Protohydrachnidia

Rhyncholimnochares sp., copyright Ray Fisher.

Belongs within: Hydrachnidia.
Contains: Eylais.

The Protohydrachnidia are a clade of water mites in which the lateral eyes lack muscles, the last pair of idiosomal dorsoventral muscles are arranged in a median position, and the ejaculatory complex has the distal-sclerite arms and apical setae reduced.

<==Protohydrachnidia
    |--Hydrovolzioidea TBG00
    |    |--Hydrovolziidae [Hydrovolziinae] WWO01
    |    |    |--Stygovolzia Imamura 1957 TBG00
    |    |    |    `--S. uenoi (Imamura 1957) TBG00
    |    |    |--Hydrovolziella Viets 1935 TBG00
    |    |    |    `--*H. lata (Walter 1935) [=Hydrovolzia lata] TBG00
    |    |    `--Hydrovolzia Thor 1905 TBG00
    |    |         |--H. baderi G96
    |    |         |--H. cancellata Walter 1906 TBG00
    |    |         |--H. gerhardi TBG00
    |    |         |--H. japonica Imamura 1961 TBG00
    |    |         |--H. marshallae WL09
    |    |         |--H. persica Bader & Sepasgozarian 1980 TBG00
    |    |         `--H. placophora (Monti 1905) TBG00
    |    `--Acherontacaridae [Acherontacarinae] TBG00
    |         |--Bharatavolzia Cook 1967 TBG00
    |         |    |--B. (Bharatavolzia) muscicola Cook 1967 S86a
    |         |    `--B. (Bharatavolziella Cook 1967) TBG00
    |         |         |--B. (B.) cooki Schwoerbel & Sepasgozarian 1980 S86a
    |         |         `--B. (B.) pallida Cook 1967 S86a
    |         `--Acherontacarus Viets 1932 (see below for synonymy) TBG00
    |              |--*A. halacaroides Viets 1932 TBG00
    |              |--A. bicornis Cook 1974 TBG00
    |              |--A. cedro Lundblad 1962 TBG00
    |              |--A. cicolanii GS96
    |              |--A. dividuus TBG00
    |              |--A. fonticolus Viets 1934 TBG00
    |              |--A. rutilans Angelier 1951 TBG00
    |              |--A. tuberculatus TBG00
    |              `--A. vietsi Angelier 1951 TBG00
    `--Eylaoidea WL09
         |  i. s.: Apheviderulix [Apheviderulicidae] WL09
         |           |--A. santana WL09
         |           `--A. welwitchioides WL09
         |--Piersigiidae WWO01
         |    |--Stygolimnocharinae WL09
         |    |    |--Parawandesia Angelier 1951 WL09, S86b
         |    |    |    `--P. chappuisi Angelier 1951 S86b
         |    |    `--Stygolimnochares Cook 1967 H98
         |    |         |--S. australica Cook 1986 H98
         |    |         `--S. elongata Cook 1967 S86b
         |    `--Piersigiinae WL09
         |         |--Austrapiersigia montana WL09
         |         `--Piersigia WL09
         |              |--P. crusta WL09
         |              |--P. intermedia Wi91
         |              `--P. limnophila WL09
         `--+--Eylais Wo91
            `--Limnocharidae WWO01
                 |--Limnocharinae WL09
                 |    |--Limnochares Latreille 1796 H98
                 |    |    |--L. (Cyclothrix) americana WP99
                 |    |    |--L. aquatica (Linnaeus 1758) Wo91 [=Acarus aquaticus L02]
                 |    |    |--L. australica Lundblad 1941 H98
                 |    |    |--L. crinita WL09
                 |    |    `--L. holoserica [=Hydrachna holoserica] G20
                 |    `--Neolimnochares Lundblad 1937 WL09, RFF05
                 |         |--N. breviscuta Lundblad 1941 RFF05
                 |         |--N. kakadu Harvey 1990 H98
                 |         |--N. petrophila Lundblad 1937 RFF05
                 |         |--N. placophora Lundblad 1937 RFF05
                 |         `--N. pugiunculata Viets 1954 RFF05
                 `--Rhyncholimnocharinae WL09
                      |--Austrolimnochares womersleyi WL09
                      `--Rhyncholimnochares Lundblad 1936 RFF05
                           |  i. s.: R. kittitanniana WL09
                           |         R. womersleyi (Lundblad 1952) [=Paralimnochares womersleyi] H98
                           |--R. (Rhyncholimnochares) RFF05
                           |    |--R. (R.) andina Lundblad 1953 RFF05
                           |    |--R. (R.) expansiseta Cook 1980 RFF05
                           |    |--R. (R.) gracilirostris Lundblad 1953 RFF05
                           |    |--R. (R.) lamellipalpis Lundblad 1936 RFF05
                           |    |--R. (R.) longipalpis Lundblad 1953 RFF05
                           |    `--R. (R.) longiscuta Lundblad 1941 RFF05
                           `--R. (Paralimnochares Lundblad 1937) RFF05
                                |--R. (P.) dipersiai (Rosso de Ferradás 1975) RFF05
                                `--R. (P.) sursumhians (Lundblad 1937) RFF05

Acherontacarus Viets 1932 [incl. Acherontacarellus Lundblad 1962, Acherontacaropsis Cook 1967, Neoacherontacarus Bader 1989] TBG00

*Type species of generic name indicated

REFERENCES

[GS96] Gerecke, R., & A. Di Sabatino. 1996. Historical zoogeography and evolution of habitat preference in water mites (Hydracarina) of the central Mediterranean region. In: Mitchell, R., D. J. Horn, G. R. Needham & W. C. Welbourn (eds) Acarology IX vol. 1. Proceedings pp. 523–527. Ohio Biological Survey: Columbus (Ohio).

[G96] Gil, M. J. 1996. Water-mite (Hydracarina) larvae of some selected genera from Spain: a phylogenetic inference. In: Mitchell, R., D. J. Horn, G. R. Needham & W. C. Welbourn (eds) Acarology IX vol. 1. Proceedings pp. 543–546. Ohio Biological Survey: Columbus (Ohio).

[G20] Goldfuss, G. A. 1820. Handbuch der Naturgeschichte vol. 3. Handbuch der Zoologie pt 1. Johann Leonhard Schrag: Nürnberg.

[H98] Halliday, R. B. 1998. Mites of Australia: A checklist and bibliography. CSIRO Publishing: Collingwood.

[L02] Latreille, P. A. 1802. Histoire Naturelle, générale et particulière des crustacés et des insectes vol. 3. Familles naturelles des genres. F. Dufart: Paris.

[RFF05] Rosso de Ferradás, B., & H. R. Fernández. 2005. Elenco y biogeografía de los ácaros acuáticos (Acari, Parasitengona, Hydrachnidia) de Sudamérica. Graellsia 61 (2): 181–224.

[S86a] Schwoerbel, J. 1986a. Acari: Limnohalacaridae and Hydrovolziidae. In: Botosaneanu, L. (ed.) Stygofauna Mundi: A Faunistic, Distributional, and Ecological Synthesis of the World Fauna inhabiting Subterranean Waters (including the Marine Interstitial) pp. 643–647. E. J. Brill/Dr W. Backhuys: Leiden.

[S86b] Schwoerbel, J. 1986b. Acari: “Hydrachnellae”. In: Botosaneanu, L. (ed.) Stygofauna Mundi: A Faunistic, Distributional, and Ecological Synthesis of the World Fauna inhabiting Subterranean Waters (including the Marine Interstitial) pp. 652–696. E. J. Brill/Dr W. Backhuys: Leiden.

