 LM87
    |--A. puncticollis Notman 1925 LM87
    |    |--A. p. puncticollis B08
    |    `--A. p. angustemaculatus Baehr 1997 B08
    |--A. rubiginosus Newman 1856 B08 [incl. A. castaneus Castelnau 1867 B08, LM87]
    |--A. rufescens B08
    |--A. scolytides Newman 1842 LM87
    |--A. seriepunctatus Notman 1925 LM87
    |--A. tasmani Blackburn 1901 LM87
    |--A. variolosus Lea 1910 LM87
    |--A. vicinus Castelnau 1867 LM87
    `--A. zonatus Castelnau 1867 LM87

*Type species of generic name indicated

REFERENCES

[B08] Baehr, M. 2008. New and rare pseudomorphine species from Western Australia (Insecta, Coleoptera, Carabidae, Pseudomorphinae). Records of the Western Australian Museum 24 (2): 151–180.

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

[OM08] Ober, K. A., & D. R. Maddison. 2008. Phylogenetic relationships of tribes within Harpalinae (Coleoptera: Carabidae) as inferred from 28S ribosomal DNA and the wingless gene. Journal of Insect Science 8: 63.

Pseudomorphini

Graphipterus serrator, copyright Hectonichus.


Belongs within: Harpalinae.
Contains: Adelotopus, Sphallomorpha.

The Pseudomorphini are a group of often flattened ground beetles found living under bark. They are often found in association with ants and larvae of some species are physogastric inquilines in ant nests (Lawrence & Britton 1991). The majority of genera in this group are ovoviviparous (Baehr 2008).

Characters (from Lawrence & Britton 1991): Antennal scape not visible from above; antennae sometimes short and clavate; head with deep antennal sulcus between eye and mandible; legs more or less concealed beneath body, with may be flattened and laterally explanate.

Pseudomorphini [Pseudomorphinae]
    |  i. s.: Cryptocephalomorpha B08
    |--+--Pseudomorpha Kirby 1825 OM08, LM87
    |  |    |--*P. excrucians Kirby 1825 LM87
    |  |    |--P. angustata OM08
    |  |    `--P. confusa Notman 1925 LM87
    |  `--Graphipterus Latreille 1802 OM08, L02 [Graphipterini]
    |       |--G. circumdatus Raffr. 1885 G89
    |       |--G. cordiger OM08
    |       |--G. limbatus OM08
    |       |--G. minutus B14
    |       |--G. serrator (Forskål 1775) [=Carabus serrator] G89
    |       |--G. trilineatus [=Carabus trilineatus] L02
    |       `--G. variegatus [=Carabus variegatus] L02
    `--+--+--Adelotopus OM08
       |  |--Paussotropus Waterhouse 1877 B08
       |  |    `--P. cylindricus (Chaudoir 1862) (see below for synonymy) B08
       |  `--Cainogenion Notman 1925 B08
       |       |--*C. ipsoides (Newman 1837) [=Adelotopus ipsoides] B08
       |       |    |--C. i. ipsoides B08
       |       |    `--C. i. occidentale Baehr 1997 B08
       |       |--C. creberrimum (Blackburn 1901) [=Adelotopus creberrimus] LM87
       |       |--C. ephippiatum (Newman 1856) [=Adelotopus ephippiatus] LM87
       |       `--C. obscurum (Castelnau 1867) [=Adelotopus obscurus; incl. A. subopacus Macleay 1871] LM87
       `--+--Orthogonius [Orthogoniinae, Orthogoniini] OM08
          |    |--O. brevilabris B55
          |    |--O. clarkei B55
          |    |--O. davidis B89
          |    |--O. duplicatus [incl. O. alternans] B89
          |    `--O. profundestriatus B89
          `--Sphallomorpha OM08

Paussotropus cylindricus (Chaudoir 1862) [=Adelotopus cylindricus, Cainogenion cylindricum; incl. *Paussotropus parellelus Waterhouse 1877] B08

*Type species of generic name indicated

REFERENCES

[B08] Baehr, M. 2008. New and rare pseudomorphine species from Western Australia (Insecta, Coleoptera, Carabidae, Pseudomorphinae). Records of the Western Australian Museum 24 (2): 151–180.

[B55] Basilewsky, P. 1955. Carabidae (Coleoptera) de l’Angola (première partie). Diamang Publicações Culturais 27: 91–137.

[B89] Bates, H. W. 1889. Viaggio di Leonardo Fea in Birmani e regioni vicine. XVI.—On some Carabidae from Burma collected by Mr. L. Fea. Annali del Museo Civico di Storia Naturale di Genova, Serie 2a, 7: 100–111.

