Pupoides

Pupoides nitidulus, copyright Emmanuel Tardy.


Belongs within: Pupilloidea.

Pupoides is a genus of very small snails found in tropical or subtropical regions, characterised by the presence of a single tooth near the top of the aperture (Stanisic et al. 2010).

Characters (from Stanisic et al. 2010): Shell very small, elongately conical, sinistral or dextral, smooth; apeture ovate with a single upper parietal denticle; outer lip strongly reflected; umbilicus a narrow chink. Colour waxy brown.

<==Pupoides Pfeiffer 1854 [incl. Themapupa Iredale 1930; Pupoididae, Pupoidinae] KKW12
    |--*P. (Pupoides) nitidulus (Pfeiffer 1839) [=Bulimus nitidulus] SS10
    `--P. (Ischnopupoides Pilsbry 1926) G88
         `--P. (I.) antiquus Yu & Wang 1977 G88

Pupoides incertae sedis:
  P. albilabris BP90
  P. beltianus (Tate 1894) W08 (see below for synonymy)
  P. coenopictus H84
  P. contrarius (Smith 1894) [=Pupa contraria] S88
  P. eremicolus (Tate 1894) [=Pupa eremicola, Themapupa eremicola] S88
  P. hordacea PF15
  P. ischnus (Tate 1894) [=Pupa ischa, Themapupa ischna; incl. T. i. latior Iredale 1937] S88
  P. lepidulus (Adams & Angas 1864) W08
  P. marginata PF15
  P. modicus BP90
  P. myoporinae S88
  P. pacificus (Pfeiffer 1846) (see below for synonymy) KKW12
  P. sinistralis (Pilsbry 1921) [=P. pacificus var. sinistralis, Themapupa sinistralis] KKW12

Pupoides beltianus (Tate 1894) W08 [=Pupa beltiana S88, Pupoides contrarius beltianus S88, Themapupa beltiana S88]

Pupoides pacificus (Pfeiffer 1846) [=Pupa pacifica, Bulimus pacificus, Themapupa pacifica; incl. T. anapacifica Iredale 1939, Pupoides (Themapupa) anapacifica, T. comperta Iredale 1940, T. dirupta Iredale 1939] KKW12

*Type species of generic name indicated

REFERENCES

[BP90] Burch, J. B., & T. A. Pearce. 1990. Terrestrial Gastropoda. In: Dindal, D. L. (ed.) Soil Biology Guide pp. 201–309. John Wiley & Sones: New York.

[G88] Gu H.-L. 1988. Fossils of land snails from Xinyu Formation (early Tertiary) of Qingjiang Basin, Jiangxi Province. Acta Palaeontologica Sinica 27 (1): 111–124.

[H84] Heller, J. 1984. Shell colours of desert landsnails. Malacologia 25 (2): 355–359.

[KKW12] Köhler, F., V. Kessner & C. Whisson. 2012. New records of non-marine, non-camaenid gastropods (Mollusca: Gastropoda) from islands off the Kimberley coast, Western Australia. Records of the Western Australian Museum 27 (1): 21–39.

[PF15] Pilsbry, H. A., & J. H. Ferriss. 1915. Mollusca of the southwestern states, VII: the Dragoon, Mule, Santa Rita, Baboquivari, and Tucson Ranges, Arizona. Proceedings of the Academy of Natural Sciences of Philadelphia 67 (2): 363–418, pls 8–15.

[S88] Solem, A. 1988. Non-camaenid land snails of the Kimberley and Northern Territory, Australia. I. Systematics, affinities and ranges. Invertebrate Taxonomy 4: 455–604.

[SS10] Stanisic, J., M. Shea, D. Potter & O. Griffiths. 2010. Australian Land Snails vol. 1. A. field guide to eastern Australian species. Bioculture Press: Mauritius.

[W08] Wilson, B. 2008. Terrestrial gastropods of Faure Island, Shark Bay, Western Australia. Records of the Western Australian Museum Supplement 75: 21–24.

Sphaerospira

Sphaerospira mortenseni, copyright Claude & Amandine Evanno.


Belongs within: Helicoidei.

Sphaerospira is a genus of large snails found in southeastern Queensland and northeastern New South Wales characterised by a black mantle (Stanisic et al. 2010).

Characters (from Stanisic et al. 2010): Shell large to very large, globose to conically globose, with a low to elevated spire; whorls rounded, sutures impressed; protoconch with weak, crowded radial ridges, teleoconch smooth, microsculpture of very fine periostracal radial ridges; lip thickened and reflected, blue-black; umbilicus closed. Colour tawny yellow with numerous black spiral bands and a dark suffusion behind the lip, to all brown with bands visible only on early whorls; rarely all-yellow. Animal with black mantle.