[TBG00] Tuzovskij, P., D. Benfatti & R. Gerecke. 2000. The water mite family Acherontacaridae Cook, 1967 nov. stat. and diagnosis of the superfamily Hydrovolzioidea Thor, 1905 (Acariformes, Hydrachnidia). Acarologia 41: 451–473.

[WL09] Walter, D. E., E. E. Lindquist, I. M. Smith, D. R. Cook & G. W. Krantz. 2009. Order Trombidiformes. In: Krantz, G. W., & D. E. Walter (eds) A Manual of Acarology 3^rd ed. pp. 233–420. Texas Tech University Press.

[WP99] Walter, D. E., & H. C. Proctor. 1999. Mites: Ecology, Evolution and Behaviour. CABI Publishing: Wallingford (UK).

[Wi91] Witte, H. 1991. The phylogenetic relationships within the Parasitengonae. In: Dusbábek, F., & V. Bukva (eds) Modern Acarology: Proceedings of the VIII International Congress of Acarology, held in České Budĕjovice, Czechoslovakia, 6–11 August 1990 vol. 2 pp. 171–182. SPB Academic Publishing: The Hague.

[Wo91] Wohltmann, A. 1991. Nutrition and respiratory rate of water mites (Actinedida: Hydrachnidia) from a phylogenetic point of view. In: Dusbábek, F., & V. Bukva (eds) Modern Acarology: Proceedings of the VIII International Congress of Acarology, held in České Budĕjovice, Czechoslovakia, 6–11 August 1990 vol. 2 pp. 543–548. SPB Academic Publishing: The Hague.

[WWO01] Wohltmann, A., H. Witte & R. Olomski. 2001. Organismal patterns causing high potential for adaptive radiation in Parasitengonae (Acari: Prostigmata). In: Halliday, R. B., D. E. Walter, H. C. Proctor, R. A. Norton & M. J. Colloff (eds) Acarology: Proceedings of the 10^th International Congress pp. 83–99. CSIRO Publishing: Melbourne.

Wandesia

Wandesia thori, copyright Faculty of Science and Mathematics, Podgorica.

Belongs within: Hydryphantidae.

Wandesia is a genus of elongate-bodied water mites with widely separated coxae and lacking dorsal plates or genital flaps.

<==Wandesia Schechtel 1912 RFF05
    |  i. s.: W. cavecerberum GS96
    |         W. saginata GS96
    |         W. troma Cook 1986 H98
    |--W. (Wandesia) S86
    |    |--W. (W.) centensis Habeeb 1973 S86
    |    |--W. (W.) extendens Cook 1980 S86
    |    |--W. (W.) racovitzai Gledhill 1970 S86
    |    `--W. (W.) thori Schechtel 1912 S86
    |--W. (Allowandesia Schwoerbel 1961) S86
    |    `--W. (A.) japonica Imamura 1956 S86
    |--W. (Mesowandesia André & Naudo 1962) S86
    |    `--W. (M.) riparia Habeeb 1954 S86
    |--W. (Partnuniella Viets 1938) RFF05
    |    |--W. (P.) chechoi Cook 1988 [=W. (Pseudowandesia) chechoi] RFF05
    |    |--W. (P.) cubana Orghidan & Georgesco 1977 S86
    |    |--W. (P.) lethaea (Besch 1964) RFF05
    |    |--W. (P.) stalagmophila Lundblad 1941 RFF05
    |    |--W. (P.) thermalis (Viets 1938) RFF05
    |    `--W. (P.) walteri Lundblad 1937 RFF05
    `--W. (Pseudowandesia Habeeb 1958) RFF05
         |--W. (P.) andiana Cook 1980 RFF05
         |--W. (P.) approximata Cook 1967 S86
         |--W. (P.) gaspensis Habeeb 1955 S86
         |--W. (P.) glareosa Weigmann & Schminke 1970 S86
         |--W. (P.) stygophila Szalay 1944 S86
         |--W. (P.) tanasachii Orgidan & Georgescu 1977 S86
         `--W. (P.) vermiformis Cook 1967 S86

*Type species of generic name indicated

REFERENCES

[GS96] Gerecke, R., & A. Di Sabatino. 1996. Historical zoogeography and evolution of habitat preference in water mites (Hydracarina) of the central Mediterranean region. In: Mitchell, R., D. J. Horn, G. R. Needham & W. C. Welbourn (eds) Acarology IX vol. 1. Proceedings pp. 523–527. Ohio Biological Survey: Columbus (Ohio).

[H98] Halliday, R. B. 1998. Mites of Australia: A checklist and bibliography. CSIRO Publishing: Collingwood.

[RFF05] Rosso de Ferradás, B., & H. R. Fernández. 2005. Elenco y biogeografía de los ácaros acuáticos (Acari, Parasitengona, Hydrachnidia) de Sudamérica. Graellsia 61 (2): 181–224.

[S86] Schwoerbel, J. 1986. Acari: “Hydrachnellae”. In: Botosaneanu, L. (ed.) Stygofauna Mundi: A Faunistic, Distributional, and Ecological Synthesis of the World Fauna inhabiting Subterranean Waters (including the Marine Interstitial) pp. 652–696. E. J. Brill/Dr W. Backhuys: Leiden.

Hygrobates

Hygrobates fluviatilis, photographed by Nigrico.

Belongs within: Hygrobatidae.

Hygrobates is a genus of mostly soft-bodied water mites with the capitulum fused to coxae I and often with a ventral projection on the palpfemur and ventral denticles on the palpgenu.

<==Hygrobates Koch 1837 RFF05
    |--H. (Hygrobates) RFF05
    |    |--H. (H.) ampliatus Viets 1936 RFF05
    |    |    |--H. a. ampliatus RFF05
    |    |    |--H. a. interpositus Viets 1954 RFF05
    |    |    `--H. a. productus Viets 1936 RFF05
    |    |--H. (H.) amplipalpis Lundblad 1953 RFF05
    |    |--H. (H.) apertus Viets 1936 RFF05
    |    |    |--H. a. apertus RFF05
    |    |    `--H. a. apertulus Viets 1954 RFF05
    |    |--H. (H.) clevamus Cook 1980 RFF05
    |    |--H. (H.) crassipes Lundblad 1937 RFF05
    |    |--H. (H.) curvipalpis Besseling 1949 RFF05
    |    |--H. (H.) dadayi Cook 1967 S86
    |    |--H. (H.) discrepans Lundblad 1937 RFF05
    |    |--H. (H.) diversidentatus Viets 1954 RFF05
    |    |--H. (H.) gracilidens Lundblad 1936 RFF05
    |    |--H. (H.) obtusidens Lundbald 1953 RFF05
    |    |--H. (H.) oxyrhynchus Viets 1954 RFF05
    |    |--H. (H.) phreaticus Cook 1967 S86
    |    |--H. (H.) plaumanni Lundblad 1936 RFF05
    |    |--H. (H.) plebejus Lundblad 1930 RFF05
    |    |    |--H. p. plebejus RFF05
    |    |    `--H. p. tamboensis Lundblad 1953 RFF05
    |    |--H. (H.) plicatus Lundblad 1942 RFF05
    |    |--H. (H.) porosus Lundblad 1938 RFF05
    |    |--H. (H.) procursus Viets 1936 RFF05
    |    |--H. (H.) sterrodermus Lundblad 1953 RFF05
    |    `--H. (H.) triangularis Lundblad 1937 RFF05
    |         |--H. t. triangularis RFF05
    |         `--H. t. lundbladi Viets 1938 RFF05
    |--H. (Hygrobatides Lundblad 1936) RFF05
    |    |--H. (H.) angustiporus Viets 1954 RFF05
    |    |--H. (H.) distendens Lundblad 1953 RFF05
    |    |--H. (H.) longimanus Lundblad 1942 RFF05
    |    |--H. (H.) pachydermis (Lundblad 1936) RFF05
    |    |--H. (H.) rufus (Lundblad 1937) RFF05
    |    `--H. (H.) transversalis Viets 1954 RFF05
    `--H. (Schubartella Viets 1937) RFF05
         |--H. (S.) longipes (Viets 1937) RFF05
         `--H. (S.) paraensis (Besch 1969) RFF05

