Pseudocaeciliinae

Heterocaecilius lachlani, from Insects of Tasmania.


Belongs within: Pseudocaecilioidea.

The Pseudocaeciliinae are a group of barklice with two-segmented tarsi, known predominantly from south-east Asia and the Pacific (New & Lienhard 2007).

<==Pseudocaeciliinae [Meplerinae, Pseudoscottiinae] NL07
    |--*Allopsocus marginalis Banks 1920 NL07
    |--Scottiella NL07
    |--Platyocaecilius Li 2002 L02b
    |    `--*P. parallelivenius Li 2002 L02b
    |--Fashengocaecilius Lienhard 2003 L03 [=Calocaecilius Li 2002 non Mockford 1974 L02b]
    |    `--*F. biaduncus (Li 1992) [=Allocaecilius biaduncus, *Calocaecilius biaduncus] L03
    |--Phyllocaecilius Li 2002 L02b
    |    `--*P. atrichus Li 2002 L02b
    |--Chorocaecilius Li 2000 L02a
    |    |--C. brunellus (Tillyard 1923) (see below for synonymy) SN08
    |    `--C. decemdentatus Li 2000 L02a
    |--Phallocaecilius Lee & Thornton 1967 L02b
    |    |--*P. hirsutus (Thornton 1961) [=Pseudocaecilius hirsutus] NL07
    |    `--P. sentosus Li 1999 L02a
    |--Thelocaecilius Li 2002 L02b
    |    |--*T. mecokeratus Li 2002 L02b
    |    `--T. papillatus (Li 1995) [=Heterocaecilius papillatus] L02b
    |--Neocaecilius Li 2000 L02a
    |    |--*N. campanula [=Heterocaecilius campanula] NL07
    |    |--N. granulatus (Lee & New 1992) NL07
    |    |--N. mutatus (Lee & New 1992) NL07
    |    `--N. qianshanensis Li 2000 L02a
    |--Mesocaecilius Okamoto 1910 NL07, L02a [Mesocaeciliinae]
    |    |--*M. quadrimaculatus Okamoto 1910 L02a
    |    |--M. bellus Li 2002 L02b
    |    |--M. decorosus Li 2002 L02b
    |    |--M. elegans Li 1997 L02b
    |    |--M. euryopterus Li 2002 L02b
    |    |--M. lepidus Li 2002 L02b
    |    `--M. paraornatus (New 1977) [=Pseudocaecilius paraornatus] L02b
    |--Ophiodopelma Enderlein 1908 L02a
    |    |--*O. ornatipenne Enderlein 1908 NL07, L02a
    |    |--O. anocella Li 2002 L02b
    |    |--O. fluctosa Li 1995 L02a
    |    |--O. glyptocephalum Lienhard 1985 NL07
    |    |--O. hieroglyphicum (Enderlein 1903) NL07
    |    |--O. multipunctatum (Hagen 1859) NL07
    |    |--O. permaculatum Lee & Thornton 1967 NL07
    |    |--O. pictipenna Lee & Thornton 1967 NL07
    |    |--O. polyspila Li 1993 L02b
    |    `--O. semiceps Lee & Thornton 1967 NL07
    |--Levucaecilius Smithers 1995 NL07, S95
    |    |--*L. greenwoodi [=Pseudocaecilius greenwoodi, Orbocaecilius greenwoodi] NL07
    |    |--L. albicrus (Thornton 1981) [=Heterocaecilius albicrus] S95
    |    |--L. apicalis (Thornton 1981) [=Heterocaecilius apicalis] S95
    |    |--L. dardanus (Lee & Thornton 1967) S95 [=Heterocaecilius dardanus NL07, Neocaecilius dardanus NL07]
    |    |--L. hilli Smithers 1995 NL07
    |    |--L. panicus (Lee & Thornton 1967) [=Heterocaecilius panicus] S95
    |    |--L. pictus (Thornton 1981) [=Heterocaecilius pictus] S95
    |    |--L. simplex (Lee & Thornton 1967) NL07, L02a [=Heterocaecilius simplex NL07]
    |    |--L. veitchi [=Heterocaecilius veitchi] S95
    |    `--L. volatus (Lee & Thornton 1967) [=Heterocaecilius volatus] S95
    |--Lobocaecilius Lee & Thornton 1967 L02a
    |    |--*L. cynara Lee & Thornton 1967 L02a
    |    |--L. arcuatus Vaughan, Thornton & New 1989 NL07
    |    |--L. bifurcus Li 1993 L02a
    |    |--L. drepanus New & Thornton 1976 NL07
    |    |--L. ligur Lee & New 1992 NL07
    |    |--L. longifurca Lee & Thornton 1967 NL07
    |    |--L. pahangensis Lee & New 1992 NL07
    |    |--L. perductivirgus Li 2002 L02b
    |    |--L. pilosus Lee & New 1992 NL07
    |    |--L. quadripartitus Li 1995 L02b
    |    |--L. thorntoni Lee & New 1992 NL07
    |    `--L. viridis Lee & New 1992 NL07
    |--Allocaecilius Lee & Thornton 1967 L02a
    |    |--*A. heterothrax Lee & Thornton 1967 L02a
    |    |--A. albidorsualis Li 1993 L02a
    |    |--A. arcuatus Li 2002 L02b
    |    |--A. elongatus Lee & Thornton 1967 NL07
    |    |--A. fusciceps Lee & Thornton 1967 NL07
    |    |--A. gombakensis Lee & New 1992 NL07
    |    |--A. guangdongicus Li 1993 L02a
    |    |--A. maculatus Lee & Thornton 1967 NL07
    |    |--A. selangorensis New & Thornton 1975 NL07
    |    |--A. sinensis Lee & Thornton 1967 L02a
    |    |--A. smithersi Lee & New 1992 NL07
    |    `--A. tenuilongus Li 1995 L02a
    |--Mepleres Enderlein 1926 SN08 [incl. Meniscopsocus Li 1993 L02b, Pseudoscottiella Badonnel 1946 L02b]
    |    |--*M. maeandricus Enderlein 1926 SN08
    |    |--M. cornutus (Lee & New 1992) NL07 [=Pseudoscottiella cornuta L02b]
    |    |--‘Pseudoscottiella’ decolor Badonnel 1955 B55
    |    |--‘Pseudoscottiella’ hyalina Badonnel 1955 B55
    |    |--‘Pseudoscottiella’ immaculata Badonnel 1955 B55
    |    |--M. indicus (Lee & New 1992) NL07
    |    |--M. limbatus (Enderlein 1908) L02a
    |    |--M. longicellus Li 2002 L02b
    |    |--M. longitudinalis Li 2002 [=M. longitudianlis] L03
    |    |--‘Pseudoscottiella’ maculata Broadhead & Richard 1982 L02b
    |    |--*Pseudoscottiella’ megops NL07
    |    |--M. nigroguttatus (Karny 1925) NL07 [=Pseudoscottiella nigraguttata L02b]
    |    |--M. ocellatus Li 2002 L02b
    |    |--M. oresbius Li 2002 L02b
    |    |--M. parvicellus Li 2002 L02b
    |    |--M. pisiformis (Lee & New 1992) NL07
    |    |--M. plenimacularis Li 2002 L02b [=Pseudoscottiella plenimacularis L02a]
    |    |--M. proboscideus Li 2002 L02b
    |    |--M. procurrens Li 2002 L02b
    |    |--M. radialis (Thornton 1984) NL07
    |    |--M. raphalis (Lee & New 1992) NL07
    |    |--M. rotundatus (New 1974) [=Pseudoscottiella rotundata, Meniscopsocus rotundatus] SN08
    |    |--M. similis (Lee & New 1992) NL07
    |    |--M. sinicus (Li 1993) L02b [=*Meniscopsocus sinicus NL07]
    |    |--M. solenocerus (Lee & New 1992) NL07
    |    |--M. suella (Thornton 1984) NL07 [=Pseudoscottiella suella L02b]
    |    |--M. tanei (Smithers 1977) [=Pseudoscottiella tanei, Meniscopsocus tanei] SN08
    |    |--M. transversus (Banks 1937) NL07
    |    |--‘Pseudoscottiella’ tuberculata Badonnel 1955 B55
    |    |--M. ulterior Li 2002 L02b
    |    |--M. unitus Li 2002 L02b
    |    |--M. urbanus (Vaughan, Thornton & New 1991) NL07
    |    |--M. vilus (Lee & New 1992) NL07
    |    |--‘Pseudoscottiella’ yeni New 1974 L02b
    |    `--M. yunnanicus Li 2002 L02b
    |--Pseudocaecilius Enderlein 1903 Y02 [incl. Hageniella Enderlein 1903 (preoc.) L02a]
    |    |--P. citricola (Ashmead 1879) NL07 (see below for synonymy)
    |    |--P. angustus (Enderlein 1903) NL07
    |    |--P. bibulbus Li 1993 L03, L02a [=P. bibalbus (l. c.) L03]
    |    |--P. bicostatus Li 2002 L02b
    |    |--P. ceratocercus Li 2002 L02b
    |    |--P. claggi (Banks 1937) NL07
    |    |--P. clunialis Vaughan, Thornton & Neo 1991 NL07
    |    |--P. cornutus Lee & Thornton 1967 NL07
    |    |--P. danaus Lee & New 1992 NL07
    |    |--P. euryocercus Li 2002 [=P. eurocerus] L03
    |    |--P. formosanus (Banks 1937) L02a
    |    |--P. galactozonalis Li 2002 [=P. galactozonacis] L03
    |    |--P. helicoides Lee & Thornton 1967 NL07
    |    |--P. immaculatus Li 2002 L02b
    |    |--P. inaequalis (Banks 1916) NL07
    |    |--P. innotatus Banks 1937 NL07
    |    |--P. kagoshimensis Y02
    |    |--P. largicellus Li 2002 L02b
    |    |--P. machadoi Badonnel 1955 B55
    |    |--P. monotaeniatus Li 2002 [=P. monotaenitus, P. monotaenius] L03
    |    |--P. nitoris (Banks 1937) NL07
    |    |--P. octomaculatus (Li 1995) [=Heterocaecilius octomaculatus] L02b
    |    |--P. otiosus (Banks 1937) NL07
    |    |--P. papillaris Li 2002 L02b
    |    |--P. plagiozonalis Li 1995 L02a
    |    |--P. productus New & Thornton 1976 NL07
    |    |--P. pusillus (Banks 1931) NL07
    |    |--P. ranus Thornton 1984 NL07
    |    |--P. serratus Lee & New 1992 NL07
    |    |--P. setifer Vaughan, Thornton & New 1991 NL07
    |    |--P. sexdentatus Li 2002 L02b
    |    |--P. tahitiensis (Karny 1926) [=Epipsocus tahitiensis] M93
    |    |--P. tenellus Enderlein 1926 NL07
    |    |--P. testaceus Enderlein 1903 NL07
    |    |--P. undecimimaculatus Li 1995 L02b
    |    |--P. utricularis Li 1999 L02a
    |    |--P. venimaculatus Li 2002 L02b
    |    |--P. wellsae Lee & New 1992 NL07
    |    `--‘Psocus’ zonatus [=*Hageniella zonata] NL07
    `--Heterocaecilius Lee & Thornton 1967 L02a [incl. Orbocaecilius Li 1999 L03, Obrocaecilius (l. c.) L03]
         |--*H. minutus Lee & Thornton 1967 SN08 [=H. minotus NL07]
         |--H. abacutidens Li 2002 L02b
         |--H. analis Vaughan, Thornton & New 1991 NL07
         |--H. anareolatus New 1993 NL07
         |--H. anomalus (Thornton 1961) Y02, L02b [=Orbocaecilius anomolus L02b]
         |--H. argutus (Li 2002) [=Obrocaecilius (l. c. for Orbocaecilius) argutus] L03
         |--H. bicruris (Li 2002) [=Obrocaecilius (l. c. for Orbocaecilius) bicruris] L03
         |--H. bidigitatus Li 1992 L02a [=Orbocaecilius bidigitatus L03]
         |--H. bifarius (Li 2002) [=Obrocaecilius (l. c. for Orbocaecilius) bifarius] L03
         |--H. brachystigmus (Li 2002) [=Obrocaecilius (l. c. for Orbocaecilius) brachystigmus] L03
         |--H. brevifurca Thornton 1984 NL07
         |--H. challengerius Lee & New 1992 NL07
         |--H. circulicellus Li 1995 L02b [=Orbocaecilius circulicellus L03]
         |--H. dasoceratus (Li 2002) [=Obrocaecilius (l. c. for Orbocaecilius) dasoceratus] L03
         |--H. deloneurous Li 2002 L02b
         |--H. dentalis Li 2002 L02b
         |--H. duodecidentus (Li 2002) [=Obrocaecilius (l. c. for Orbocaecilius) duodecidentus] L03
         |--H. fasciatus Lee & Thornton 1967 NL07
         |--H. flaviformis Lee & New 1992 NL07
         |--H. fumosus Thornton 1984 NL07
         |--H. fuscipalpus Lee & Thornton 1967 NL07
         |--H. fuscus Y02
         |--H. gressitti Lee & Thornton 1967 NL07
         |--H. hepaticivenus Li 2002 L02b
         |--H. hoplites Lee & New 1992 NL07
         |--H. huananensis Li 2002 L02b
         |--H. huaxiensis Li 2002 L02b
         |--H. imparilis (Li 2002) [=Obrocaecilius (l. c. for Orbocaecilius) imparilis] L03
         |--H. jinghongicus Li 1995 L02a
         |--H. kobus Lee & Thornton 1967 NL07
         |--H. lachlani (Enderlein 1903) [=Pseudocaecilius lachlani] SN08
         |--H. leei New 1993 NL07
         |--H. liangi Li 1991 L02b
         |--H. loiaoi New 1991 L02a
         |--H. longicornis (Li 2002) [=Obrocaecilius (l. c. for Orbocaecilius) longicornis] L03
         |--H. longipenna Lee & Thornton 1967 NL07
         |--H. longispinus Lee & Thorton 1967 NL07
         |--H. maculans Li 1991 L02b
         |--H. maculifornis (Thornton 1961) L02b [=Pseudocaecilius maculifrons L02a]
         |--H. maculosus Lee & New 1992 NL07
         |--H. marginatus Lee & New 1992 NL07
         |--H. mecotrichus Li 2002 L02b
         |--H. odontothelus Li 2002 L02b
         |--H. orientalis Lee & New 1992 NL07
         |--H. ornatus (Enderlein 1903) [=Pseudocaecilius ornatus] NL07
         |--H. papillosus Vaughan, Thornton & New 1991 NL07
         |--H. paulicellus (Li 1999) NL07, L02a [=*Orbocaecilius paulicellus L02a]
         |--H. platokeratis Li 1993 L02a
         |--H. pseudoanomalus Li 1993 L02b [=Obrocaecilius (l. c. for Orbocaecilius) pseudoanomalus L03]
         |--H. quadrangulus Li 1992 L02b
         |--H. reticulatus Lee & New 1992 NL07
         |--H. rhadnus Thornton 1984 NL07
         |--H. sedecimidentus (Li 2002) [=Obrocaecilius (l. c. for Orbocaecilius) sedecimidentus] L03
         |--H. stenokeratis Li 1993 L02a
         |--H. stiliger Lee & Thornton 1967 NL07
         |--H. taipingensis Garcia Aldrete 1999 NL07
         |--H. tartareus Li 2002 L02b
         |--H. vulturius (Li 2002) [=Obrocaecilius (l. c. for Orbocaecilius) vulturius] L03
         `--H. xihuicus (Li 2002) [=Obrocaecilius (l. c. for Orbocaecilius) xihuicus] L03