<==Sphaerospira Mörch 1867 [incl. Varohadra Iredale 1933] SS10
    |--*S. fraseri (Griffith & Pidgeon 1833) (see below for synonymy) SS10
    |--S. arthuriana (Cox 1873) [incl. Varohadra halleyae Iredale 1937] SS10
    |--S. bencarlessi Stanisic & Potter in Stanisic, Shea et al. 2010 SS10
    |--S. blomfieldi (Cox 1864) (see below for synonymy) SS10
    |--S. informis Mousson 1869 [incl. S. informis dietrichae Iredale 1937, S. informis fringilla Iredale 1937] SS10
    |--S. mortenseni (Iredale 1929) SS10
    |--S. mossmani (Brazier 1875) [=Helix (Hadra) mossmani] SS10
    |--S. oconnellensis (Cox 1871) (see below for synonymy) SS10
    |--S. rockhamptonensis (Cox 1873) (see below for synonymy) SS10
    `--S. sidneyi (Iredale 1933) [=Bentosites blomfieldi sidneyi] SS10

Sphaerospira blomfieldi (Cox 1864) [incl. Thersites concors Fulton 1904, S. blomfieldi warroensis Hedley & Musson 1892] SS10

*Sphaerospira fraseri (Griffith & Pidgeon 1833) [=Helix fraseri; incl. H. capucinus Férussac 1835, H. coarctata Férussac 1839, Sphaerospira fraseri feriarum Iredale 1937, H. fraseri flavescens Hedley 1888 non H. flavescens Pfeiffer 1848, Bentosites blomfieldi latior Iredale 1933, Sphaerospira parallela Iredale 1937, S. fraseri permuta Iredale 1937] SS10

Sphaerospira oconnellensis (Cox 1871) [incl. S. oconnellensis caroli Iredale 1933, S. oconnellensis jacksoni Iredale 1933] SS10

Sphaerospira rockhamptonensis (Cox 1873) [incl. S rockhamptonensis decreta Iredale 1937, Helix moresbyi Angas 1876, H. planibasis Brazier 1881] SS10

*Type species of generic name indicated

REFERENCES

[SS10] Stanisic, J., M. Shea, D. Potter & O. Griffiths. 2010. Australian Land Snails vol. 1. A. field guide to eastern Australian species. Bioculture Press: Mauritius.

Austrochloritis

Austrochloritis sp., copyright Catching the Eye.


Belongs within: Helicoidei.

Austrochloritis is a genus of snails found in southeastern Australia with a teleoconch sculpture including periostracal setae and wavy ridgelets (Stanisic et al. 2010).

Characters (from Stanisic et al. 2010): Shell small to large, subdiscoidal to turbinate with a flat to elevated spire; whorls rounded, shouldered to angulate or keeled, sutures moderately to strongly impressed; protoconch sculpture of scattered pustules and vague rugose radial ridges, teleoconch sculpture of coarse radial ridges to smooth, with curved or straight, crowded to widely spaced periostracal setae, microsculpture of wavy periostracal ridgelets; lip reflected, often with a sulcus and swelling immediately behind; umbilicus very narrow to moderately open, V-shaped. Colour dark to light brown, lip white to brown. Animal dark grey or black to orange-brown.