Hygrobates incertae sedis:
  H. australicus Cook 1986 H98
  H. calliger Piersig 1896 VBB91
  H. calvotus WL09
  H. fluviatilis (Ström 1768) GS91
  H. hamatus Viets 1935 H98
  H. longipalpis (Hermann 1804) PH91
  H. longiporus CS91
  H. neocalliger WL09
  H. nigromaculatus Lebert 1879 D91
  H. salamandrarum WL09
  H. setosus DPD16
  H. trigonicus Koenike 1895 D91

*Type species of generic name indicated

REFERENCES

[CS91] Cicolani, B., & A. Di Sabatino. 1991. Sensitivity of water mites to water pollution. In: Dusbábek, F., & V. Bukva (eds) Modern Acarology: Proceedings of the VIII International Congress of Acarology, held in České Budĕjovice, Czechoslovakia, 6–11 August 1990 vol. 1 pp. 465–474. SPB Academic Publishing: The Hague.

[DPD16] Dabert, M., H. Proctor & J. Dabert. 2016. Higher-level molecular phylogeny of the water mites (Acariformes: Prostigmata: Parasitengonina: Hydrachnidiae). Molecular Phylogenetics and Evolution 101: 75–90.

[D91] Davids, C. 1991. Water mites: the impact of larvae and adults on their host and prey populations. In: Dusbábek, F., & V. Bukva (eds) Modern Acarology: Proceedings of the VIII International Congress of Acarology, held in České Budĕjovice, Czechoslovakia, 6–11 August 1990 vol. 1 pp. 497–501. SPB Academic Publishing: The Hague.

[GS91] Gerecke, R., & J. Schwoerbel. 1991. Water quality and water mites (Acari, Actinedida) in the Upper Danube region, 1959-1984. In: Dusbábek, F., & V. Bukva (eds) Modern Acarology: Proceedings of the VIII International Congress of Acarology, held in České Budĕjovice, Czechoslovakia, 6–11 August 1990 vol. 1 pp. 483–491. SPB Academic Publishing: The Hague.

[H98] Halliday, R. B. 1998. Mites of Australia: A checklist and bibliography. CSIRO Publishing: Collingwood.

[PH91] Punčochár, P., & J. Hrbáček. 1991. Water mites in the plankton of Hubenov Reservoir and their relations to fish stock composition. In: Dusbábek, F., & V. Bukva (eds) Modern Acarology: Proceedings of the VIII International Congress of Acarology, held in České Budĕjovice, Czechoslovakia, 6–11 August 1990 vol. 1 pp. 449–457. SPB Academic Publishing: The Hague.

[RFF05] Rosso de Ferradás, B., & H. R. Fernández. 2005. Elenco y biogeografía de los ácaros acuáticos (Acari, Parasitengona, Hydrachnidia) de Sudamérica. Graellsia 61 (2): 181–224.

[S86] Schwoerbel, J. 1986. Acari: “Hydrachnellae”. In: Botosaneanu, L. (ed.) Stygofauna Mundi: A Faunistic, Distributional, and Ecological Synthesis of the World Fauna inhabiting Subterranean Waters (including the Marine Interstitial) pp. 652–696. E. J. Brill/Dr W. Backhuys: Leiden.

[VBB91] Valdecasas, A. G., A. Baltanás & E. Bello. 1991. A preliminary assessment of single rocks for sampling water mites. In: Dusbábek, F., & V. Bukva (eds) Modern Acarology: Proceedings of the VIII International Congress of Acarology, held in České Budĕjovice, Czechoslovakia, 6–11 August 1990 vol. 1 pp. 503–507. SPB Academic Publishing: The Hague.

[WL09] Walter, D. E., E. E. Lindquist, I. M. Smith, D. R. Cook & G. W. Krantz. 2009. Order Trombidiformes. In: Krantz, G. W., & D. E. Walter (eds) A Manual of Acarology 3^rd ed. pp. 233–420. Texas Tech University Press.

Corticacarus

Venter of idiosoma of male Corticacarus argentinensis, from Goldschmidt & Ramírez Sánchez (2020).

Belongs within: Hygrobatidae.

Corticacarus is a genus of hygrobatid water mites with relatively laterally placed legs IV with the glands on coxae IV placed centrally, posterior to the suture between coxae III and IV, and with a ventral projection on the palpfemur.

<==Corticacarus Lundblad 1936 RFF05 [incl. Notorheobates Imamura 1984 H98]
    |--C. (Corticacarus) RFF05
    |    |--C. (C.) coarctipalpis Lundblad 1942 RFF05
    |    |--C. (C.) coeruleus Lundblad 1942 RFF05
    |    |--C. (C.) columbianus Lundblad 1941 RFF05
    |    |--C. (C.) crassipalpis Lundblad 1942 RFF05
    |    |--C. (C.) davecooki (Cook 1988) RFF05
    |    |--C. (C.) dentipalpis Lundblad 1936 RFF05
    |    |    |--C. d. dentipalpis RFF05
    |    |    `--C. d. schadei Lundblad 1942 RFF05
    |    |--C. (C.) edentulus Lundblad 1942 RFF05
    |    |--C. (C.) exilis Cook 1988 RFF05
    |    |--C. (C.) jemobus Cook 1988 RFF05
    |    |--C. (C.) malleifer Lundblad 1936 RFF05
    |    |    |--C. m. malleifer RFF05
    |    |    `--C. m. affinis Lundblad 1942 RFF05
    |    |--C. (C.) patagonicus Besch 1964 RFF05
    |    |--C. (C.) placophorus Schwoerbel 1986 RFF05
    |    |--C. (C.) pravanus Cook 1988 RFF05
    |    |--C. (C.) rimorus Cook 1988 RFF05
    |    |--C. (C.) rostratus Cook 1988 RFF05
    |    |--C. (C.) sovus Cook 1988 RFF05
    |    |--C. (C.) stukabus Cook 1988 RFF05
    |    `--C. (C.) umadus Cook 1988 RFF05
    |--C. (Corticacarellus Lundblad 1937) RFF05
    |    |--C. (C.) acutidens (Lundblad 1938) RFF05
    |    |--C. (C.) labialis Lundblad 1936 RFF05
    |    |--C. (C.) latidens (Viets 1953) RFF05
    |    `--C. (C.) vietsi Lundblad 1953 RFF05
    |--C. (Corticacarides Lundblad 1941) RFF05
    |    |--C. (C.) nilsoni Besch 1964 RFF05
    |    `--C. (C.) plaumanni Lundblad 1941 RFF05
    |--C. (Lundbladacarus Motas & Tanasachi 1960) RFF05
    |    |--C. (L.) anchistus Cook 1988 RFF05
    |    |--C. (L.) angelescui Motas & Tanasachi 1960 RFF05
    |    |--C. (L.) argentinensis Cook 1988 RFF05
    |    |--C. (L.) brundini Besch 1964 RFF05
    |    |--C. (L.) chilensis Viets 1967 RFF05
    |    |--C. (L.) divisus Cook 1988 RFF05
    |    |--C. (L.) icotus Cook 1988 RFF05
    |    |--C. (L.) penai Cook 1988 RFF05
    |    |--C. (L.) schusteri Viets 1977 RFF05
    |    `--C. (L.) xystus Besch 1964 RFF05
    |--C. (Paracorticacarus Lundblad 1953) RFF05
    |    |--C. (P.) brassanus Cook 1980 RFF05
    |    |--C. (P.) brogavus Cook 1980 RFF05
    |    |--C. (P.) coldomus Cook 1980 RFF05
    |    |--C. (P.) copinae Rosso de Ferradás 1991 RFF05
    |    |--C. (P.) curvirostris Lundblad 1953 RFF05
    |    |--C. (P.) longirostris Lundblad 1953 RFF05
    |    |--C. (P.) minutissimus (Lundblad 1953) RFF05
    |    |--C. (P.) motasi Lundblad 1953 RFF05
    |    |--C. (P.) multiscutus Lundblad 1953 RFF05
    |    |    |--C. m. multiscutus RFF05
    |    |    |--C. m. divisus Lundblad 1953 RFF05
    |    |    `--C. m. reductus Lundblad 1953 RFF05
    |    |--C. (P.) noodti Besch 1964 RFF05
    |    |--C. (P.) schwoerbeli Cook 1980 RFF05
    |    |--C. (P.) setipes Lundblad 1953 RFF05
    |    |--C. (P.) smithi Cook 1980 RFF05
    |    |--C. (P.) suchezae Fernández & Rosso de Ferradás 2001 RFF05
    |    `--C. (P.) testudo Lundblad 1953 RFF05
    |         |--C. t. testudo RFF05
    |         `--C. t. aberratus Lundblad 1953 RFF05
    |--C. (Polycorticacarellus Lundblad 1953) RFF05
    |    |--C. (P.) multiporus Lundblad 1953 RFF05
    |    `--C. (P.) pereirai Cook 1980 RFF05
    `--C. (Tetracorticacarellus Lundblad 1941) RFF05
         `--C. (T.) scutatus Lundblad 1941 RFF05