Chorocaecilius brunellus (Tillyard 1923) [=Caecilius brunellus, Heterocaecilius brunellus, Pseudocaecilius brunellus; incl. H. diogenes Lee & Thornton 1967, *Chorocaecilius diogenes] SN08

Pseudocaecilius citricola (Ashmead 1879) NL07 [=Psocus citricola M93; incl. Pse. elutus var. africanus Badonnel 1931 L02a, Elipsocus criniger Perkins 1899 M93, Kilauella criniger M93, Pseudocaecilius criniger M93, *Pse. elutus Enderlein 1903 NL07, L02b, Trichopsocus (*Pseudocaecilius) elutus L02a, Trichopsocus indicatus Navás 1934 M93, Pse. medianicostatus Li 1993 L02a, Caecilius pretiosus Banks 1920 M93, Pse. pretiosus M93, Pse. wolcotti Banks 1924 M93]

*Type species of generic name indicated

REFERENCES

[B55] Badonnel, A. 1955. Psocoptères de l'Angola. Diamang Publicações Culturais 26: 11–267.

[L02a] Li F. 2002a. Psocoptera of China vol. 1. Science Press: Beijing.

[L02b] Li F. 2002b. Psocoptera of China vol. 2. Science Press: Beijing.

[L03] Lienhard, C. 2003. Nomenclatural amendments concerning Chinese Psocoptera (Insecta), with remarks on species richness. Revue Suisse de Zoologie 110 (4): 695–721.

[M93] Mockford, E. L. 1993. North American Psocoptera (Insecta). Sandhill Crane Press, Inc.

[NL07] New, T. R., & C. Lienhard. 2007. The Psocoptera of Tropical South-east Asia. Brill: Leiden.

[SN08] Schmidt, E. R., & T. R. New. 2008. The Psocoptera (Insecta) of Tasmania, Australia. Memoirs of Museum Victoria 65: 71–152.

[S95] Smithers, C. N. 1995. Psocoptera (Insecta) of Christmas Island. Invertebrate Taxonomy 9: 529–561.

[Y02] Yoshizawa, K. 2002. Phylogeny and higher classification of suborder Psocomorpha (Insecta: Psocodea: ‘Psocoptera’). Zoological Journal of the Linnean Society 136: 371–400.

Philotarsidae

Female Aaroniella badonelli, copyright Barbara Thurlow.


Belongs within: Peripsocoidea.

The Philotarsidae are a group of barklice best studied in the Australasian region (New & Lienhard 2007).

Characters (from New & Lienhard 2007): Fore wing length ca 2.2–3.8 mm. Lacinial apex bicuspid, outer cusp broad. Tarsi two- or three-segmented; claws with subapical tooth; pulvillus slender. Macropterous or occasionally brachypterous. Fore wing with areola postica free; margin and veins setose; M+Cu usually with one rank of setae; at least some hind wing veins setose; marginal setae of both wings may cross in opposing ranks. Abdomen without ventral eversible vesicles. Male with hypandrium simple, without paired lateral lobes; aedeagus rounded at apex; external parameres not or scarcely protruding beyond tip of aedeagal arch. Female subgenital plate with simple apical lobe; gonapophyses complete, external valve large and setose.

<==Philotarsidae [Philotarsinae]
    |--Abelopsocus Schmidt & New 2008 SN08
    |    |--*A. fenestratus (Schmidt & Thornton 1993) [=Latrobiella fenestrata, Philotarsopsis fenestrata] SN08
    |    |--A. basipunctatus (Thornton, Wong & Smithers 1977) (see below for synonymy) SN08
    |    |--A. cruciclypeus (New & Thornton 1981) (see below for synonymy) SN08
    |    |--A. pardus (Thornton, Wong & Smithers 1977) [=Aaroniella parda, Latrobiella parda, Philotarsopsis parda] SN08
    |    `--A. truganiniae Schmidt & New 2008 SN08
    |--Philotarsus Kolbe 1880 L02
    |    |--*P. picicornis (Fabricius 1793) M93 (see below for synonymy)
    |    |--P. flaviceps (Stephens 1836) Y02, M93 (see below for synonymy)
    |    |--P. kwakiuti Mockford 1951 M93
    |    |--P. parviceps Roesler 1954 G74
    |    |--P. quercicola Y02
    |    |--P. sinensis Li 1997 L02
    |    |--P. zangdaicus Li 2002 L02
    |    `--P. zangxiaoicus Li 2002 L02
    |--Haplophallus Thornton 1959 SN08
    |    |--*H. orientalis Thornton 1959 SN08
    |    |--H. chinensis Li 1995 L02
    |    |--H. comptoni Vaughan, Thornton & New 1989 NL07
    |    |--H. cubitalis New & Thornton 1976 NL07
    |    |--H. leopardina (Williner 1943) [=Philotarsus leopardina] B86
    |    |--H. oblongatus Thornton & Smithers 1977 NL07
    |    |--H. rhabdocladus Li 2002 L02
    |    |--H. separatus Thornton & Smithers 1977 NL07
    |    |--H. sinus Thornton & New 1977 SN08
    |    |--H. subclarus Mockford 2003 M03
    |    |--H. talilus Thornton & New 1977 NL07
    |    `--H. trachopetalus Li 1999 L03, L02 [=H. trachopterus (l. c.) L03]
    |--Philotarsopsis Tillyard 1923 [incl. Latrobiella Thornton 1981] SN08
    |    |--P. guttata (Tillyard 1923) (see below for synonymy) SN08
    |    |--P. alticola (Thornton & Smithers 1977) NL07
    |    |--P. bundoorensis [=*Latrobiella bundoorensis] NL07
    |    |--P. elongata (Thornton & Smithers 1977) NL07
    |    |--P. hellyeri Schmidt & New 2008 SN08
    |    |--P. kaindiensis (Thornton & Smithers 1977) NL07
    |    |--P. manus (Thornton & New 1977) NL07
    |    |--P. paraguttata (Thornton & New 1977) [=Haplophallus paraguttatus, Latrobiella paraguttata] SN08
    |    |--P. pseudosigma (Thornton & Smithers 1977) NL07
    |    |--P. setosa (Thornton & Smithers 1977) NL07
    |    |--P. sigma (Thornton & Smithers 1977) NL07
    |    |--P. sigmoides (Thornton & Smithers 1977) NL07
    |    `--P. stigmata (Thornton & Smithers 1977) NL07
    `--Aaroniella Mockford 1951 SN08
         |--*A. maculosa (Aaron 1883) M93 [=Elipsocus maculosus SN08, Philotarsus maculosus M93]
         |--A. achrysa (Banks 1941) [=Graphocaecilius achrysus] M93
         |--A. antennata Thornton & Smithers 1977 NL07
         |--A. badonneli (Danks 1950) [=Philotarsus badonneli; incl. A. eertmoedi Mockford 1979] M93
         |--A. bakeri Thornton 1981 NL07
         |--A. curtifurca Li 2002 L02
         |--A. flexa Li 2002 L02
         |--A. grandiocula Li 2002 L02
         |--A. gressitti Thornton, Lee & Chui 1972 NL07
         |--A. guttulata (Banks 1916) NL07
         |--A. gyratigra Li 2002 L02
         |--A. kepongensis New & Lee 1992 NL07
         |--A. lobata Vaughan, Thornton & New 1989 NL07
         |--A. lombokensis Thornton 1981 NL07
         |--A. multipunctata Li 1995 L02
         |--A. nebulosa Vaughan, Thornton & New 1991 NL07
         |--A. pardina Li 1999 L02
         |--A. pedunculata Thornton & Smithers 1977 NL07
         |--A. pulchra Thornton 1959 NL07, L02
         |--A. rawlingsi Smithers 1969 SN08
         |--A. spenceri Thornton 1981 NL07
         |--A. sudarmani Vaughan, Thornton & New 1991 NL07
         `--A. ternata Li 1999 L02