<==Austrochloritis Pilsbry 1890 SS10
    |--*A. porteri (Cox 1866) [=Helix porteri] SS10
    |--A. abbotti Stanisic in Stanisic, Shea et al. 2010 SS10
    |--A. abrotonus Shea & Griffiths in Stanisic, Shea et al. 2010 SS10
    |--A. ascensa Iredale 1943 SS10
    |--A. ballengarra Stanisic in Stanisic, Shea et al. 2010 SS10
    |--A. bellengerensis (Cox 1871) [incl. Helix bellingenensis Brazier 1871] SS10
    |--A. brevipila (Pfeiffer 1849) [incl. Chloritis novocambrica Gude 1906] SS10
    |--A. congarinni Stanisic & Potter in Stanisic, Shea et al. 2010 SS10
    |--A. cunninghamiana Stanisic & Potter in Stanisic, Shea et al. 2010 SS10
    |--A. disjuncta (Gude 1906) SS10
    |--A. illawarra Shea & Griffiths in Stanisic, Shea et al. 2010 SS10
    |--A. iluka Stanisic & Potter in Stanisic, Shea et al. 2010 SS10
    |--A. kanangra Shea & Griffiths in Stanisic, Shea et al. 2010 SS10
    |--A. kaputarensis Stanisic in Stanisic, Shea et al. 2010 SS10
    |--A. kiparra Stanisic in Stanisic, Shea et al. 2010 SS10
    |--A. kosciuszkoensis Shea & Griffiths in Stanisic, Shea et al. 2010 SS10
    |--A. laurieton Shea & Griffiths in Stanisic, Shea et al. 2010 SS10
    |--A. liverpoolensis Shea & Griffiths in Stanisic, Shea et al. 2010 SS10
    |--A. marksandersi Stanisic in Stanisic, Shea et al. 2010 SS10
    |--A. metuenda Iredale 1938 SS10
    |--A. monita Shea & Griffiths in Stanisic, Shea et al. 2010 SS10
    |--A. nambucca Iredale 1943 SS10
    |--A. niangala Shea & Griffiths in Stanisic, Shea et al. 2010 SS10
    |--A. nundinalis Iredale 1943 SS10
    |--A. paucisetosa Stanisic in Stanisic, Shea et al. 2010 SS10
    |--A. seaviewensis Stanisic & Potter in Stanisic, Shea et al. 2010 SS10
    |--A. speculoris Shea & Griffiths in Stanisic, Shea et al. 2010 SS10
    |--A. stanisici Shea & Griffiths in Stanisic, Shea et al. 2010 SS10
    |--A. sydneyensis Shea & Griffiths in Stanisic, Shea et al. 2010 SS10
    |--A. timorensis Stanisic in Stanisic, Shea et al. 2010 SS10
    |--A. toonumbar Shea & Griffiths in Stanisic, Shea et al. 2010 SS10
    |--A. warrumbunglensis Shea & Griffiths in Stanisic, Shea et al. 2010 SS10
    `--A. wollemiensis Shea & Griffiths in Stanisic, Shea et al. 2010 SS10

*Type species of generic name indicated

REFERENCES

[SS10] Stanisic, J., M. Shea, D. Potter & O. Griffiths. 2010. Australian Land Snails vol. 1. A. field guide to eastern Australian species. Bioculture Press: Mauritius.

Steorra

Steorra bala, copyright Claude & Amandine Evanno.


Belongs within: Helicoidei.

Steorra is a genus of large snails with a banded or monochrome shell found in northeastern Queensland (Stanisic et al. 2010).

Characters (from Stanisic et al. 2010): Shell large to very large, solid to fairly thin, turbinately globose to trochoidally globose with a low to elevated conical spire; whorls rounded to subangulate, sutures impressed; protoconch with weak radial ridges, teleoconch smooth, microsculpture of crowded pustules or periostracal scales on the early whorls, and widely spaced to crowded periostracal microradial ridges on the later whorls; lip reflected, occasionally thickened; umbilicus narrowly open to closed. Colour tawny yellow with numerous dark brown spiral bands, dark brown suffusion behind lip and dark brown circum-umbilical patch, to all-over monochrome dark reddish brown; lip reflected, dark brown.

<==Steorra Stanisic in Stanisic, Shea et al. 2010 SS10
    |--*S. mourilyani (Brazier 1875) (see below for synonymy) SS10
    |--S. bala (Brazier 1878) [=Helix bala] SS10
    |--S. bellaria (Iredlae 1937) [=Gnarosophia bellaria] SS10
    |--S. cookensis (Brazier 1875) (see below for synonymy) SS10
    |--S. crookshanksi Stanisic in Stanisic, Shea et al. 2010 SS10
    |--S. estherlilleyae Stanisic in Stanisic, Shea et al. 2010 SS10
    |--S. jimfergusoni Stanisic & Potter in Stanisic, Shea et al. 2010 SS10
    |--S. mazee (Brazier 1878) [=Helix mazee] SS10
    |--S. mulgravensis (Brazier 1872) (see below for synonymy) SS10
    |--S. rawnesleyi (Brazier 1873) [=Helix (Camaena) rawnesleyi] SS10
    |--S. tomsoni (Brazier 1876) [=Helix (Hadra) tomsoni] SS10
    `--S. worsfoldi Stanisic in Stanisic, Shea et al. 2010 SS10

Steorra cookensis (Brazier 1875) [=Helix (Hadra) cookensis; incl. H. bebias Brazier 1878, Pallidelix spurlingi Iredale 1937] SS10

*Steorra mourilyani (Brazier 1875) [=Helix (Hadra) mourilyani; incl. Varohadra banfieldi Iredale 1937, H. (H.) hanni Brazier 1876, H. (H.) hilli Brazier 1875, Varohadra hubbardi Iredale 1937, H. (H.) johnstonei Brazier 1875, V. russelli Iredale 1937] SS10

Steorra mulgravensis (Brazier 1872) [=Helix mulgravensis; incl. Gnarosophia palmensis austrina Iredale 1933, H. palmensis meridionalis Brazier 1881, G. mitifica Iredale 1933, H. incei multifasciata Cox 1864 non H. multifasciata Weinland & Anton 1857, H. (Hadra) palmensis Brazier 1876] SS10

*Type species of generic name indicated

REFERENCES

[SS10] Stanisic, J., M. Shea, D. Potter & O. Griffiths. 2010. Australian Land Snails vol. 1. A. field guide to eastern Australian species. Bioculture Press: Mauritius.