Corticacarus incertae sedis:
  C. aligenitalis Cook 1986 H98
  C. aloonus Cook 1986 H98
  C. angulicoxalis Viets 1978 H98
  C. australicus Viets 1978 H98
  C. cooki (Imamura 1984) [=Notorheobates cooki] H98
  C. cramerae Cook 1986 H98
  C. gennadus Cook 1986 H98
  C. gracilipalpis Viets 1978 H98
  C. hirsutus Viets 1978 H98
  C. incurvatus Viets 1953 RFF05
  C. longipalpis Cook 1986 H98
  C. nabidus Cook 1986 H98
  C. pachydermis Cook 1986 H98
  C. parvus Cook 1986 H98
  C. prasadi Cook 1986 H98
  C. tadilopus Cook 1986 h98
  C. tasmanicus Cook 1986 H98
  C. togalus Cook 1986 H98
  C. victorianus Viets 1978 [=C. victoriensis (l. c.)] H98

*Type species of generic name indicated

REFERENCES

[H98] Halliday, R. B. 1998. Mites of Australia: A checklist and bibliography. CSIRO Publishing: Collingwood.

[RFF05] Rosso de Ferradás, B., & H. R. Fernández. 2005. Elenco y biogeografía de los ácaros acuáticos (Acari, Parasitengona, Hydrachnidia) de Sudamérica. Graellsia 61 (2): 181–224.

Australiobates

Venter of female Australiobates gomorus, from Goldschmidt & Ramírez Sánchez (2020).

Belongs within: Hygrobatidae.

Australiobates is a genus of hygrobatid water mites bearing a posterior process on the capitulum, found in Australasia, South Africa and South America.

<==Australiobates Lundblad 1941 RFF05
    |--A. cekalovici Besch 1964 RFF05
    |--A. curtipalpis Cook 1988 RFF05
    |--A. emalus Cook 1986 H98
    |--A. gledhilli Cook 1986 H98
    |--A. gomorus Cook 1988 RFF05
    |--A. klaasseni Besch 1964 RFF05
    |--A. linderi Lundblad 1941 H98
    |--A. litatus Cook 1988 RFF05
    |--A. mutatus Viets 1978 H98
    |--A. neolinderi Cook 1986 H98
    |--A. ogavus Cook 1988 RFF05
    |--A. ombatus Cook 1986 H98
    |--A. perplexus Cook 1986 H98
    |--A. queenslandensis Cook 1986 H98
    |--A. radagus Cook 1986 H98
    |--A. vantermus Cook 1988 RFF05
    |--A. ventriscutatus Cook 1986 H98
    |--A. vimorus Cook 1986 H98
    `--A. violaceus Lundblad 1941 H98

*Type species of generic name indicated

REFERENCES

[H98] Halliday, R. B. 1998. Mites of Australia: A checklist and bibliography. CSIRO Publishing: Collingwood.

[RFF05] Rosso de Ferradás, B., & H. R. Fernández. 2005. Elenco y biogeografía de los ácaros acuáticos (Acari, Parasitengona, Hydrachnidia) de Sudamérica. Graellsia 61 (2): 181–224.

Aspidiobates

Aspidiobates sp., copyright Melissa Carew.

Belongs within: Hygrobatidae.

Aspidiobates is a genus of water mites found in Australasia and South America characterised by a dorsal shield of one large anterior platelet and multiple smaller posterior platelets, three or four pairs of genital acetabula, and legs without swimming hairs.

<==Aspidiobates Lundblad 1941 RFF05
    |--A. aethes Harvey & Cook 1988 H98
    |--A. bidewel Harvey & Cook 1988 H98
    |--A. geometricus Cook 1986 H98
    |--A. harveyi Cook 1988 RFF05
    |--A. pilbara Harvey 1988 H98
    |--A. scutatus Lundblad 1941 H98
    |--A. similis Cook 1986 H98
    |--A. spatiosus Viets 1969 H98
    |--A. unidorsalis Smit 1942 H98
    `--A. wittenoom Harvey 1988 H98

*Type species of generic name indicated

REFERENCES

[H98] Halliday, R. B. 1998. Mites of Australia: A checklist and bibliography. CSIRO Publishing: Collingwood.

[RFF05] Rosso de Ferradás, B., & H. R. Fernández. 2005. Elenco y biogeografía de los ácaros acuáticos (Acari, Parasitengona, Hydrachnidia) de Sudamérica. Graellsia 61 (2): 181–224.

Omartacaridae

Venter of Omartacarus elongatus, from Cook (1963).

Belongs within: Hygrobatoidea.

The Omartacaridae are a group of interstitial water mites with a disjunct distribution in southern Asia, Australia and the Americas. Species have a soft integument, somewhat elongated idiosoma, all coxal plates grouped together with the medial edges of anterior coxal group noticeably longer than those of the posterior group, and suture lines between coxal plates III–IV extending at right angles to the midline or slightly posterolaterally (Walter et al. 2009).

Omartacaridae RFF05
    |--Maharashtracarus Cook 1967 [Maharashtracarinae] S86
    |    |--M. neotropicus Cook 1980 S86
    |    `--M. phreaticus Cook 1967 S86
    `--Omartacarus Cook 1963 [Omartacarinae] RFF05
         |  i. s.: O. australicus Viets 1978 H98
         |         O. blythi Cook 1986 H98
         |         O. bobidus Cook 1986 H98
         |         O. ferradasae Cook 1988 RFF05
         |         O. lomedus Cook 1986 H98
         |         O. paraelongatus Fernández & Grosso 1991 RFF05
         |         O. pauciporus Cook 1986 H98
         |         O. tucumanensis Fernández 1992 RFF05
         |--O. (Omartacarus) S86
         |    |--O. (O.) elongatus Cook 1963 S86
         |    `--O. (O.) motasi Cook 1980 S86
         `--O. (Omartacarellus Cook 1974) S86
              `--O. (O.) brevipalpis Cook 1974 S86

*Type species of generic name indicated

REFERENCES

[H98] Halliday, R. B. 1998. Mites of Australia: A checklist and bibliography. CSIRO Publishing: Collingwood.