Abelopsocus basipunctatus (Thornton, Wong & Smithers 1977) [=Aaroniella basipunctata, Latrobiella basipunctata, Philotarsopsis basipunctata] SN08

Abelopsocus cruciclypeus (New & Thornton 1981) [=Haplophallus cruciclypeus, Latrobiella cruciclypeus, Philotarsopsis cruciclypeus] SN08

Philotarsopsis guttata (Tillyard 1923) [=Philotarsis guttatus, Haplophallus guttatus, Latrobiella guttata; incl. *Philotarsopsis delicata Tillyard 1923, Philotarsis greyi Edwards 1950] SN08

Philotarsus flaviceps (Stephens 1836) Y02, M93 [=Psocus flaviceps M93, Caecilius flaviceps M93, Elipsocus flaviceps M93]

*Philotarsus picicornis (Fabricius 1793) M93 [=Hemerobius picicornis L02, Psocus picicornis M93; incl. Philotarsus flaviceps var. fuscoguttata Enderlein 1901 G74, Caecilius irroratus Curtis 1837 M93, Psocus lasiopterus Burmeister 1839 M93, C. lasiopterus M93, Mesopsocus poecilopterus Navás 1913 M93, Psocus pusillus Zetterstedt 1840 non Hemerobius pusillus Müller 1776 M93, Philotarsus picicornis var. stigma Kolbe 1880 G74, Ps. striatulus Stephens 1836 non Latreille 1795 M93]

*Type species of generic name indicated

REFERENCES

[B86] Badonnel, A. 1986. Psocoptères de Colombie (Insecta, Psocoptera). Spixiana 9 (2): 179–223.

[G74] Günther, K. K. 1974. Staubläuse, Psocoptera. Die Tierwelt Deutschlands 61: 1–315.

[L02] Li F. 2002. Psocoptera of China vol. 1. Science Press: Beijing.

[L03] Lienhard, C. 2003. Nomenclatural amendments concerning Chinese Psocoptera (Insecta), with remarks on species richness. Revue Suisse de Zoologie 110 (4): 695–721.

[M93] Mockford, E. L. 1993. North American Psocoptera (Insecta). Sandhill Crane Press, Inc.

[M03] Mockford, E. L. 2003. New species and records of Psocoptera from the Kuril Islands. Mitt. Mus. Nat.kd. Berl., Dtsch. Entomol. Z. 50 (2): 191–230.

[NL07] New, T. R., & C. Lienhard. 2007. The Psocoptera of Tropical South-east Asia. Brill: Leiden.

[SN08] Schmidt, E. R., & T. R. New. 2008. The Psocoptera (Insecta) of Tasmania, Australia. Memoirs of Museum Victoria 65: 71–152.

[Y02] Yoshizawa, K. 2002. Phylogeny and higher classification of suborder Psocomorpha (Insecta: Psocodea: ‘Psocoptera’). Zoological Journal of the Linnean Society 136: 371–400.

Peripsocidae

Peripsocus madescens, copyright Tom Murray.


Belongs within: Peripsocoidea.

The Peripsocidae are a group of barklice that are most common on bark or dead foliage, more rarely on low vegetation (New & Lienhard 2007).

Characters (from New & Lienhard 2007): Fore wing length ca 1.8–2.4 mm. Lacinial apex slender, bicuspid, outer cusp usually broader than inner. Vertex smooth or with few small setae. Tarsi two-segmented; claws with subapical tooth, pulvillus slender. Fore wing lacking areola postica; pterostigma of normal form, more or less expanded in central region. Wing setation usually reduced or absent, occasionally more conspicuous; hind wing glabrous, veins Rs and M fused for a length. Male with hypandrium simple; phallosome a closed frame, with pair of poriferous parameres; aedeagal arch usually with distinct apical constriction or process; endophallus with symmetrical sclerites. Female subgenital plate often with distinct, commonly bilobed, apical region; gonapophyses complete; dorsal valve large, usually setose at apex; external valve large, setose.

