Elaphrinae

Elaphrus lapponicus, from here.


Belongs within: Geadephaga.

The Elaphrinae are a group of carabid beetles characterised by the presence of a setigerous puncture in each scrobe, and two setigerous punctures above each eye (Ball & Bousquet 2001).

Characters (from Ball & Bousquet 2001): Size moderate to large (total length ca. 6-18 mm); head capsule ventrally without antennal grooves; antennal insertions laterad, each between base of mandible and anterior margin of eye; antenna filiform, antennomeres 2-4 not nodose, 2-6 without unusually long setae, eleven antennomeres; labrum with six setae; mandible with seta in scrobe; terminal palpomeres about cylindrical; pterothorax with mesonotal scutellum dorsally visible, not concealed beneath elytra; front coxal cavities closed by insertion of tip of proepimeron each side in cavity of lateral projection of prosternum, middle and hind coxal cavities disjunct-confluent; elytra extended to apex of abdominal sternum VIII, with apical margins subsinuate, striate or not dorsally; metathoracic wings large; front tibia anisochaete, antenna cleaner sulcate, grade B; abdomen with six (II-VII) pregenital sterna normally exposed, sterna III and IV separated by suture; male genitalia with parameres similar in form, and both moderately long, setose preapically and apically; ovipositor with gonocoxae rotated 90° in repose, without ramus, gonocoxa with two articles, gonocoxite 2 without accessory lobe; and reproductive tract without spermatheca 1, with spermatheca 2; secretions of pygidial glands higher saturated acids, discharge by oozing.

<==Elaphrinae [Elaphrini, Elaphritae] B14
    |--Diacheila Motschulsky 1844 BB01
    |--Nebriosoma Castelnau 1867 LM87
    |    `--*N. fallax Castelnau 1867 LM87
    |--Calyptogonia Sloane 1920 LM87
    |    `--*C. atra Sloane 1920 [=C. ater] LM87
    |--Decogmus Sloane 1915 LM87
    |    `--*D. chalybeus Sloane 1915 LM87
    |--Belthisa Bonelli 1810 BB01
    |    |--B. catenaria Brown 1944 BB01
    |    `--B. multipunctata BB01
    |         |--B. m. multipunctata BB01
    |         `--B. m. aurata Fischer von Waldheim 1828 BB01
    `--Elaphrus Fabricius 1775 BB01
         |  i. s.: E. aquaticus L02
         |         E. californicus OM08
         |         E. cupreus C01
         |         E. riparius C01
         |         E. sugai Nakane 1987 I92
         |         E. uliginosus C01
         |         E. viridis Horn 1878 B14
         |--E. (Arctelaphrus Semenov 1926) BB01
         |    `--E. (A.) lapponicus Gyllenhal 1810 BB01
         |--E. (Elaphroterus Semenov 1895) BB01
         `--E. (Neoelaphrus Hatch 1951) BB01

*Type species of generic name indicated

REFERENCES

[BB01] Ball, G. E., & Y. Bousquet. 2001. Carabidae Latreille, 1810. In: Arnett, R. H., Jr & M. C. Thomas (eds) American Beetles vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia pp. 32–132. CRC Press: Boca Raton.

[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature's gems. Ivy Press: Lewes (United Kingdom).

[C01] Csiki, E. 1901. Bogarak [Coleopteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 75–120. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[I92] Iwahashi, J. (ed.) 1992. Reddo Deeta Animaruzu: a pictorial of Japanese fauna facing extinction. JICC: Tokyo.

[L02] Latreille, P. A. 1802. Histoire Naturelle, générale et particulière des crustacés et des insectes vol. 3. Familles naturelles des genres. F. Dufart: Paris.

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

[OM08] Ober, K. A., & D. R. Maddison. 2008. Phylogenetic relationships of tribes within Harpalinae (Coleoptera: Carabidae) as inferred from 28S ribosomal DNA and the wingless gene. Journal of Insect Science 8: 63.

Derodontidae

Derodontus maculatus, copyright Bonnie Ott.


Belongs within: Scirtoidea.

The Derodontidae are a group of small beetles with paired ocelli that feed on fungi.