[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature's gems. Ivy Press: Lewes (United Kingdom).

[G89] Gestro, R. 1889. Viaggio ab Assab nel Mar Rosso dei signori G. Doria ed O. Beccari con il R. Avviso «Esploratore» dal 16 Novembre 1879 al 26 Febbraio 1880.—IV. Coleotteri. Annali del Museo Civico di Storia Naturale di Genova, Serie 2a, 7: 5–72.

[L02] Latreille, P. A. 1802. Histoire Naturelle, générale et particulière des crustacés et des insectes vol. 3. Familles naturelles des genres. F. Dufart: Paris.

Lawrence, J. F., & E. B. Britton. 1991. Coleoptera (beetles). In: CSIRO. The Insects of Australia: A textbook for students and research workers vol. 2 pp. 543–683. Melbourne University Press: Carlton (Victoria).

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

[OM08] Ober, K. A., & D. R. Maddison. 2008. Phylogenetic relationships of tribes within Harpalinae (Coleoptera: Carabidae) as inferred from 28S ribosomal DNA and the wingless gene. Journal of Insect Science 8: 63.

Neocarenum

Neocarenum angustatum, copyright Museums Victoria.


Belongs within: Scaritinae.

Neocarenum is an Australian genus of scaritine ground beetles characterised by moniliform antennae, sub-securiform labial palpi and bidentate anterior tibiae.

<==Neocarenum Castelnau 1867 LM87
    |--N. singulare Castelnau 1867 [=N. singularis] LM87
    |--N. angustatum Sloane 1894 LM87
    |--N. blackburni Sloane 1895 LM87
    |--N. cylindripenne Bates 1874 LM87
    |--N. dingo Sloane 1916 LM87
    |--N. elongatum (Macleay 1864) [=Carenum elongatum; incl. N. kreusleri Castelnau 1867] LM87
    |--N. parviceps Sloane 1894 LM87
    |--N. retusum Bates 1874 LM87
    |--N. rugulosum Macleay 1869 LM87
    `--N. spenceri Sloane 1897 LM87

*Type species of generic name indicated

REFERENCES

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

Conopterum

Conopterum sp., copyright Reiner Richter.


Belongs within: Scaritinae.

Conopterum is an Australian genus of scaritine ground beetles, characterised by slender, filiform antennae and bidentate anterior tibiae.

<==Conopterum Chaudoir 1868 [incl. Carenidium Chaudoir 1868] LM87
    |--*C. insigne Chaudoir 1868 LM87
    |--C. aberrans (Sloane 1897) [=Eutoma aberrans] LM87
    |--C. atrum (Sloane 1916) [=Carenidium atrum] LM87
    |--C. bicornutum Macleay 1887 LM87
    |--C. bifurcum (Sloane 1916) [=Carenidium bifurcum] LM87
    |--C. chaudoirii (Macleay 1887) [=Carenidium chaudoirii] LM87
    |--C. damelii (Macleay 1869) [=Carenidium damelii] LM87
    |--C. darlingense (Macleay 1887) [=Carenidium darlingense] LM87
    |--C. frenchi (Sloane 1916) [=Carenidium frenchi] LM87
    |--C. gagatinum (Macleay 1864) [=Carenum gagatinum, *Carenidium gagatinum] LM87
    |--C. leai (Sloane 1897) [=Carenidium leai] LM87
    |--C. longipenne (Sloane 1907) [=Carenidium longipenne] LM87
    |--C. modestum Sloane 1888 LM87
    |--C. mucronatum (Macleay 1866) [=Carenum mucronatum] LM87
    |--C. pertenue (Sloane 1916) [=Carenidium pertenue] LM87
    |--C. purpuratum (Sloane 1905) [=Carenidium purpuratum] LM87
    |--C. pyripenne Sloane 1900 LM87
    |--C. riverinae (Macleay 1865) (see below for synonymy) LM87
    |--C. sapphirinum (Bates 1874) [=Carenidium sapphirinum] LM87
    |--C. septentrionale (Macleay 1887) [=Carenidium septentrionale] LM87
    |--C. spaldingii (Macleay 1878) [=Carenidium spaldingii] LM87
    |--C. superbum (Castelnau 1867) (see below for synonymy) LM87
    `--C. tropicale (Macleay 1887) [=Carenidium tropicale] LM87

Conopterum riverinae (Macleay 1865) [=Carenum riverinae; incl. Ca. amabile Castelnau 1867, Conopterum barnardi Macleay 1887, Co. incornutum Macleay 1887, Monocentrum insigne Chaudoir 1868, Co. littorale Macleay 1887, Co. violaceum Macley 1887] LM87

Conopterum superbum (Castelnau 1867) [=Carenum superbum; incl. Carenidium kreuslerae Macleay 1869, Carenidium lacustre Macleay 1873] LM87

*Type species of generic name indicated

REFERENCES

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

Carenum

Carenum optimum, copyright Noël Mal.