<==Peripsocidae [Peripsocinae]
    |--Orbiperipsocus Li 2002 [=Orbiperipaocus] L03
    |    `--*O. fractiflexus Li 2002 L03
    |--Campanulatus Li 2002 [=Canpanulatus] L03
    |    |--*C. jaculatorius Li 2002 [=C. jaculotoria, C. jaculotorius] L03
    |    |--C. lagenarius Li 2002 L02
    |    `--C. pictus (Thornton 1962) [=Peripsocus pictus] L02
    |--Bicuspidatus Li 1993 L02
    |    |--*B. pulchipunctatus Li 1993 L02
    |    |--B. guangdongensis Li 1993 L02
    |    `--B. sigillatus Li 1993 L02
    |--Properipsocus Li 1995 L02
    |    |--*P. gracilis Li 1995 L02
    |    |--P. bicornis (Thornton 1962) [=Peripsocus bicornis, Periterminalis bicornis] L02
    |    `--P. quadartus Li 2002 [=P. qradartus] L03
    |--Kaestneriella Roesler 1943 M93
    |    |--*K. pilosa M93
    |    |--K. fumosa (Banks 1903) [=Peripsocus fumosus] M93
    |    |--K. guatemalensis Y02
    |    `--K. tenebrosa Mockford & Sullivan 1990 M93
    |--Pericupsocus Li 2002 L02
    |    |--*P. cuspidatus Li 2002 L02
    |    |--P. alboguttatus (Dalman 1823) L02
    |    |--P. crenulatus (Thornton & Wong 1968) L02
    |    `--P. digitalis Li 2002 L02
    |--Cycloperipsocus Li 1993 SN08 [=Cycloperopsocus (l. c.) L03]
    |    |--*C. pangi Li 1993 SN08
    |    |--C. diplocyclus Li 2002 L02
    |    |--C. edwardsi (New 1973) [=Peripsocus edwardsi] SN08
    |    `--C. plurimaculatus (Li 1992) [=Peripsocus plurimaculatus] L02
    |--Periterminalis Li 1997 L02 [=Periteriminalis (l. c.) L03]
    |    |--*P. crytomeriae Li 1997 L02
    |    |--P. hiatus (Thornton & Wong 1968) [=Peripsocus hiatus] NL07
    |    |--P. lata Li 2002 [=P. latus] L03
    |    |--P. longicuspis Li 2002 L02
    |    `--P. shanxiensis Li 2002 L02
    |--Turriperipsocus Li 1997 L02
    |    |--*T. cunninghamiae Li 1997 L02, L03, L02 [=Peripsocus cunninghamius L02, T. cunninghamius (l. c.) L03]
    |    |--T. chiangensis (New 1973) [=Peripsocus chiangensis] NL07
    |    |--T. cupressisugus Li 2002 L02
    |    |--T. decimidentatus Li 1997 L02
    |    |--T. hypsodontus Li 2002 L02
    |    `--T. leshanensis Li 2002 L02
    |--Coniperipsocus Li 1997 L02 [=Comiperipsocus (l. c.) L03]
    |    |--*C. yuae Li 1997 L02
    |    |--C. jindingensis Li 2002 L03
    |    |--C. melanolomus Li 2002 L02
    |    |--C. minutissimus Li 2002 L02
    |    |--C. proceridentalis Li 2002 L02
    |    |--C. quadrangulus (Li 1992) [=Diplopsocus quadrangulus] L03
    |    `--C. quadrifascius (Li & Mockford 1993) [=Diplopsocus quadrifascius] L02
    |--Diplopsocus Li & Mockford 1993 L03
    |    |--*D. cupressicolus Li & Mockford 1993 L02 [=D. cupressicola (l. c.) L03]
    |    |--D. albostigmus Li & Mockford 1993 L02
    |    |--D. anoplus (Thornton & Wong 1968) NL07
    |    |--D. chebalingicus Li 1993 L02
    |    |--D. circularis (New & Thornton 1976) NL07
    |    |--D. fasciatus (Thornton 1959) NL07
    |    |--D. fujianicus Li & Mockford 1993 L02
    |    |--D. hainanensis Li & Mockford 1993 L02
    |    |--D. huananensis Li 2002 L02
    |    |--D. irregularis Li 1995 L02
    |    |--D. jiangxiensis Li & Mockford 1993 L02
    |    |--D. liupanshanensis Li 2002 [=D. liupenshanensis] L03
    |    |--D. lophacanthus Li & Mockford 1993 L02
    |    |--D. magniocellatus Li 1993 L03
    |    |--D. melanostictus Li & Mockford 1993 L02
    |    |--D. ningmingicus Li & Mockford 1993 L02
    |    |--D. oblongus Li & Mockford 1993 L02
    |    |--D. palmiformis Li & Mockford 1993 L02
    |    |--D. parabolicus Li & Mockford 1993 L02
    |    |--D. parviocellatus Li 1993 L02
    |    |--D. phaeophlebicus Li 2002 [=D. phaeophlebius] L03
    |    |--D. phagococcus Li & Mockford 1993 L02
    |    |--D. philopinnus Li & Mockford 1993 L02
    |    |--D. porosus Li & Mockford 1993 L02
    |    |--D. resupinatus Li & Mockford 1993 L02
    |    |--D. rhombeus Li & Mockford 1993 L02
    |    |--D. scrobiculatus Li 2002 L02
    |    |--D. selene (Thornton & Wong 1968) NL07
    |    |--D. shanxiensis Li & Mockford 1993 L02
    |    |--D. sichuanicus Li 1997 L02
    |    |--D. sylvaticus Li 2002 L02
    |    |--D. tibeticus Li & Mockford 1993 L02
    |    |--D. xilingxiaensis Li 1997 L02
    |    `--D. yangi Li & Mockford 1993 L02
    `--Peripsocus Hagen 1866 SN08 [incl. Peripsocopsis Tillyard 1923 G74]
         |  i. s.: P. africanus Enderlein 1902 E07
         |         P. ammonus Li 2002 L02
         |         P. annectens Li 2002 L02
         |         P. anura Vaughan, Thornton & New 1991 NL07
         |         P. apicatus Vaugan, Thornton & New 1989 NL07 [=Properipsocus apicatus L02]
         |         P. apiculatus Li 2002 L02
         |         P. attenuatus Li 2002 L02
         |         P. badimaculatus Li 1993 L02
         |         P. baishanzuicus Li 1995 L02
         |         P. baiyunshanicus Li 2002 L02
         |         P. barunus Vaughan, Thornton & New 1991 NL07
         |         P. beijingensis Li 2002 L02
         |         P. biacanthus Li & Yang 1988 L02
         |         P. bifasciarius Li 2002 L02
         |         P. bifasciatus SN08
         |         P. bifidus Thornton 1984 NL07
         |         P. bivari Baz 1988 L98
         |         P. brachyura Thornton & Wong 1968 NL07
         |         P. brinchangensis New & Lee 1991 NL07
         |         P. bucephalus Li 1992 L02
         |         P. bucerus Li 1992 L02
         |         P. bulbus Li 2002 L02
         |         P. camerunus B55
         |         P. cassideus Li 1999 L02
         |         P. caudatus Li 1997 L02
         |         P. changbaishanicus Li 2002 L02
         |         P. circinus Thornton & Wong 1968 NL07
         |         P. cochleus Schmidt & New 2008 SN08
         |         P. conoidalis Li 2002 L02
         |         P. consobrinus Pearman 1951 L98
         |         P. constrictus Thornton & Wong 1968 NL07
         |         P. corollaris Li 2002 L02
         |         P. crassicosta New & Lee 1991 NL07
         |         P. crenulatus Thornton & Wong 1968 NL07
         |         P. cylindratus Li 2002 L02
         |         P. decurvatus Li 1995 L02
         |         P. denticulatus Thornton & Wong 1968 NL07
         |         P. disdentus Li 1997 L03, L02 [=P. distentus (l. c.) L03]
         |         P. dongbeiensis Li 2002 L02
         |         P. duodecimidentus Li 2002 L02
         |         P. elongatus New & Thornton 1976 NL07
         |         P. equispineus Li 2002 [=P. eguispineus] L03
         |         P. exilis Li 1997 L02
         |         P. falsipictus Li 2002 L02
         |         P. ferrugineus Thornton & Wong 1968 NL07
         |         P. fici Smithers 1984 T13
         |         P. forcipatus Li 1993 L02
         |         P. forficatus Li 2002 L02
         |         P. fornicalis Li 1999 L02
         |         P. frimensis New & Lee 1991 NL07
         |         P. fulvescens Navás 1920 NL07
         |         P. furcellatus Li 2002 L02
         |         P. grandispineus Li 2002 L02
         |         P. guandishanicus Li 2002 L02
         |         P. hainanensis Li 2002 L02
         |         P. haplacanthus Li 1993 L02
         |         P. hedinianus Enderlein 1936 L02
         |         P. hickmani SN08
         |         P. hongkongensis Thornton & Wong 1968 NL07
         |         P. huashanensis Li 2002 L02
         |         P. ignis Okamoto 1910 NL07