Figuladra

Pale banded snail Figuladra mattea, from the Queensland Museum.


Belongs within: Helicoidei.

Figuladra is a genus of snails found in southeastern Queensland possessing a globose, almost smooth shell and red mantle (Stanisic et al. 2010).

Characters (from Stanisic et al. 2010): Shell large to very large, loosely to tightly coiled, depressedly globose to trochoidally globose with a dome-shaped spire; whorls rounded and often laterally compressed, sutures impressed; protoconch with weak, crowded radial ridges, teleoconch smooth, microsculpture of extremely fine, periostracal microradial threads; lip reflected; umbilicus narrowly open to almost closed. Colour yellowish with few to numerous brown spiral bands to monochrome yellow; lip white, often with dark brown suffusion behind it and extending to umbilical area. Animal pinkish orange with red mantle.

<==Figuladra Iredale 1933 SS10
    |--F. incei (Philippi 1846) SS10
    |    |--F. i. incei (see below for synonymy) SS10
    |    |--*F. i. curtisiana (Pfeiffer 1864) (see below for synonymy) SS10
    |    `--F. i. lessoni (Pfeiffer 1846) (see below for synonymy) SS10
    |--F. appendiculata (Reeve 1854) [incl. Helix thatcheri Cox 1870] SS10
    |--F. barneyae Stanisic in Stanisic, Shea et al. 2010 SS10
    |--F. bayensis (Brazier 1875) [=Helix (Hadra) bayensis] SS10
    |--F. challisi (Cox 1873) SS10
    |--F. mattea (Iredale 1933) SS10
    |--F. muirorum Stanisic in Stanisic, Shea et al. 2010 SS10
    |--F. narelleae Stanisic & Potter in Stanisic, Shea et al. 2010 SS10
    |--F. pallida (Hedley & Musson 1892) [=Helix rockhamptonensis pallida, Hadra rockhamptonensis pallida] SS10
    |--F. reducta (Iredale 1937) [=Varohadra bayensis reducta] SS10
    `--F. volgiola (Iredale 1933) [incl. Helix andersoni Cox 1872 non Blanford 1869] SS10

*Figuladra incei curtisiana (Pfeiffer 1864) [=Helix curtisiana; incl. Varohadra curtisiana exedra Iredale 1933, H. (Hadra) parsoni Cox 1872, H. seminigra Morelet 1864] SS10

Figuladra incei incei (Philippi 1846) [incl. Helix (Camaena) praetermissi; incl. Varohadra keppelensis degener Iredale 1937, V. keppelensis Iredale 1937] SS10

Figuladra incei lessoni (Pfeiffer 1846) [incl. Helix (Camaena) aureedensis Brazier 1872, Varohadra bernhardi Iredale 1933, H. (Hadra) yeppoonensis Beddome 1897] SS10

*Type species of generic name indicated

REFERENCES

[SS10] Stanisic, J., M. Shea, D. Potter & O. Griffiths. 2010. Australian Land Snails vol. 1. A. field guide to eastern Australian species. Bioculture Press: Mauritius.

Achatinina

Opeas pyrgula, copyright H. Zell.


Belongs within: Heterobranchia.
Contains: Streptaxidae, Achatininae, Stenogyrinae, Subulininae.

The Achatinina are a clade of snails united by molecular phylogenetic analysis, members of which have genitalia with a greatly reduced or absent stimulator. The carnivorous Streptaxoidea, characterised by a radula with lanceolate teeth, form the sister group of the remaining species in the Achatinoidea.