Belongs within: Scaritinae.

Carenum is an Australian genus of scaritine ground beetles, characterised by ovate elytra, bidentate anterior tibiae, an inner subapical puncture on the anterior femora, and a truncate clypeus.

<==Carenum Bonelli 1813 (see below for synonymy) LM87
    |--C. bonellii Westwood 1842 (see below for synonymy) LM87
    |--C. acutipes Sloane 1897 LM87
    |--C. adelaidae (Blackburn 1888) [=Eutoma adelaidae] LM87
    |--C. affine Macleay 1864 LM87
    |--C. amplicolle Sloane 1897 LM87
    |--C. angustipenne Macleay 1871 LM87
    |--C. anthracinum Macleay 1864 (see below for synonymy) LM87
    |--C. batesi Masters 1885 [=C. planipenne Bates 1874 non Macleay 1873] LM87
    |--C. bellum Sloane 1917 LM87
    |--C. brevicolle Sloane 1894 LM87
    |--C. breviforme Bates 1874 LM87
    |--C. brevipenne (Macleay 1887) [=Eutoma brevipenne] LM87
    |--C. brisbanense Castelnau 1867 [=C. brisbanensis; incl. C. opacum Macleay 1869] LM87
    |--C. browni Sloane 1916 LM87
    |--C. carbonarium Castelnau 1867 [incl. C. vicinum Sloane 1890] LM87
    |--C. cavipenne (Bates 1874) [=Eutoma cavipenne] LM87
    |--C. cognatum Sloane 1895 LM87
    |--C. concinnum Sloane 1905 LM87
    |--C. convexum Chaudoir 1868 LM87
    |--C. coracinum Macleay 1865 [incl. Corenoscaphus lucidus Macleay 1887] LM87
    |--C. cordipenne Sloanea 1897 LM87
    |--C. coruscum Macleay 1864 [=C. corruscum (l. c.)] LM87
    |--C. cupreomarginatum Blackburn 1888 (see below for synonymy) LM87
    |--C. cupripenne Macleay 1863 LM87
    |--C. decorum Sloane 1888 LM87
    |--C. devastator Castelnau 1867 LM87
    |--C. devisii Macleay 1883 LM87
    |--C. digglesi (Macleay 1869) [=Eutoma digglesi] LM87
    |--C. dispar Macleay 1869 LM87
    |--C. distinctum Macleay 1864 LM87
    |--C. ducale Sloane 1905 [=C. frenchi Sloane 1895 non Eutoma frenchi Sloane 1894] LM87
    |--C. elegans Macleay 1864 (see below for synonymy) LM87
    |--C. emarginatum Sloane 1900 [incl. C. emarginatum aequalis Sloane 1916] LM87
    |--C. episcopale (Castelnau 1867) [=Eutoma episcopalis] LM87
    |--C. eximium Sloane 1916 LM87
    |--C. felix Sloane 1888 [=C. (*Paliscaphus) felix] LM87
    |--C. filiforme (Castelnau 1867) [=Eutoma filiforme] LM87
    |--C. floridum Sloane 1917 [=C. lepidum Sloane 1916 non Sloane 1890] LM87
    |--C. formosum Sloane 1907 LM87
    |--C. foveolatum (Macleay 1888) non Macleay 1864 (not preoc. if in dif. gen.) [=Calliscapterus foveolatus] LM87
    |--C. frenchi (Sloane 1894) [=Eutoma frenchi] LM87
    |--C. froggatti Sloane 1897 LM87
    |--C. fugitivum Blackburn 1888 LM87
    |--C. fulgidum Sloane 1917 LM87
    |--C. gratiosum (Sloane 1897) [=Eutoma gratiosum] LM87
    |--C. habile Sloane 1892 LM87
    |--C. habitans Sloane 1890 [=C. (Calliscapterus) habitans] LM87
    |--C. imitator Sloane 1897 LM87
    |--C. inconspicuum Blackburn 1888 LM87
    |--C. ineditum Macleay 1869 LM87
    |--C. interiore Sloane 1888 [=C. (Platythorax) interiore] LM87
    |--C. interruptum Macleay 1865 (see below for synonymy) LM87
    |--C. iridescens Sloane 1894 LM87
    |--C. janthinum Macleay 1883 [=C. ianthinum] LM87
    |--C. kingii Macleay 1869 [incl. C. propinquum Macleay 1869] LM87