[RFF05] Rosso de Ferradás, B., & H. R. Fernández. 2005. Elenco y biogeografía de los ácaros acuáticos (Acari, Parasitengona, Hydrachnidia) de Sudamérica. Graellsia 61 (2): 181–224.

[S86] Schwoerbel, J. 1986. Acari: “Hydrachnellae”. In: Botosaneanu, L. (ed.) Stygofauna Mundi: A Faunistic, Distributional, and Ecological Synthesis of the World Fauna inhabiting Subterranean Waters (including the Marine Interstitial) pp. 652–696. E. J. Brill/Dr W. Backhuys: Leiden.

Walter, D. E., E. E. Lindquist, I. M. Smith, D. R. Cook & G. W. Krantz. 2009. Order Trombidiformes. In: Krantz, G. W., & D. E. Walter (eds) A Manual of Acarology 3^rd ed. pp. 233–420. Texas Tech University Press.

Austraturus

Austraturus tumidipalpis, from Viets (1978).

Belongs within: Aturidae.

Austraturus is an Australian genus of aturid water mites with the dorsal shield divided into seven platelets, and with the pedipalp having several small ventral projections on the femur and paired ventral setae placed subdistally on the tibia.

<==Austraturus Viets 1978 H98
    |--A. concavus Cook 1986 H98
    |--A. coomensis Imamura 1984 H98
    |--A. elongatus Cook 1986 H98
    |--A. exilis Cook 1986 H98
    |--A. longipalpis Cook 1986 H98
    |--A. projectus Cook 1986 H98
    |--A. tumidipalpis Viets 1978 H98
    |--A. uncicoxalis Cook 1986 [=A. unicoxalis (l. c.)] H98
    |--A. victoriensis Cook 1986 H98
    `--A. vietsi Cook 1986 H98

*Type species of generic name indicated

REFERENCES

[H98] Halliday, R. B. 1998. Mites of Australia: A checklist and bibliography. CSIRO Publishing: Collingwood.

Axonopsella

Axonopsella misionum male, from Rosso de Ferradás (1995).

Belongs within: Aturidae.

Axonopsella is a genus of aturid water mites found in Australasia and South America. The dorsal plate is comprised of a single shield and projections from coxa IV cover the openings of legs IV.

<==Axonopsella Lundblad 1930 RFF05
    |--A. (Chilaxonopsella Cook 1988) RFF05
    |    |--A. (C.) barri Cook 1988 RFF05
    |    |--A. (C.) chilensis Cook 1988 RFF05
    |    |--A. (C.) doloma Cook 1988 RFF05
    |    `--A. (C.) tanda Cook 1988 RFF05
    |--A. (Coaxonopsella Cook 1980) RFF05
    |    `--A. (C.) jujuyensis Cook 1980 RFF05
    |--A. (Humboldtaxonopsella Orghidan & Gruia 1981) RFF05
    |    `--A. (H.) linaresi Orghidan & Gruia 1981 RFF05
    |--A. (Luciaxonopsella Rosso de Ferradás 1995) RFF05
    |    `--A. (L.) misionum Rosso de Ferradás 1995 RFF05
    |--A. (Neaxonopsella Lundblad 1937) RFF05
    |    |--A. (N.) argentinensis Cook 1980 RFF05
    |    |--A. (N.) caripense Orghidan & Gruia 1981 RFF05
    |    |--A. (N.) hamatoides Lundblad 1937 RFF05
    |    |--A. (N.) mamillata Viets 1954 RFF05
    |    `--A. (N.) trifida Lundblad 1937 RFF05
    |--A. (Paraxonopsella Lundblad 1937) RFF05
    |    |--A. (P.) bifida Lundblad 1937 RFF05
    |    |--A. (P.) filunguis Lundblad 1937 RFF05
    |    `--A. (P.) hamata Lundblad 1936 RFF05
    `--A. (Rostaxonopsella Orghidan & Gruia 1987) RFF05
         `--A. (R.) nana Orghidan & Gruia 1987 RFF05

Axonopsella incertae sedis:
  A. affinis Lundblad 1943 RFF05
  A. andina Lundblad 1953 RFF05
  A. australica Cook 1986 H98
  A. bakeri WL09
  A. cerucha Cook 1986 H98
  A. chordella Cook 1986 H98
  A. davidsi Cook 1986 H98
  A. diamphida Cook 1986 H98
  A. diatropa Cook 1986 H98
  A. eremita Cook 1986 H98
  A. expansipes Cook 1986 H98
    |--A. e. expansipes H98
    `--A. e. zodala Cook 1986 H98
  A. ferradasae Cook 1986 H98
  A. fragala Cook 1986 H98
  A. groota Cook 1986 H98
  A. harveyi Cook 1986 H98
  A. henrechi Orghidan & Gruia 1981 RFF05
  A. hopkinsi Cook 1986 H98
  A. interstitialis Cook 1986 H98
  A. jacunda Cook 1986 H98
  A. koola Cook 1986 H98
  A. laeta Cook 1986 H98
  A. magna Cook 1986 H98
  A. oldora Cook 1986 H98
  A. plaumanni Lundblad 1936 RFF05
  A. ricala Cook 1986 H98
  A. spinigera Lundblad 1930 RFF05
  A. thori Cook 1986 H98
  A. trimaculata Cook 1986 H98
  A. truza Cook 1986 H98
  A. vaginalis Viets 1936 RFF05
  A. vietsi Lundblad 1937 RFF05
  A. walteri Cook 1986 H98
  A. wasta Cook 1986 H98
  A. wootoomba Cook 1986 H98
  A. zotela Cook 1986 H98

*Type species of generic name indicated

REFERENCES

[H98] Halliday, R. B. 1998. Mites of Australia: A checklist and bibliography. CSIRO Publishing: Collingwood.

[RFF05] Rosso de Ferradás, B., & H. R. Fernández. 2005. Elenco y biogeografía de los ácaros acuáticos (Acari, Parasitengona, Hydrachnidia) de Sudamérica. Graellsia 61 (2): 181–224.

[WL09] Walter, D. E., E. E. Lindquist, I. M. Smith, D. R. Cook & G. W. Krantz. 2009. Order Trombidiformes. In: Krantz, G. W., & D. E. Walter (eds) A Manual of Acarology 3^rd ed. pp. 233–420. Texas Tech University Press.

Albia

Albia sp., from Water Mites of North America.

Belongs within: Aturidae.

Albia is a cosmopolitan genus of aturid water mites with the dorsal shield composed of a single plate but without projections covering the openings of legs IV.

<==Albia Thor 1899 RFF05
    |  i. s.: A. australica Cook 1986 H98
    |         A. brokenensis Smit 1992 H98
    |         A. lundbladi Cook 1986 H98
    |         A. neogaea WL09
    |         A. rectifrons Viets 1935 H98
    |--A. (Anchistalbia Cook 1974) RFF05
    |    `--A. (A.) nobilis Lundblad 1936 RFF05
    `--A. (Dentalbia Cook 1967) S86
         |--A. (D.) dentipalpis Cook 1967 S86
         `--A. (D.) phreatica Cook 1967 S86

*Type species of generic name indicated

REFERENCES

[H98] Halliday, R. B. 1998. Mites of Australia: A checklist and bibliography. CSIRO Publishing: Collingwood.