Achatinina BR17
    |--Streptaxoidea BR17
    |    |--Streptaxidae BR05
    |    `--Diaphera Albers 1850 [Diapheridae] BR17
    |         `--*D. cumingiana (Pfeiffer 1845) [=Cylindrella cumingiana] BR17
    `--Achatinoidea [Achatinacea] BR05
         |--Micractaeon Verdcourt 1993 [Micractaeonidae] BR05
         |    `--*M. kakamegaensis Verdcourt 1993 BR17
         |--Aillya Odhner 1927 BR05 [Aillyida, Aillyidae BR17, Aillyoidea]
         |    `--*A. camerunensis Odhner 1927 BR17
         |--Ferussaciidae [Ferrusacidae, Ferussaciinae] SS10
         |    |--Coilostele TTE93
         |    |--Ferussacia Risso 1826 [=Ferrussacia (l. c.)] BR05
         |    |    `--*F. gronoviana Risso 1826 BR17
         |    `--Geostilbia Crosse 1867 SS10
         |         `--G. gundlachi (Pfeiffer 1850) (see below for synonymy) SS10
         `--Achatinidae [Achatinae, Achatinida, Subulinidae, Subulinoidea] SS10
              |  i. s.: Leptocala mollicella FGN07
              |         Oreohomorus P02
              |         Subulona FGN07
              |           |--S. bumbaensis (Spence 1923) [=Homorus (Subulona) bumbaensis] PB27
              |           `--S. decollata (Morelet 1873) FGN07
              |         Striosubulina FGN07
              |         Dictyoglessula FGN07
              |         Oleata FGN07
              |         Pileata FGN07
              |         Eremopeas Pilsbry 1906 SS10
              |           |--*E. interioris (Tate 1894) SS10 (see below for synonymy)
              |           `--E. tuckeri (Pfeiffer 1846) SS10 (see below for synonymy)
              |         Prosopeas achatinaceum (Pfeiffer 1846) (see below for synonymy) S88
              |         Allopeas Baker 1935 SS10
              |           |--*A. gracile (Hutton 1834) SS10 (see below for synonymy)
              |           `--A. clavulinus (Potiez & Michaud 1838) SS10
              |         Paropeas Pilsbry 1906 SS10
              |           |--*P. acutissimum (Mousson 1857) [=Bulimus acutissimus] SS10
              |           `--P. achatinaceum (Pfeiffer 1846) SS10
              |         Eutomopeas BR17
              |         Leptinaria lamellata BR17, A66 [=Tornatellina (Leptinaria) lamellata A66]
              |         Archachatina PB27
              |         Columna flammea PB27, D66
              |         Pseudachatina PB27
              |         Atopocochlis PB27
              |         Pseudotrochus jansseni Dupuis & Putzeys 1922 PB27
              |         Thomea PB27
              |         Krapfiella PB27
              |         Mabiliella PB27
              |         Metachatina PB27
              |         Cochlitoma PB27
              |         Hypolysia PB27
              |         Xerocerastus PB27
              |         Zootecus PB27
              |         Chilonopsis PB27
              |--Achatininae BR05
              |--Stenogyrinae BR05
              |--Subulininae BR17
              |--Trichodina Ancey 1888 BR17 [=Petriola Dall 1905 BR17; Petriolinae BR05]
              |    `--*T. marmorea (Reeve 1850) [=Achatina marmorea, *Petriola marmorea] BR17
              |--Rumina Risso 1826 [Ruminidae, Rumininae] BR05
              |    `--*R. decollata (Linnaeus 1758) [=Helix decollata] BR17
              |--Coeliaxis Adams & Angas 1865 [=Caeliaxis (l. c.); Caeliaxidae, Coeliaxinae] BR05
              |    `--*C. layardi (Adams & Angas 1865) [=Subulina layardi] BR17
              |--Pyrgina Greef 1882 BR05 [Pyrgininae BR17]
              |    `--*P. umbilicata Greef 1882 BR17
              |--Cryptelasmus Pilsbry 1907 BR05 [Cryptelasminae BR17]
              |    `--*C. canteroianus (Pfeiffer 1857) [=Balea canteroiana] BR17
              |--Thyrophorella Greef 1882 BR05 [Thyrophorellidae, Thyrophorellinae BR17, Thyrophorelloidea]
              |    `--*T. thomensis Greef 1882 BR17
              |--Opeas Albers 1850 [Opeatinae] BR05
              |    |--*O. goodallii (Miller 1822) BR17 (see below for synonymy)
              |    |--O. minutum E99
              |    |--O. pyrgula Schmacker & Boettger 1891 S88
              |    |--O. thomasi Connolly 1923 PB27
              |    `--O. venustum PB27
              |--Cecilioides Férussac 1814 BR05 (see below for synonymy)
              |    |--*C. acicula (Müller 1774) [=Buccinum acicula] BR17
              |    |--C. eulima CC06
              |    |--C. gundlachi [incl. Caecilioides spencei Dupuis & Putzeys 1922] PB27
              |    `--C. stuhlmanni PB27
              `--Glessulinae [Glessulidae] BR17
                   |--Tortaxis BR17
                   |--Glessula Martens 1860 BR05
                   |    `--*G. gemma (Reeve 1850) [=Achatina gemma] BR17
                   `--Rishetia Godwin-Austen 1920 BR17, BR05 [Rishetiinae]
                        `--*R. tenuispira (Benson 1836) [=Achatina tenuispira] BR17