    |--C. laevigatum Macleay 1864 (see below for synonymy) LM87
    |--C. laevipenne Macleay 1863 [incl. C. laevicolle Sloane 1888] LM87
    |--C. laterale Macleay 1865 LM87
    |--C. leai Sloane 1916 LM87
    |--C. lepidum Sloane 1890 LM87
    |--C. levissimum (Sloane 1900) [=Eutoma levissime] LM87
    |--C. lobatum Sloane 1900 LM87
    |--C. longulum Sloane 1916 [incl. C. longulum atroviridis Sloane 1916] LM87
    |--C. macleayi Blackburn 1888 LM87
    |--C. magnificum (Macleay 1887) [=Eutoma magnificum] LM87
    |--C. marginatum (Boisduval 1835) [=*Arnidius marginatus] LM87
    |--C. montanum Sloane 1916 LM87
    |--C. morosum Sloane 1907 LM87
    |--C. nickerli Ancey 1880 LM87
    |--C. nitidipes Sloane 1916 LM87
    |--C. obsoletum Macleay 1888 LM87
    |--C. occidentale Sloane 1897 LM87
    |--C. odewahnii Castelnau 1867 [incl. C. ordinatum Macleay 1869] LM87
    |--C. opacicolle Sloane 1897 LM87
    |--C. optimum Sloane 1895 LM87
    |--C. ovale Sloane 1900 LM87
    |--C. parvulum Macleay 1873 LM87
    |--C. perplexum White 1841 LM87
    |--C. planipenne Macleay 1873 LM87
    |--C. politissimum (Chaudoir 1868) [=Eutoma politissimum] LM87
    |--C. politum Westwood 1842 LM87
    |--C. porphyreum Bates 1874 LM87
    |--C. pulchrum Sloane 1897 LM87
    |--C. puncticolle Macleay 1864 LM87
    |--C. punctipenne (Macleay 1883) [=Eutoma punctipenne] LM87
    |--C. purpuratum (Castelnau 1867) [=Eutoma purpurata] LM87
    |--C. purpureum Sloane 1897 LM87
    |--C. pusillum Macleay 1883 LM87
    |--C. quadripunctatum Macleay 1863 [=*Carenoscaphus quadripunctatus] LM87
    |--C. rectangulare Macleay 1864 [=*Platythorax rectangularis] LM87
    |--C. reflexum Sloane 1897 LM87
    |--C. regulare Sloane 1900 LM87
    |--C. rutilans Sloane 1907 LM87
    |--C. scaritoides Westwood 1843 (see below for synonymy) LM87
    |--C. serratipes Sloane 1900 LM87
    |--C. simile Macleay 1865 [incl. C. ovipenne Macleay 1871, C. triste Macleay 1869] LM87
    |--C. smaragdulum Westwood 1842 LM87
    |--C. speciosum Sloane 1888 [=C. (Calliscapterus) speciosus] LM87
    |--C. splendens Castelnau 1867 LM87
    |--C. splendidum Macleay 1863 LM87
    |--C. striatopunctatum Macleay 1865 LM87
    |--C. subcostatum Macleay 1865 LM87
    |--C. subcyaneum Macleay 1869 [incl. C. rugatum Blackburn 1888, C. sulcaticeps Sloane 1897] LM87
    |--C. submetallicum Macleay 1871 LM87
    |--C. subplanatum Bates 1874 LM87
    |--C. subporcatulum Macleay 1865 [incl. C. politulum Macleay 1871, C. viridimarginatum Macleay 1871] LM87
    |--C. sumptuosum Westwood 1842 LM87
    |--C. terraereginae Macleay 1883 LM87
    |--C. tibiale Sloane 1894
    |--C. tinctilatum (Newman 1838) (see below for synonymy) LM87
    |--C. transversicolle Chaudoir 1868 LM87
    |--C. tumidipes Sloane 1900 LM87
    |--C. venustum Sloane 1897 LM87
    |--C. versicolor Sloane 1897 LM87
    |--C. violaceum Macleay 1864 LM87
    |--C. virescens Sloane 1894 [incl. C. virescens viridiventris Sloane 1916] LM87
    |--C. viridiaeneum (Macleay 1888) [=Calliscapterus viridiaeneus] LM87
    |--C. viridicolor (Sloane 1895) [=Eutoma viridicolor] LM87
    `--C. viridissimum (Macleay 1888) [=Carenoscaphus viridissimus] LM87