[RFF05] Rosso de Ferradás, B. & H. R. Fernández. 2005. Elenco y biogeografía de los ácaros acuáticos (Acari, Parasitengona, Hydrachnidia) de Sudamérica. Graellsia 61 (2): 181–224.

[S86] Schwoerbel, J. 1986. Acari: “Hydrachnellae”. In: Botosaneanu, L. (ed.) Stygofauna Mundi: A Faunistic, Distributional, and Ecological Synthesis of the World Fauna inhabiting Subterranean Waters (including the Marine Interstitial) pp. 652–696. E. J. Brill/Dr W. Backhuys: Leiden.

[WL09] Walter, D. E., E. E. Lindquist, I. M. Smith, D. R. Cook & G. W. Krantz. 2009. Order Trombidiformes. In: Krantz, G. W., & D. E. Walter (eds) A Manual of Acarology 3^rd ed. pp. 233–420. Texas Tech University Press.

Frontipodopsis

Frontipodopsis sp., from Goldschmidt et al. (2022).

Belongs within: Hygrobatoidea.

Frontipodopsis is a genus of water mites with a disjunct distribution in southern Eurasia, Australia and the Americas. They are characterised by a strongly laterally compressed idiosoma, and dorsoventrally expanded and laterally flattened segments on leg IV (Walter et al. 2009).

Frontipodopsis Walter 1919 [Frontipodopsidae, Frontipodopsinae] RFF05
    |--F. australica Cook 1986 H98
    |--F. chilensis Cook 1988 RFF05
    |--F. curvipalpis Cook 1986 H98
    |--F. mesoamericana Cook 1980 S86
    |--F. nearctica Cook 1963 S86
    |--F. obscura Cook 1986 H98
    |--F. reticulatifrons Szalay 1945 S86
    |    |--F. r. reticulatifrons S86
    |    |--F. r. indica Cook 1967 S86
    |    |--F. r. okinawensis Imamura 1957 S86
    |    `--F. r. persicus Schwoerbel & Sepas. 1980 S86
    |--F. staheli Walter 1919 RFF05
    |    |--F. s. staheli RFF05
    |    `--F. s. mandibularis Lundblad 1943 RFF05
    `--F. sudafricanus WL09

*Type species of generic name indicated

REFERENCES

[H98] Halliday, R. B. 1998. Mites of Australia: A checklist and bibliography. CSIRO Publishing: Collingwood.

[RFF05] Rosso de Ferradás, B. & H. R. Fernández. 2005. Elenco y biogeografía de los ácaros acuáticos (Acari, Parasitengona, Hydrachnidia) de Sudamérica. Graellsia 61 (2): 181–224.

[S86] Schwoerbel, J. 1986. Acari: “Hydrachnellae”. In: Botosaneanu, L. (ed.) Stygofauna Mundi: A Faunistic, Distributional, and Ecological Synthesis of the World Fauna inhabiting Subterranean Waters (including the Marine Interstitial) pp. 652–696. E. J. Brill/Dr W. Backhuys: Leiden.

[WL09] Walter, D. E., E. E. Lindquist, I. M. Smith, D. R. Cook & G. W. Krantz. 2009. Order Trombidiformes. In: Krantz, G. W., & D. E. Walter (eds) A Manual of Acarology 3^rd ed. pp. 233–420. Texas Tech University Press.

Recifella

Dorsum and palp of female Recifella excavata, from Rosso de Ferradás & Böttger (1997).

Belongs within: Unionicolidae.

Recifella is a genus of unionicolid water mites found in Australia and South America, with complete dorsal and ventral shields and three or four pairs of glandularia on the dorsal shield.

<==Recifella Viets 1935 RFF05
    |--R. (Recifella) RFF05
    |    |--R. (R.) angulipes (Lundblad 1938) RFF05
    |    |--R. (R.) clavata (Lundblad 1937) RFF05
    |    |--R. (R.) gibberipalpis (Lundblad 1936) RFF05
    |    |--R. (R.) laminipes (Viets 1935) RFF05
    |    |--R. (R.) latipes (Lundblad 1936) RFF05
    |    |--R. (R.) lundbladi (Besseling 1949) RFF05
    |    `--R. (R.) sinuatipes (Lundblad 1941) RFF05
    |--R. (Eorecifella Cook 1980) RFF05
    |    |--R. (E.) balteata (Lundblad 1943) RFF05
    |    |--R. (E.) cruciata (Lundblad 1936) RFF05
    |    |    |--R. c. cruciata RFF05
    |    |    `--R. c. brevis (Lundblad 1943) RFF05
    |    |--R. (E.) diplothoracica (Lundblad 1937) RFF05
    |    |--R. (E.) elliptica (Walter 1919) RFF05
    |    |    |--R. e. elliptica RFF05
    |    |    |--R. e. amabilis (Lundblad 1943) RFF05
    |    |    |--R. e. schadei (Lundblad 1943) RFF05
    |    |    `--R. e. validipalpis (Lundblad 1937) RFF05
    |    |--R. (E.) excavata (Lundblad 1936) RFF05
    |    |--R. (E.) excavatoides (Viets 1975) RFF05
    |    |--R. (E.) opimipalpis (Viets 1977) RFF05
    |    |--R. (E.) pusilla (Lundblad 1937) RFF05
    |    `--R. (E.) undulata (Lunblad 1936) RFF05
    `--R. (Monokoenikea Lundblad 1941) RFF05
         `--R. (M.) melini (Lundblad 1930) RFF05

Recifella incertae sedis:
  R. agnosta Cook 1986 H98
  R. australica Cook 1986 H98
  R. baltoona Cook 1986 H98
  R. bifida Cook 1986 H98
  R. columbiana (Lundblad 1953) RFF05
  R. doomba Cook 1986 H98
  R. flagellata Cook 1986 H98
  R. inka (Lundblad 1944) RFF05
  R. paucipora (Lundblad 1943) RFF05
  R. pectinatus Smit 1992 H98
  R. plana Cook 1986 H98
  R. pulchra (Lundblad 1937) RFF05
  R. szalayi Cook 1986 H98
  R. tinka (Viets 1975) [=Koenikea tinka] H98
  R. triradiata Cook 1986 H98
  R. truncata (Besseling 1949) RFF05
  R. umala Cook 1986 H98
  R. vonjama Cook 1986 H98

*Type species of generic name indicated

REFERENCES

[H98] Halliday, R. B. 1998. Mites of Australia: A checklist and bibliography. CSIRO Publishing: Collingwood.

[RFF05] Rosso de Ferradás, B., & H. R. Fernández. 2005. Elenco y biogeografía de los ácaros acuáticos (Acari, Parasitengona, Hydrachnidia) de Sudamérica. Graellsia 61 (2): 181–224.

Koenikea

Koenikea sp., copyright David E. Reed.

Belongs within: Unionicolidae.

Koenikea is a genus of unionicolid water mites found in Australia and the Americas, with complete dorsal and ventral shields and five or six pairs of glandularia on the dorsal shield.