*Allopeas gracile (Hutton 1834) SS10 [=Bulimus gracilis SS10, Lamellaxis (Allopeas) gracilis S88, Opeas gracile S88, Stenogyra gracilis TC89; incl. O. ardelio Iredale 1941 SS10, O. oparanum (Pfeiffer 1846) S88]

Cecilioides Férussac 1814 BR05 [=Caecilianella Bourguignat 1856 BR05, Caecilioides BR05; Caeciliae, Caecilianellea, Cecilioididae, Cecilioidinae BR17]

*Eremopeas interioris (Tate 1894) SS10 [=Stenogyra interioris SS10, Pseudopeas (Eremopeas) interioris S88]

Eremopeas tuckeri (Pfeiffer 1846) SS10 [=Bulimus tuckeri S88, Prosopeas (Eremopeas) tuckeri S88; incl. B. walli Cox 1864 SS10]

Geostilbia gundlachi (Pfeiffer 1850) [incl. Cecilioides aperta (Swainson 1840) (n. n.), *G. caledonica Crosse 1867] SS10

*Opeas goodallii (Miller 1822) BR17 [=Bulimus goodallii BR17, Helix goodalli non Ferussac 1821 S88; incl. B. pumilus Pfeiffer 1840 S88, Opeas pumilum S88]

Prosopeas achatinaceum (Pfeiffer 1846) [=Bulimus achatinaceus; incl. Achatina javanica Reeve 1849, Opeas javanicum] S88

*Type species of generic name indicated

REFERENCES

[A66] Adams, H. 1866. List of land and freshwater shells collected by Mr. E. Bartlett on the Upper Amazons, and on the River Ucayali, eastern Peru, with descriptions of new species. Proceedings of the Zoological Society of London 1866: 440–445.

[BR05] Bouchet, P., & J.-P. Rocroi. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1–2): 1–397.

[BR17] Bouchet, P., J.-P. Rocroi, B. Hausdorf, A. Kaim, Y. Kano, A. Nützel, P. Parkhaev, M. Schrödl & E. E. Strong. 2017. Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia 61 (1–2): 1–526.

[CC06] Cameron, R. A. D., L. M. Cook, G. A. Goodfriend & M. B. Seddon. 2006. Fossil land snails of Porto Santo, Madeiran archipelago: change and stasis in Pleistocene to Recent times. Malacologia 49 (1): 25–59.

[D66] Dohrn, H. 1866. Synopsis of the birds of Ilha do Principe, with some remarks on their habits and descriptions of new species. Proceedings of the Zoological Society of London 1866: 324–332.

[E99] Esu, D. 1999. Contribution to the knowledge of Neogene climate changes in western and central Europe by means of non-marine molluscs. In: J. Agustí, L. Rook & P. Andrews (eds) The Evolution of Neogene Terrestrial Ecosystems in Europe pp. 328–354. Cambridge University Press.

[FGN07] Fontaine, B., O. Gargominy & E. Neubert. 2007. Land snail diversity of the savanna/forest mosaic in Lopé National Park, Gabon. Malacologia 49 (2): 313–338.

[P02] Pickford, M. 2002. Early Miocene grassland ecosystem at Bukwa, Mount Elgon, Uganda. Comptes Rendus Palevol 1: 213–219.

[PB27] Pilsbry, H. A., & J. Bequaert. 1927. The aquatic mollusks of the Belgian Congo, with a geographical and ecological account of Congo malacology. Bulletin of the American Museum of Natural History 53 (2): 69–602, pls 10–77.

[S88] Solem, A. 1988. Non-camaenid land snails of the Kimberley and Northern Territory, Australia. I. Systematics, affinities and ranges. Invertebrate Taxonomy 4: 455–604.

[SS10] Stanisic, J., M. Shea, D. Potter & O. Griffiths. 2010. Australian Land Snails vol. 1. A. field guide to eastern Australian species. Bioculture Press: Mauritius.

[TC89] Tapparone Canefri, T. 1889. Viaggio di Leonardo Fea in Birmania e regioni vicine. XVIII.—Molluschi terrestri e d’acqua dolce. Annali del Museo Civico di Storia Naturale di Genova, Serie 2a, 7: 295–359.

[TTE93] Tracey, S., J. A. Todd & D. H. Erwin. 1993. Mollusca: Gastropoda. In: Benton, M. J. (ed.) The Fossil Record 2 pp. 131–167. Chapman & Hall: London.