         |         P. jianfenglingicus Li 2002 L02
         |         P. jiangxiensis Li 2002 L02
         |         P. jilinicus Li 2002 L02
         |         P. jinggangshanicus Li 2002 L02
         |         P. jinshaanensis Li 2002 L02
         |         P. jinxiuensis Li 2002 L02
         |         P. kunmingiensis Li 2002 L02
         |         P. laoshanicus Li 2002 L02
         |         P. laricis Li 2002 L02
         |         P. latispineus Li 2002 L02
         |         P. leleupi Badonnel 1976 L98
         |         P. leptorrhizus Li 1997 L02
         |         P. lifashengi Lienhard 2003 [=P. orientalis Li 2002 non Garcia Aldrete 1999] L03
         |         P. limi New & Thornton 1976 NL07
         |         P. longifurcus Li 2002 L02
         |         P. louguantaiensis Li 2002 L02
         |         P. lunaris Li 1993 L02
         |         P. luotongshanicus Li 2002 L02
         |         P. machadoi Badonnel 1955 B55
         |         P. macrosiphus Li 1993 L02
         |         P. magnimammus Li 2002 L02
         |         P. maoricus (Tillyard 1923) SN08 (see below for synonymy)
         |         P. medifasciarius Li 2002 [=P. medifaciarius] L03
         |         P. medimacularis Li 1995 L02
         |         P. medispineus Li 2002 L02
         |         P. megalophus Li 1995 L02
         |         P. melaleucae New 1971 SN08
         |         P. meridionalis Li 1999 L02
         |         P. microcheilus Li 1993 [=P. microcheilius] L03
         |         P. milleri (Tillyard 1923) [=Peripsocopsis milleri; incl. Peripsocus eucalypti Edwards 1950] SN08
         |         P. mingshanicus Li 1997 L02
         |         P. mirabilis Li 1995 L02
         |         P. morulops (Tillyard 1923) [=Peripsocopsis morulops] SN08
         |         P. mutilatus Badonnel 1986 B86
         |         P. nanjingensis Li 2002 L02
         |         P. nanus Navás 1922 NL07
         |         P. nasutus B86
         |         P. nitens Thornton & Wong 1968 NL07, L98
         |         P. octoidentus Li 2002 L02
         |         P. oculatus Enderlein 1926 NL07
         |         P. ‘oculatus’ Badonnel 1955 non Enderlein 1926 [=P. ocuatus] B55
         |         P. oculimacularis Li 2002 L02
         |         P. odontopetalus Li 2002 [=P. codntopetalus] L03
         |         P. oligodontus Li 2002 L02
         |         P. optimalis Li 1997 L02
         |         P. orbiculatus Li 1993 L02
         |         P. orebius Li 2002 L02
         |         P. orientalis Garcia Aldrete 1999 NL07
         |         P. oxydontus Li 2002 L02
         |         P. pamae Schmidt & New 2008 SN08
         |         P. papillatus Vaughan, Thornton & New 1989 NL07
         |         P. parareicherti Vaughan, Thornton & New 1989 NL07
         |         P. paraspinosus New & Lee 1991 NL07
         |         P. parvulus Kolbe 1880 (see below for synonymy) G74
         |         P. parvus Li 1995 L02
         |         P. phaeochilus Li 2002 L02
         |         P. plagiotropus Li 2002 L02
         |         P. platyopterus Li 1997 L02
         |         P. plstypus Li 1995 L03, L02 [=L. platylpus (l. c.) L03, L. platypus (l. c.) L03]
         |         P. polygonalis Li 1995 L02
         |         P. polyoacanthus Li 1993 L02
         |         P. pseudoquercicolus Thornton 1962 L02
         |         P. qingchengshanicus Li 2002 L02
         |         P. qingdaoensis Li 1993 L02
         |         P. quadratidentalis Li 2002 L02
         |         P. quadratiprocessus Li 1993 L02
         |         P. quadripunctatus Badonnel 1955 L02 [incl. P. quadripunctatus f. brachypterus Badonnel 1955 B55]
         |         P. quattuordecimus Li 1995 L02
         |         P. quercicola Enderlein 1906 NL07
         |         P. reduncus Li 2002 L02
         |         P. reflexibilis Li 2002 L02
         |         P. reicherti Enderlein 1903 NL07
         |         P. rhombicus Li 2002 L02
         |         P. rhomboacanthus Li 2002 [=P. rhombocanthus] L03
         |         P. scalpratus Li 2002 L02
         |         P. scapiformis Li 2002 L02
         |         P. sedcimidentalis Li 1997 [=P. sedecimidentalis] L03
         |         P. sexidentus Li 2002 L02
         |         P. shilinensis Li 2002 L02
         |         P. siculiformis Li 2002 L02
         |         P. ‘similis’ Enderlein 1903 non P. phaeopterus var. similis Kolbe 1880 NL07
         |         P. ‘similis’ Badonnel 1955 nec Kolbe 1880 nec Enderlein 1903 B55
         |         P. singularis Banks 1937 L02
         |         P. spinosus Thornton 1959 L02
         |         P. stenopterus Thornton & Wong 1964 L02
         |         P. stigmatus Thornton & Wong 1968 NL07
         |         P. stigmostigmus Li 2002 L02
         |         P. stipiatus Li 1993 L02
         |         P. suffitus Enderlein 1903 NL07
         |         P. suoxiyuicus Li 2002 L02
         |         P. taipingensis New & Lee 1991 NL07
         |         P. terricolis Badonnel 1986 L02
         |         P. tillyardi New 1973 SN08
         |         P. tinctus Vaughan, Thornton & New 1989 NL07
         |         P. transivenus Li 2002 L02
         |         P. tredecimus Li 1997 L02
         |         P. trigonoispineus Li 2002 L02
         |         P. umbrosus Navás 1911 (n. d.) L98
         |         P. undecimidentus Li 2002 L02
         |         P. undulatus Li 2002 L02
         |         P. uniformis New & Thornton 1976 NL07
         |         P. valvulus Thornton & Wong 1968 NL07
         |         P. variatus Soehardjan & Hamann 1959 NL07 [=Cycloperipsocus variatus L02]
         |         P. varidentatus Li 2002 [=P. septemidentus Li 2001 (n. n.), P. varidentus] L03
         |         P. vescus Li 1997 L02
         |         P. viriosus Li 1995 L02
         |         P. wuhoi Li 2002 L02
         |         P. wuyishanicus Li 1999 L02
         |         P. xanthochilus Li 2002 L02
         |         P. xihuensis Li 2002 L02
         |         P. yongi New & Lee 1991 NL07
         |         P. yuleki Galil in Galil & Halperin 1983 L98
         |         P. zhangispineus Li 2002 L02
         |         P. zhangliani Li 2002 [=P. zhangliangi, P. zhengliangi] L03
         |         P. ziguiensis Li 1997 L02
         |--Group III M93
         |    |--P. stagnivagus Chapman 1930 M93
         |    `--Subgroup B M93
         |         |--*P. phaeopterus (Stephens 1836) M93 (see below for synonymy)
         |         `--P. didymus Roesler 1939 M93, G74 (see below for synonymy)
         `--Group I M93
              |--Subgroup A M93
              |    |--P. madidus (Hagen 1861) [=Psocus madidus; incl. Ps. permadidus Walsh 1863, Caecilius permadidus] M93
              |    |--P. nigrescens M93
              |    |--P. pauliani Badonnel 1949 M93
              |    |--P. quadrifasciatus (Harris 1869) (see below for synonymy) M93
              |    `--P. reductus Badonnel 1943 M93
              `--Subgroup B M93
                   |--P. alachuae Mockford 1971 M93
                   |--P. alboguttatus (Dalman 1823) (see below for synonymy) M93
                   |--P. maculosus Mockford 1971 M93
                   |--P. madescens (Walsh 1863) [=Psocus madescens] M93
                   |--P. minimus Mockford 1971 M93
                   |--P. potosi Mockford 1971 M93
                   `--P. teutonicus M93