Carenum Bonelli 1813 [incl. Arnidius Boisduval 1835, Calliscapterus Macleay 1887, Carenoscaphus Macleay 1887, Chariscapterus Sloane 1888, Eutoma Newman 1838, Paliscaphus Sloane 1888, Platythorax Macleay 1887] LM87

Carenum anthracinum Macleay 1864 [incl. C. cyanipenne Macleay 1869, C. ebeninum Castelnau 1867, C. gracile Sloane 1895, C. nitescens Macleay 1869] LM87

Carenum bonellii Westwood 1842 [=Scarites cyaneus Bonelli 1813 non Fabricius 1775, *C. cyaneum; incl. C. chaudoiri Macleay 1869, C. scitulum Macleay 1864, C. viridipenne Westwood 1849, C. westwoodii Castelnau 1867] LM87

Carenum cupreomarginatum Blackburn 1888 [=*Chariscapterus cupreomarginatus; incl. Ca. (Chariscapterus) opulens Sloane 1888] LM87

Carenum elegans Macleay 1864 [incl. Car. campestre Macleay 1865, *Calliscapterus campestris, Car. rufipes Macleay 1869] LM87

Carenum interruptum Macleay 1865 [incl. C. arenarium Sloane 1888, C. castelnaui Chaudoir 1868, C. obscurum Macleay 1865, C. occultum Macleay 1871, C. purpureomarginatum Macleay 1887, C. sexpunctatum Macleay 1869] LM87

Carenum laevigatum Macleay 1864 [incl. C. frontale Macleay 1865, C. murrumbidgense Macleay 1865, C. punctulatum Macleay 1864, C. schomburgkii Castelnau 1867] LM87

Carenum scaritoides Westwood 1843 [incl. C. ambiguum Macleay 1865, C. atronitens Macleay 1864, C. atronitens Castelnau 1867 non Macleay 1864, C. gawlerense Macleay 1869, C. ignotus Sloane 1892, C. intermedium Westwood 1849, C. nigerrimum Macleay 1865, C. oblongum Macleay 1864, C. striatopunctulatum Macleay 1865, C. subquadratum Macleay 1865] LM87

Carenum tinctilatum (Newman 1838) [=*Eutoma tinctilatus; incl. C. bipuncatum Macleay 1863, C. glaberrimum Macleay 1865, Eutoma laevis Castelnau 1867, E. loddonensis Castelnau 1867, E. mastersi Macleay 1869, E. newmanni Castelnau 1867, E. punctulatum Macleay 1887 non Carenum punctulatum Macleay 1864, Carenum subrugulosum Macleay 1865, C. substriatulum Macleay 1865, C. undulatum Macleay 1865] LM87

*Type species of generic name indicated

REFERENCES

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

Scaraphites

Scaraphites sp., from Friends of Queens Park Bushland.


Belongs within: Scaritinae.

Scaraphites is an Australian genus of scaritine ground beetles characterised by filiform palpi and fore tibiae with the second external tooth nearer the apex than the inner apical spine.

<==Scaraphites Westwood 1842 LM87
    |--S. rotundipennis (Dejean 1825) (see below for synonymy) LM87
    |--S. confusus (Westwood 1842) [=Scarites (Scaraphites) confusus] LM87
    |--S. hirtipes Macleay 1864 [incl. S. assimilis Sloane 1894, S. crenaticollis Macleay 1864] LM87
    |--S. humeralis Castelnau 1867 LM87
    |--S. laticollis Macleay 1866 LM87
    |    |--S. l. laticollis LM87
    |    `--S. l. gigas Castlenau 1867 LM87
    |--S. lenaeus Westwood 1842 LM87
    |    |--S. l. lenaeus LM87
    |    |--S. l. latipennis Macleay 1863 LM87
    |    `--S. l. pacificus Sloane 1888 LM87
    |--S. lucidus Chaudoir 1863 LM87
    |--S. martinii Castelnau 1867 LM87
    `--S. silenus (Westwood 1842) (see below for synonymy) LM87

Scaraphites rotundipennis (Dejean 1825) [=Scarites rotundipennis; incl. Scaraphites insulanus Sloane 1888, Scaraphites intermedius Macleay 1865, Scarites (*Scaraphites) macleaii Westwood 1842] LM87

Scaraphites silenus (Westwood 1842) [=Scarites silenus; incl. Scarites bacchus Westwood 1842, Scaraphites heros Castelnau 1867, Scaraphites mastersi Macleay 1869] LM87

*Type species of generic name indicated

REFERENCES

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.