<==Koenikea Wolcott 1900 RFF05
    |--K. (Koenikea) V77
    |    |--K. (K.) acuta Lundblad 1943 RFF05
    |    |    |--K. a. acuta RFF05
    |    |    `--K. a. dentata Viets 1975 RFF05
    |    |--K. (K.) affinis Lundblad 1938 RFF05
    |    |--K. (K.) angustipalpis Viets 1975 RFF05
    |    |--K. (K.) bicornis Lundblad 1941 RFF05
    |    |--K. (K.) bipapillata Lundblad 1941 RFF05
    |    |--K. (K.) biscutata Daday 1905 RFF05
    |    |--K. (K.) brunnea Lundblad 1937 RFF05
    |    |--K. (K.) coerulescens Lundblad 1936 RFF05
    |    |    |--K. c. coerulescens RFF05
    |    |    |--K. c. rosea Lundblad 1936 RFF05
    |    |    `--K. c. vietsi Lundblad 1937 RFF05
    |    |--K. (K.) convexa Daday 1905 (n. d.) RFF05
    |    |--K. (K.) corniculata Viets 1975 RFF05
    |    |--K. (K.) falcaria Viets 1977 V77
    |    |--K. (K.) fluctuata Viets 1975 RFF05
    |    |--K. (K.) furcatimaculata Lundblad 1936 RFF05
    |    |--K. (K.) hastata Viets 1977 V77
    |    |--K. (K.) lata Viets 1975 RFF05
    |    |--K. (K.) longipes Lundblad 1936 RFF05
    |    |--K. (K.) medioposita Lundblad 1936 RFF05
    |    |    |--K. m. medioposita V77
    |    |    |--K. m. asterifera Lundblad 1943 V77
    |    |    `--K. m. transversa Lundblad 1943 V77
    |    |--K. (K.) octopapillata Lundblad 1930 RFF05
    |    |--K. (K.) opimipalpis Viets 1977 V77
    |    |--K. (K.) pallida Lundblad 1937 RFF05
    |    |--K. (K.) papillata Viets 1975 RFF05
    |    |--K. (K.) paraguayensis Rosso de Ferradás & Böttger 1998 RFF05
    |    |--K. (K.) plicata Lundblad 1936 RFF05
    |    |--K. (K.) procursa Viets 1977 V77
    |    |--K. (K.) quadricornuta Walter 1919 RFF05
    |    |--K. (K.) rectangulata Lundblad 1930 RFF05
    |    |--K. (K.) rectirostris Lundblad 1936 RFF05
    |    |    |--K. r. rectirostris RFF05
    |    |    |--K. r. crassoalata Lundblad 1943 RFF05
    |    |    |--K. r. ramosa Lundblad 1943 RFF05
    |    |    `--K. r. retractidens Lundblad 1938 RFF05
    |    |--K. (K.) retrocornuta Rosso de Ferradás & Böttger 1997 RFF05
    |    |--K. (K.) siolii Viets 1975 RFF05
    |    |--K. (K.) triangularis Lundblad 1938 RFF05
    |    |--K. (K.) unguiculata Lundblad 1943 RFF05
    |    `--K. (K.) victoriola Rosso de Ferradás 1998 RFF05
    |--K. (Diplokoenikea Lundblad 1936) RFF05
    |    |--K. (D.) clavigera Lundblad 1943 RFF05
    |    |    |--K. c. clavigera RFF05
    |    |    `--K. c. assimilis Lundblad 1943 RFF05
    |    |--K. (D.) curvirostris Walter 1919 RFF05
    |    |--K. (D.) flagellaris Lundblad 1938 RFF05
    |    |--K. (D.) grossa Lundblad 1938 RFF05
    |    |--K. (D.) pectinata Lundblad 1936 RFF05
    |    |--K. (D.) pectinifera Lundblad 1938 RFF05
    |    |--K. (D.) quinquemaculata Lundblad 1943 RFF05
    |    |--K. (D.) scutica Viets 1977 V77
    |    `--K. (D.) simulans Lundblad 1943 RFF05
    |--K. (Jurucuia Viets 1954) RFF05
    |    `--K. (J.) oblongata (Viets 1954) RFF05
    |--K. (Koenikella Lundblad 1936) RFF05
    |    |--K. (K.) crassipalpis Lundblad 1936 RFF05
    |    `--K. (K.) cyanozona Lundblad 1937 RFF05
    |--K. (Neokoenikea Lundblad 1936) RFF05
    |    `--K. (N.) armipes Lundblad 1936 RFF05
    |--K. (Notomideopsis Wolcott 1905) RFF05
    |    |--K. (N.) acanthifera Lundblad 1953 RFF05
    |    |--K. (N.) acutidens Lundblad 1943 RFF05
    |    |--K. (N.) brasiliensis Lundblad 1930 RFF05
    |    |--K. (N.) chilensis Cook 1988 RFF05
    |    |--K. (N.) circumcincta Lundblad 1937 RFF05
    |    |--K. (N.) consimilis Viets 1975 RFF05
    |    |--K. (N.) crassa Lundblad 1937 RFF05
    |    |--K. (N.) curicola Viets 1975 RFF05
    |    |--K. (N.) curvipes Lundblad 1938 RFF05
    |    |--K. (N.) diabolica Viets 1975 RFF05
    |    |--K. (N.) elegans Lundblad 1938 RFF05
    |    |--K. (N.) hamulata Lundblad 1936 RFF05
    |    |--K. (N.) imitata Lundblad 1930 RFF05
    |    |--K. (N.) lunipes Lundblad 1938 RFF05
    |    |--K. (N.) nordenskioldi Cook 1980 RFF05
    |    |--K. (N.) obscura Lundblad 1937 RFF05
    |    |--K. (N.) rectipes Lundblad 1943 RFF05
    |    |--K. (N.) soona Cook 1988 RFF05
    |    |--K. (N.) spinosa Daday 1905 RFF05
    |    `--K. (N.) tigrina Lundblad 1936 RFF05
    |         |--K. t. tigrina RFF05
    |         `--K. t. crassiseta Lundblad 1943 RFF05
    |--K. (Parakoenikea Lundblad 1936) RFF05
    |    |--K. (P.) curvipalpis Lundblad 1936 RFF05
    |    `--K. (P.) scutellum Besch 1969 RFF05
    |--K. (Pseudokoenikea Lundblad 1941) RFF05
    |    |--K. (P.) horrida Lundblad 1930 RFF05
    |    `--K. (P.) rutae Lundblad 1930 RFF05
    |         |--K. r. rutae RFF05
    |         `--K. r. furcaticornis Viets 1956 RFF05
    |--K. (Sespekoenikea Cook 1974) S86
    |    `--K. (S.) expansipalpis Cook 1974 S86
    `--K. (Tamboella Lundblad 1953) RFF05
         `--K. (T.) multiseta Lundblad 1953 RFF05

Koenikea incertae sedis:
  K. australica Lundblad 1941 H98
  K. balteata Lundblad 1943 V77
  K. bispina WL09
  K. branacha Cook 1986 H98
  K. colorata Viets 1980 [=Recifella colorata] H98
  K. crinita Cook 1986 H98
  K. curtiseta Cook 1986 H98
  K. decapora Cook 1986 H98
  K. distans Viets 1980 H98
  K. elliptica Walter 1919 V77
    |--K. e. elliptica V77
    |--K. e. amabilis Lundblad 1943 V77
    |--K. e. schadei Lundblad 1943 V77
    `--K. e. validipalpis Lundblad 1937 V77
  K. haldemani PW01
  K. jacunda Cook 1986 H98
  K. lemba Cook 1986 H98
  K. longiuscula Lundblad 1936 RFF05
  K. pauciseta Cook 1980 RFF05
  K. piota Cook 1986 H98
  K. plaumanni Lundblad 1937 RFF05
  K. pseudodistans Cook 1986 H98
  K. pulchra Lundblad 1937 V77
  K. purpurea Smit 1992 H98
  K. rodosa Cook 1986 H98
  K. rutala Cook 1986 H98
  K. schubarti Viets 1977 V77
  K. sorpresa Cook 1986 H98
  K. spinipes PW01
  K. timmsi Viets 1977 H98
  K. verrucosa Lundblad 1947 H98
  K. voloma Cook 1986 H98

*Type species of generic name indicated

REFERENCES

[H98] Halliday, R. B. 1998. Mites of Australia: A checklist and bibliography. CSIRO Publishing: Collingwood.