Peripsocus alboguttatus (Dalman 1823) [=Psocus alboguttatus; incl. Ps. pupillatus Walker 1853, Peripsocus pupillatus, Ps. quadrimaculatus Stephens 1836 non Latreille 1794] M93

Peripsocus didymus Roesler 1939 M93, G74 [incl. P. didymus silesiaca Obr 1948 G74, P. didymus truncatus Badonnel 1943 G74]

Peripsocus maoricus (Tillyard 1923) SN08 [=*Peripsocopsis maoricus NL07, SN08; incl. Peripsocus macropterus Edwards 1950 SN08]

Peripsocus parvulus Kolbe 1880 [=P. alboguttatus var. parvulus; incl. P. parvulus f. longipennis Roesler 1939] G74

*Peripsocus phaeopterus (Stephens 1836) M93 [=Psocus phaeopterus SN08; incl. Pe. fortunatus Navás 1916 L98, Ps. nigricornis Stephens 1836 M93, Ps. obscurus Rambur 1842 M93, Peripsocus phaeopterus var. similis Kolbe 1880 G74]

Peripsocus quadrifasciatus (Harris 1869) [=Psocus quadrifasciatus; incl. Pe. subpupillatus ab. quadriramosus Enderlein 1901, Ps. subfasciatus Rambur 1842 non Zetterstedt 1840, Pe. subfasciatus, Pe. subpupillatus McLachlan 1883] M93

*Type species of generic name indicated

REFERENCES

[B55] Badonnel, A. 1955. Psocoptères de l'Angola. Diamang Publicações Culturais 26: 11–267.

[B86] Badonnel, A. 1986. Psocoptères de Colombie (Insecta, Psocoptera). Spixiana 9 (2): 179–223.

[E07] Enderlein, G. 1907. Wissenschaftliche Ergebnisse der Schwedischen Zoologischen Expedition nach dem Kilimandjaro, dem Meru und den Umgebenden Massaisteppen Deutsch-Ostafrikas 1905–1906 vol. 15. Corrodentia pt 2. Copeognatha. Almqvist & Wiksells Boktryckeri-A. B.: Uppsala.