Succineoidei

Amber snail Succinea putris, copyright Charles J. Sharp.


Belongs within: Heterobranchia.

The Succineoidei are a group of terrestrial molluscs whose members have optic tentacles that retract that do not invert, and elasmognathous jaws with lateral winged processes and a greatly enlarged attachment plate (Stanisic et al. 2010). The cosmopolitan Succineidae are thin-shelled, conical snails whereas the Australasian Athoracophoridae are brightly coloured slugs.

Succineoidei [Elasmognatha, Heterurethra, Succineida, Succineiformes, Succineoinei]
    |--Athoracophoridae [Athoracophoracea, Athoracophoroidea, Tracheopulmonata] SS10
    |    |--Aneitella virgata T84
    |    |--Triboniophorus Humbert 1863 SS10
    |    |    `--*T. graeffei Humbert 1863 (see below for synonymy) SS10
    |    |--Vomanus Iredale 1915 P61
    |    |    `--*V. marmoreus (Hutton 1879) [=Konophora marmoreus] P61
    |    |--Amphiconophora Suter 1897 P61
    |    |    |--*A. gigantea (Suter 1909) P61
    |    |    `--A. verrucosa (Simroth 1889) P61
    |    |--Aneitea Gray 1860 [Aneiteadae, Aneiteidae, Aneiteinae] BR05
    |    |    |--*A. macdonaldii Gray 1860 BR17
    |    |    `--A. simrothi T84
    |    |--Athoracophorus Gould 1852 (see below for synonymy) BR05
    |    |    `--*A. bitentaculatus (Quoy & Gaimard 1832) P61 [=Limax bitentaculatus P61, *Janella bitentaculata BR17]
    |    |         |--A. b. bitentaculatus P61
    |    |         `--A. b. rufovenosus Suter 1909 P61
    |    `--Pseudaneitea Cockerell 1891 P61
    |         |--*P. papillata (Hutton 1879) [=Athoracophorus papillatus] P61
    |         |--P. dendyi (Suter 1897) P61
    |         |--P. huttoni (Suter 1909) P61
    |         |--P. martensi (Suter 1909) P61
    |         |--P. schauinslandi (Plate 1897) P61
    |         |--P. simrothi (Suter 1896) P61
    |         `--P. sorenseni Powell 1955 P61
    `--Succineidae [Diplotrema, Succineacea, Succineoidea] BR05
         |  i. s.: Quickia FGN07
         |         Spirancinea Iredale 1945 SS10
         |           `--*S. norfolkensis (Sykes 1900) (see below for synonymy) SS10
         |         Papyrotheca G88
         |         Omalonyx matheroni T84
         |--Catinella Pease 1870 [Catinellinae] BR05
         |    |--*C. rubida Pease 1870 BR17
         |    `--C. avara BP90
         `--Succineinae BR05
              |--Hyalimax Adams & Adams 1855 [Hyalimacidae, Hyalimacinae] BR05
              |    `--*H. perlucidus (Quoy & Gaimard 1832) [=Limax perlucidus] BR17
              |--Austrosuccinea Iredale 1937 [incl. Arborcinea Iredale 1937] S88
              |    |--*A. australis (Férussac 1861) [=Succinea australis] SS10
              |    |--A. macgillivrayi (Cox 1864) [incl. Succinea nortoni Cox 1864] SS10
              |    `--A. strigillata (Adams & Angas 1864) W08
              |--Oxyloma Westerlund 1885 [Oxylomatinae, Oxylominae] BR05
              |    |--*O. dunkeri (Pfeiffer 1865) [=Succinea dunkeri] BR17
              |    |--O. elegans (Risso 1826) CP06
              |    |--O. pfeifferi (Rossmässler 1835) V09, CP06
              |    `--O. retusa BP90
              `--Succinea Draparnaud 1801 SS10 [incl. Papusuccinea Iredale 1941 S88; Succineae]
                   |--*S. putris (Linnaeus 1758) SS10 [=Helix putris SS10, Bulimus putris PB27]
                   |--S. aperta Lea 1838 F27
                   |--‘Papusuccinea’ archeyi (Powell 1933) P61
                   |--S. avara B59
                   |--S. baumanni Sturany 1894 V07
                   |--S. boissyi (Deshayes 1863) TTE93
                   |--S. caducca Mighels 1845 J49
                   |--S. chudeaui [incl. S. chudeaui var. leyneni Dupuis & Putzeys 1923] PB27
                   |--S. congoensis PB27
                   |--S. coxi Finlay 1927 [=S. aperta Cox 1868 non Lea 1838] F27
                   |--S. grosvenori B59
                   |--S. ovalis BP90
                   |--S. propinqua T84
                   |--S. rotundata Gould 1845 [incl. S. patula Mighels 1845 non Lamarck in de Blainville 1827] J49
                   |--S. strubelli [=*Papusuccinea strubelli] P61
                   `--S. wilsoni Lea 1864 BF89