[PW01] Proctor, H., & K. Wilkinson. 2001. Coercion and deceit: water mites (Acari: Hydracarina) and the study of intersexual conflict. In: Halliday, R. B., D. E. Walter, H. C. Proctor, R. A. Norton & M. J. Colloff (eds) Acarology: Proceedings of the 10^th International Congress pp. 155–169. CSIRO Publishing: Melbourne.

[RFF05] Rosso de Ferradás, B., & H. R. Fernández. 2005. Elenco y biogeografía de los ácaros acuáticos (Acari, Parasitengona, Hydrachnidia) de Sudamérica. Graellsia 61 (2): 181–224.

[S86] Schwoerbel, J. 1986. Acari: “Hydrachnellae”. In: Botosaneanu, L. (ed.) Stygofauna Mundi: A Faunistic, Distributional, and Ecological Synthesis of the World Fauna inhabiting Subterranean Waters (including the Marine Interstitial) pp. 652–696. E. J. Brill/Dr W. Backhuys: Leiden.

[V77] Viets, K. O. 1977. Neue Koenikea-Arten aus Brasilien (Acari: Hydrachnellae: Unionicolidae). Senckenbergiana Biologica 57 (4-6): 341–358.

[WL09] Walter, D. E., E. E. Lindquist, I. M. Smith, D. R. Cook & G. W. Krantz. 2009. Order Trombidiformes. In: Krantz, G. W., & D. E. Walter (eds) A Manual of Acarology 3^rd ed. pp. 233–420. Texas Tech University Press.

Piona

Male Piona nodata, from here.

Belongs within: Pionidae.

Piona is a genus of pionid water mites in which males bear a dorsal concavity on the tibia of leg IV containing several stout, peg-like setae.

<==Piona Koch 1842 RFF05
    |--P. aculeatipalpis Viets 1954 RFF05
    |--P. acutidens Lundblad 1941 RFF05
    |--P. alpicola WP99
    |--P. amazonica Viets 1954 RFF05
    |--P. anisitsi Daday 1905 RFF05
    |--P. atrorubra Lundblad 1937 RFF05
    |--P. australica Viets 1980 H98
    |--P. carnea (Koch 1836) L91
    |--P. chilensis Viets 1968 RFF05
    |--P. coccinea (Koch 1836) PH91
    |--P. confinis Lundblad 1943 RFF05
    |--P. conglobata BK91
    |--P. constricta PH91
    |--P. cumberlandensis (Rainbow 1906) (see below for synonymy) H98
    |--P. davidcooki Rosso de Ferradás & Böttger 1997 RFF05
    |--P. deformis Lundblad 1941 RFF05
    |--P. dejecta Viets 1937 RFF05
    |--P. dissipata Lundblad 1938 RFF05
    |--P. erratica Marshall 1940 RFF05
    |--P. fissa Viets 1940 RFF05
    |--P. indica WL09
    |--P. interrupta WL09
    |--P. juncta Lundblad 1936 RFF05
    |    |--P. j. juncta RFF05
    |    `--P. j. inconspicua Lundblad 1943 RFF05
    |--P. lapointei WL09
    |--P. limnetica WL09
    |--P. longipalpis WP99
    |--P. marchanti Harvey 1996 H98
    |--P. mitchelli WL09
    |--P. murleyi Harvey 1996 H98
    |--P. neumani (Koenike 1883) PH91
    |--P. nodata (Müller 1776) L91
    |--P. oxyura Viets 1954 RFF05
    |--P. panniculata Viets 1940 RFF05
    |--P. paucipora (Thor 1897) PH91
    |--P. plaumanni Lundblad 1943 RFF05
    |--P. plenipalpis Viets 1954 RFF05
    |--P. praegracilis Viets 1937 RFF05
    |--P. (Tetrapiona) pseudouncata (Piersig 1906) SL71
    |--P. punctatissima Viets 1954 RFF05
    |--P. puripalpis Viets 1984 H98
    |--P. pusilla (Neumann 1875) PH91
    |--P. rectimarginata Viets 1954 RFF05
    |--P. reflexidens Viets 1940 RFF05
    |--P. robustipalpis Viets 1954 RFF05
    |--P. rotunda (Kramer 1879) RFF05
    |    |--P. r. rotunda RFF05
    |    |--P. r. acutipes Viets 1954 RFF05
    |    `--P. r. rotundiformis Lundblad 1938 RFF05
    |--P. rotundoides (Thor 1897) PH91
    |--P. serratiseta Viets 1956 RFF05
    |--P. setipes Cook 1980 RFF05
    |--P. sicaria Lundblad 1937 RFF05
    |--P. stjoerdalensis (Thor 1897) PH91
    |--P. sudamericana Viets 1910 RFF05
    |--P. tenuipalpis Viets 1954 RFF05
    |--P. thoracica Lundblad 1941 RFF05
    |--P. (Piona) uncataexigua Viets 1949 SL71
    |--P. ungularis Lundblad 1943 RFF05
    |--P. unidens Viets 1954 RFF05
    |--P. variabilis BK91
    `--P. venezuelensis Viets 1956 RFF05

Piona cumberlandensis (Rainbow 1906) [=Atax cumberlandensis, Unionicola cumberlandensis, Unionocola (l. c.) cumberlandensis; incl. Piona uncatiformis Lundblad 1941] H98

*Type species of generic name indicated

REFERENCES

[BK91] Biesiadka, E., & W. Kowalik. 1991. Water mites (Hydracarina) as indicators of trophy and pollution in lakes. In: Dusbábek, F., & V. Bukva (eds) Modern Acarology: Proceedings of the VIII International Congress of Acarology, held in České Budĕjovice, Czechoslovakia, 6–11 August 1990 vol. 1 pp. 475–481. SPB Academic Publishing: The Hague.

[H98] Halliday, R. B. 1998. Mites of Australia: A checklist and bibliography. CSIRO Publishing: Collingwood.

[L91] Leimann, J. 1991. Structure and formation of the sperm package of Piona carnea (Koch, 1836) (Prostigmata, Hydrachnidia), a copulating water mite. In: Dusbábek, F., & V. Bukva (eds) Modern Acarology: Proceedings of the VIII International Congress of Acarology, held in České Budĕjovice, Czechoslovakia, 6–11 August 1990 vol. 2 pp. 449–454. SPB Academic Publishing: The Hague.

[PH91] Punčochár, P., & J. Hrbáček. 1991. Water mites in the plankton of Hubenov Reservoir and their relations to fish stock composition. In: Dusbábek, F., & V. Bukva (eds) Modern Acarology: Proceedings of the VIII International Congress of Acarology, held in České Budĕjovice, Czechoslovakia, 6–11 August 1990 vol. 1 pp. 449–457. SPB Academic Publishing: The Hague.

[RFF05] Rosso de Ferradás, B., & H. R. Fernández. 2005. Elenco y biogeografía de los ácaros acuáticos (Acari, Parasitengona, Hydrachnidia) de Sudamérica. Graellsia 61 (2): 181–224.

[SL71] Spain, A. V., & M. Luxton. 1971. Catalog and bibliography of the Acari of the New Zealand subregion. Pacific Insects Monograph 25: 179–226.

[WL09] Walter, D. E., E. E. Lindquist, I. M. Smith, D. R. Cook & G. W. Krantz. 2009. Order Trombidiformes. In: Krantz, G. W., & D. E. Walter (eds) A Manual of Acarology 3^rd ed. pp. 233–420. Texas Tech University Press.

[WP99] Walter, D. E., & H. C. Proctor. 1999. Mites: Ecology, Evolution and Behaviour. CABI Publishing: Wallingford (UK).