[G74] Günther, K. K. 1974. Staubläuse, Psocoptera. Die Tierwelt Deutschlands 61: 1–315.

[L02] Li F. 2002. Psocoptera of China vol. 2. Science Press: Beijing.

[L98] Lienhard, C. 1998. Faune de France. France et Régions Limitrophes. 83. Psocoptères Euro-Méditerranéens. Fédération Française des Sociétés de Sciences Naturelles: Paris.

[L03] Lienhard, C. 2003. Nomenclatural amendments concerning Chinese Psocoptera (Insecta), with remarks on species richness. Revue Suisse de Zoologie 110 (4): 695–721.

[M93] Mockford, E. L. 1993. North American Psocoptera (Insecta). Sandhill Crane Press, Inc.

[NL07] New, T. R., & C. Lienhard. 2007. The Psocoptera of Tropical South-east Asia. Brill: Leiden.

[SN08] Schmidt, E. R., & T. R. New. 2008. The Psocoptera (Insecta) of Tasmania, Australia. Memoirs of Museum Victoria 65: 71–152.

[T13] Taylor, C. K. 2013. Annotated bibliography for Barrow Island terrestrial invertebrates. Records of the Western Australian Museum Supplement 83: 135–144.

[Y02] Yoshizawa, K. 2002. Phylogeny and higher classification of suborder Psocomorpha (Insecta: Psocodea: ‘Psocoptera’). Zoological Journal of the Linnean Society 136: 371–400.

Stenopsocidae

Stenopsocus stigmaticus, copyright Sanja565658.


Belongs within: Caeciliusetae.

The Stenopsocidae are a group of foliage-associated barklice that are most diverse in China (New & Lienhard 2007).

Characters (from New & Lienhard 2007): Fore wing length ca 3.5–4.5 mm. Mandibles usually elongate; lacinial apex narrow, incipiently bicuspid or with small rounded denticles. Tarsi two-segmented, claws without subapical tooth, pulvillus broad. Fore wing veins with short setae in one rank; hind wing margin bare or with few setae on radial fork region; fore wing with crossvein from pterostigma to Rs, crossvein from Cu1a to M. Two or three ventral abdominal vesicles present. Female with external gonapophysis valve strongly reduced, lacking seta; ventral and dorsal valves slender, elongage; apex of subgenital plate rounded or slightly bilobed; spermatheca with small basal pouch.

<==Stenopsocidae [Stenopsocinae]
    |--Malostenopsocus Li 1992 L02
    |    |--*M. yunnanicus Li 1992 L02
    |    |--M. cubitalis (Thornton 1984) [=Stenopsocus cubitalis] NL07
    |    |--M. expansus Li 1992 L02
    |    |--M. immaculatus Li 1992 L02
    |    |--M. intertextus Li 1992 L02
    |    |--M. mucronatus Li 1992 L02
    |    |--M. parallelinervius Li 1992 L02
    |    |--M. plurifasciatus Li 1995 L02
    |    `--M. sulphurepterus Li 1992 L02
    |--Graphopsocus Kolbe 1880 [incl. Teratopsocus Reuter 1893] L02
    |    |--*G. cruciatus (Linnaeus 1768) L02 (see below for synonymy)
    |    |--G. borealis Li 2002 L02
    |    |--G. choui Li 1992 L02
    |    |--G. epicharis Li 2002 L02
    |    |--G. euneurus Li 2002 L02
    |    |--G. evanidus Li 2002 L02
    |    |--G. flavipunctatus Li 2002 L02
    |    |--G. infirmus Banks 1920 NL07
    |    |--G. intrans Li 2002 L02
    |    |--G. leptostigmus Li 2002 L02
    |    |--G. liaoningensis Li 2002 L02
    |    |--G. luojiashanensis Li 2002 [=G. luojiashnensis] L03
    |    |--G. megistigmus Li 2002 L02
    |    |--G. mexicanus M93
    |    |--G. obcuneatus Li 2002 L02
    |    |--G. panduratus Li 1989 L02
    |    |--G. platynotus Li 2002 L02
    |    |--G. putuoshanensis Li 2002 L03 [=G. patuoshanensis L02, G. prtuoshanensis L03]
    |    |--G. shaanxiensis Li 1989 L02
    |    |--G. sinuolatus Li 2002 L02
    |    |--G. subaequalis Banks 1920 NL07
    |    `--G. urceolatus Li 1993 L02
    `--Stenopsocus Hagen 1866 [incl. Cubipilis Li 1993] M03
         |--*S. immaculatus (Stephens 1836) (see below for synonymy) L98
         |--S. adisoemartoi Vaughan, Thornton & New 1989 NL07
         |--S. albipileus Smithers 1974 NL07
         |--S. albus Li 1992 L02
         |--‘Cubipilis’ anchorocaulis Li 1993 L02
         |--S. angustifurcus Li 2002 L02
         |--S. angustistriatus Li 2002 L02
         |--S. anthracinus Li 1989 L03, L02 [=S. anthraeinus (l. c.) L03]
         |--S. aphidiformis Enderlein 1906 L02
         |--S. aureus Li 2002 L02
         |--‘Cubipilis’ baishanzuensis Li 1995 L02
         |--‘Cubipilis’ beijingensis Li 2002 L02
         |--S. bellatulus Li 1989 L02
         |--S. betulus Li 2002 L02
         |--‘Cubipilis’ bicoloratus Li 2002 L02
         |--S. bicoloripes Enderlein 1936 L02
         |--S. biconicus Li 2002 L02
         |--S. biconvexus Li 1997 L02
         |--S. bimaculatus Li 2002 L02
         |--S. bipunctatus Li 2002 L02
         |--S. bombusus Li 2002 L02
         |--S. brachychelus Li & Yang 1988 L03, L02 [=S. brachychelur L03, S. brachychilus (l. c.) L03]
         |--S. brachycladus Li 1989 L02
         |--S. brachyodicrus Li 2002 L02
         |--S. brevicapitus Li 1997 L02
         |--S. brevivalvaris Li 2002 L02
         |--S. capacimacularus Li 1993 L02
         |--S. cassideus Li 1992 L02
         |--S. ceuthozibrinus Li 2002 L02
         |--S. changbaishanicus Li 2002 L02
         |--‘Cubipilis’ chebalingensis Li 1993 L02
         |--S. chusanensis Navas 1933 L02
         |--S. concisus Li 2002 L02
         |--‘Cubipilis’ coronatus Li 2002 L02
         |--‘Cubipilis’ cunnatus Li 2002 L02
         |--S. dactylinus Li 1997 L02
         |--‘Cubipilis’ denivalvis Li 1993 L02
         |--S. dichospilus Li 2002 [=S. dechospilus] L03
         |--S. dictyodromus Li 1993 L02
         |--S. disphaeroides Li 2002 L02
         |--S. dissimilis Banks 1937 NL07
         |--S. emeishanicus Li 2002 [=S. emishanicus] L03
         |--S. eucallus Li & Yang 1988 L02
         |--S. externus Banks 1937 L02
         |--‘Cubipilis’ falcatus Li 2002 L02
         |--S. fanjingshanicus Li & Yang 1988 L02
         |--‘Cubipilis’ fastigiatus Li 2002 L02
         |--S. faungi Li 1999 L02
         |--‘Cubipilis’ flavicapitus Li 2002 L02
         |--S. flavicaudatus Li 2002 L02
         |--S. flavifrons Li 1989 L02
         |--S. flavinigrus Li 2002 L02
         |--S. floralis Li 2002 L02
         |--S. foliaceus Li 1997 L02
         |--S. formosanus Banks 1937 L02
         |--S. frontalis Li 1989 L02
         |--S. frontimaculatus Li 1992 L02
         |--S. fulivertex Li 2002 L02
         |--‘Cubipilis’ furcimaculatus Li 2002 L02
         |--‘Cubipilis’ gannanensis Li 2002 L02
         |--S. gansuensis Li 2002 L02
         |--S. genostictus Li 2002 L02
         |--S. gibbulosus Li 1995 L02
         |--S. gracilimaculatus Li 2002 L02
         |--S. gracillimus Li & Yang 1988 L02
         |--S. guizhouiensis Li 2002 L02
         |--*Cubipilis’ hamaocaulis Li 1993 L02
         |--S. hemiostictus Li 2002 L02
         |--S. hexagonus Li 2002 L02
         |--S. huangshanicus Li 2002 L02
         |--S. hunanicus Li 1992 L02
         |--S. isotomus Li 2002 L02
         |--S. jocosus Banks 1939 NL07
         |--S. kunmingiensis Li 2002 [=S. kuanmingiensis] L03
         |--S. lachlani Kolbe 1880 [=S. immaculatus var. lachlani] G74
         |--S. lacteus Li 1997 L02
         |--S. laterimaculatus Li 2002 [=S. latermaculatus] L03
         |--‘Cubipilis’ leucoresbius Li 2002 [=C. leucorsbicus] L03
         |--S. lifashengi Mockford 2003 M03
         |--S. liuae Li 2002 L02
         |--S. liupanshanensis Li 2002 [=S. liupanshanicus] L03
         |--‘Cubipilis’ locularis Li 2002 L02
         |--S. longicuspis Li 1997 L02
         |--S. longitudinalis Li 2002 L02
         |--S. macrocheirus Li 2002 L02
         |--‘Cubipilis’ macrostigmis Li 1995 L02
         |--S. maculosus Li & Yang 1988 L02
         |--‘Cubipilis’ majusculus Li 2002 L02
         |--S. makii Takahasi 1938 L02
         |--S. maximalis Li 1997 L02
         |--S. melanocephalus Li 1997 L02
         |--‘Cubipilis’ mesozonalis Li 2002 L02
         |--S. metasticus Li 2002 [=S. metostictus] L03
         |--S. naevicapitatus Li 2002 L02
         |--S. niger Enderlein 1906 M03
         |--S. nigricellus Okamoto 1907 M03
         |--S. obscurus Li 1997 L02
         |--S. oculimaculatus Li 1992 L02
         |--‘Cubipilis’ orbiculatus Li 1995 L02
         |--‘Cubipilis’ ovalimacularis Li 2002 L02
         |--‘Cubipilis’ palmatus Li 2002 L02
         |--S. parviforficatus Li 2002 L02
         |--S. pavonicus Li 2002 L02
         |--S. paxillivalvaris Li 2002 [=S. pasillivalvaris] L03
         |--‘Cubipilis’ pellucidus Li 2002 L02
         |--S. percussus Li 1995 L02
         |--S. periostictus Li 2002 L02
         |--S. perspicuus Li 1997 L02
         |--‘Cubipilis’ phaeostigmis Li 1995 L02
         |--S. phaeostigmus Li 1992 L02
         |--S. phaneostriatus Li 2002 L02
         |--S. pilosus Enderlein 1926 NL07
         |--S. platynotus Li 1995 L02
         |--S. platyocephalus Li 2002 L02
         |--‘Cubipilis’ platytaenius Li 2002 L02
         |--S. podorphus Li 1997 L02
         |--S. polyceratus Li 2002 L02
         |--‘Cubipilis’ pyriformis Li 2002 L02
         |--‘Cubipilis’ quaternatus Li 2002 L02
         |--S. radimaculatus Li 1989 L02
         |--S. revolutus Li 1993 L03, L02 [=S. revolatus (l. c.) L03]
         |--S. rubellus Thornton 1984 NL07
         |--S. shennongjiaensis Li 2002 L02
         |--S. sichuanicus Li 2002 L02
         |--S. silvaticus Li 2002 L02
         |--‘Cubipilis’ spilipsocius Li 1997 L02
         |--S. spongiosus Li 2002 L02
         |--S. stigmaticus (Imhoff & Labram 1845) [=Psocus stigmaticus; incl. S. lineolatus Navás 1916] G74
         |--S. striolatus Li 1992 L02
         |--S. symipsarous Li 2002 L02
         |--S. thermophilus Li 2002 L02
         |--‘Cubipilis’ tianmushanensis Li 2002 L02
         |--S. tibialis Banks 1937 L02
         |--S. tonkinensis Enderlein 1903 NL07
         |--S. tribulbus Li 1993 L02
         |--‘Cubipilis’ trinotatus Li 1995 L02
         |--S. tripartibilis Li 2002 L02
         |--S. trisetus Li 2002 L02
         |--S. turgidus Li 1997 L02
         |--S. uniformis (Hagen 1859) [=Graphopsocus uniformis] NL07
         |--S. wuxiaensis Li 1997 L02
         |--S. xanthophaeus Li 2002 L02
         |--S. xanthostigmus Li 2002 L02
         |--S. xiangxiensis Li 1992 L02
         |--S. xilingxianicus Li 1997 L03, L02 [=S. xilingxianus (l. c.) L03]
         |--‘Cubipilis’ yuensis Li 1993 L02
         `--S. zonatus Li 1989 L02