Athoracophorus Gould 1852 [=Janella Gray 1850 non Grateloup 1838; Athoracophorinae, Janellidae, Janellinae] BR05

*Succinea norfolkensis (Sykes 1900) [=Succinea norfolkensis; incl. Su. humerosa Preston 1913, Su. nepeanensis Preston 1913] SS10

*Triboniophorus graeffei Humbert 1863 [incl. T. brisbanensis Pfeiffer 1900, T. krefftii Keferstein 1865, Aneitea graeffei rosea Hedley 1893, T. schuttei Keferstein 1865] SS10

*Type species of generic name indicated

REFERENCES

[BR05] Bouchet, P., & J.-P. Rocroi. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1–2): 1–397.

[BR17] Bouchet, P., J.-P. Rocroi, B. Hausdorf, A. Kaim, Y. Kano, A. Nützel, P. Parkhaev, M. Schrödl & E. E. Strong. 2017. Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia 61 (1–2): 1–526.

[B59] Branson, B. A. 1959. An ecologic observation on Succinea avara. Nautilus 72 (4): 145–146.

[BP90] Burch, J. B., & T. A. Pearce. 1990. Terrestrial Gastropoda. In: Dindal, D. L. (ed.) Soil Biology Guide pp. 201–309. John Wiley & Sones: New York.

[BF89] Burnham, B. H., & P. E. Fell. 1989. Distribution of Melampus bidentatus (Say) and Succinea wilsoni (Lea) within a tidal marsh in eastern Connecticut. Nautilus 103 (3): 109–112.

[CP06] Cameron, R. A. D., B. M. Pokryszko, A. Riedel & A. Wiktor. 2006. Checklists, systematics and the CLECOM initiative: an alternative view from Europe. Malacologia 49 (1): 225–230.

[F27] Finlay, H. J. 1927. New specific names for austral Mollusca. Transactions and Proceedings of the New Zealand Institute 57: 488–533.

[FGN07] Fontaine, B., O. Gargominy & E. Neubert. 2007. Land snail diversity of the savanna/forest mosaic in Lopé National Park, Gabon. Malacologia 49 (2): 313–338.

[G88] Gray, J. 1988. Evolution of the freshwater ecosystem: the fossil record. Palaeogeography, Palaeoclimatology, Palaeoecology 62: 1–214.

[J49] Johnson, R. I. 1949. Jesse Wedgwood Mighels with a bibliography and a catalogue of his species. Occasional Papers on Mollusks 1 (14): 213–231.

[PB27] Pilsbry, H. A., & J. Bequaert. 1927. The aquatic mollusks of the Belgian Congo, with a geographical and ecological account of Congo malacology. Bulletin of the American Museum of Natural History 53 (2): 69–602, pls 10–77.

[P61] Powell, A. W. B. 1961. Shells of New Zealand: An illustrated handbook 4th ed. Whitcombe and Tombs Limited: Christchurch.

[S88] Solem, A. 1988. Non-camaenid land snails of the Kimberley and Northern Territory, Australia. I. Systematics, affinities and ranges. Invertebrate Taxonomy 4: 455–604.

[SS10] Stanisic, J., M. Shea, D. Potter & O. Griffiths. 2010. Australian Land Snails vol. 1. A. field guide to eastern Australian species. Bioculture Press: Mauritius.

[T84] Tillier, S. 1984. Patterns of digestive tract morphology in the limacisation of helicarionid, succineid and athoracophorid snails and slugs (Mollusca: Pulmonata). Malacologia 25 (1): 173–192.

[TTE93] Tracey, S., J. A. Todd & D. H. Erwin. 1993. Mollusca: Gastropoda. In: Benton, M. J. (ed.) The Fossil Record 2 pp. 131–167. Chapman & Hall: London.

[V07] Verdcourt, B. 2007. Miscellaneous notes on tropical African non-marine Mollusca. Annales Historico-Naturales Musei Nationalis Hungarici 99: 189–197.

[V09] Verdcourt, B. (ed.) 2009. Additions to the wild fauna and flora of the Royal Botanic Gardens, Kew. XXVI. Miscellaneous records. Kew Bulletin 64 (1): 183–194.

[W08] Wilson, B. 2008. Terrestrial gastropods of Faure Island, Shark Bay, Western Australia. Records of the Western Australian Museum Supplement 75: 21–24.