Nomen nudum: Stenopsocus psyllipennis Costa 1886 L98

*Graphopsocus cruciatus (Linnaeus 1768) L02 [=Hemerobius cruciatus L02, Psocus cruciatus M93, Stenopsocus cruciatus M93; incl. Graphopsocus cruciatus var. brevipennis Enderlein 1903 M93, Psocus costalis Stephens 1836 M93, *Teratopsocus maculipennis Reuter 1893 NL07, L98, P. nervosus Stephens 1836 M93, Stenopsocus cruciatus var. nervosus M93, Hemerobius quadrifasciatus Fabricius 1787 M93, H. quadripunctatus Fabricius 1787 G74, Psocus quatuorpunctatus L95, P. subocellatus Stephens 1836 M93]

*Stenopsocus immaculatus (Stephens 1836) [=Psocus immaculatus; incl. P. flavescens Stephens 1836, P. flavicans Zetterstedt 1840 nec Hemerobius flavicans Linnaeus 1758 nec P. flavicans Stephens 1836, P. rufescens Stephens 1836, P. strigosus Burmeister 1839, P. subfumipennis Zetterstedt 1840, P. venosus Stephens 1836] L98

*Type species of generic name indicated

REFERENCES

[G74] Günther, K. K. 1974. Staubläuse, Psocoptera. Die Tierwelt Deutschlands 61: 1–315.

[L95] Latreille, P. A. 1795. Extrait d'un mémoire pour servir de suite à l'histoire des termès, ou fourmis blanches. Bulletin de la Société Philomatique, a ses Correspondans 41–42: 1–3.

[L02] Li F. 2002. Psocoptera of China vol. 1. Science Press: Beijing.

[L98] Lienhard, C. 1998. Faune de France. France et Régions Limitrophes. 83. Psocoptères Euro-Méditerranéens. Fédération Française des Sociétés de Sciences Naturelles: Paris.

[L03] Lienhard, C. 2003. Nomenclatural amendments concerning Chinese Psocoptera (Insecta), with remarks on species richness. Revue Suisse de Zoologie 110 (4): 695–721.

[M93] Mockford, E. L. 1993. North American Psocoptera (Insecta). Sandhill Crane Press, Inc.

[M03] Mockford, E. L. 2003. New species and records of Psocoptera from the Kuril Islands. Mitt. Mus. Nat.kd. Berl., Dtsch. Entomol. Z. 50 (2): 191–230.

[NL07] New, T. R., & C. Lienhard. 2007. The Psocoptera of Tropical South-east Asia. Brill: Leiden.