Characters (from Lawrence & Britton 1991): Head with paired ocelli and series of canals and bridges; antenna eleven-segmented with three-segmented club; pronotum divided into several cells by longitudinal ridges and transverse elevations; elytra with number of deep pits at bases of striae; fore coxae projecting, trochantins concealed, fore coxal cavities externally closed, internally open; mid coxal cavities narrowly separated, partly closed laterally by metepisterna; metasternum with transverse suture; tarsi pentamerous; hind coxal plates well developed; abdomen with five ventrites, none connate, a pair of longitudinal carinae near middle of ventrite I. Larva elongate, slightly tapered posteriorly, with dorsal surfaces granulate, tuberculate and usually with paired processes on all segments except 10th, which is cylindrical and posteroventrally oriented. Head transverse, with no epicranial stem and contiguous frontal arms, and spiracles borne on tubular processes.

Derodontidae B14
    |  i. s.: Peltastica [Peltasticinae] B14
    |           |--P. amurensis Reitter 1879 B14
    |           `--P. tuberculata B14
    |--Laricobius MW15
    `--+--Nothoderodontus MW15
       |    |--N. darlingtoni LB91
       |    `--N. gourlayi LB91
       `--Derodontus [Derodontinae] MW15
            |--D. esotericus B14
            |--D. japonicus B14
            |--D. macularis (Fuss 1850) B14
            |--D. maculatus B14
            `--D. trisignatus B14

*Type species of generic name indicated

REFERENCES

[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature's gems. Ivy Press: Lewes (United Kingdom).

[LB91] Lawrence, J. F., & E. B. Britton. 1991. Coleoptera (beetles). In: CSIRO. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 2 pp. 543–683. Melbourne University Press: Carlton (Victoria).

[MW15] McKenna, D. D., A. L. Wild, K. Kanda, C. L. Bellamy, R. G. Beutel, M. S. Caterino, C. W. Farnum, D. C. Hawks, M. A. Ivie, M. L. Jameson, R. A. B. Leschen, A. E. Marvaldi, J. V. McHugh, A. F. Newton, J. A. Robertson, M. K. Thayer, M. F. Whiting, J. F. Lawrence, A. Ślipiński, D. R. Maddison & B. D. Farrell. 2015. The beetle tree of life reveals that Coleoptera survived end-Permian mass extinction to diversify during the Cretaceous terrestrial revolution. Systematic Entomology 40 (4): 835–880.

Bostrichoidea

Orphilus subnitidus, from here.


Belongs within: Coleoptera.
Contains: Bostrichidae, Ptinidae, Dermestes, Megatominae.

The Bostrichoidea are a group of beetles predominantly adapted for dry habitats, united by modified cryptonephridism, aedeagal structure and the lack of a basal mandibular mola in the larva (Lawrence & Britton 1991).

<==Bostrichoidea
    |  i. s.: Endecatomus [Endecatomidae] B14
    |           `--E. dorsalis Mellié 1848 B14
    |--+--Bostrichidae MW15
    |  `--Ptinidae MW15
    `--Dermestidae [Dermestoidea] MW15
         |  i. s.: Reesa vespulae LB91
         |         Neoanthrenus YHT05
         |         Hexanodes AY04
         |         Thorictus A61 [Thorictidae, Thorictinae LB91]
         |           |--T. capensis A61
         |           |--T. castaneus Germ. 1834 G89
         |           |--T. germari Lucas 1846 E12
         |           |--T. mauritanicus Lucas 1846 E12
         |           `--T. puncticollis Lucas 1846 E12
         |         Trinodinae B14
         |           |--Trichelodes LB91 [Trichelodini AY04]
         |           |    `--T. delicatula LB91
         |           `--Thylodrini B14
         |                |--Thylodrias contractus Motschulsky 1839 B14
         |                `--Trichodrias B14
         |         Attagenus Latreille 1802 L02 [Attageninae LB91]
         |           |--A. macellarius [=Dermestes macellarius] L02
         |           |--A. pellio B70
         |           |--A. piceus C01
         |           |--A. posticalis Fairm. 1879 G89
         |           |--A. schaefferi C01
         |           |--A. trifasciatus [=Dermestes trifasciatus] L02
         |           |--A. unicolor OM91
         |           |    |--A. u. unicolor OM91
         |           |    `--A. u. japonicus OM91
         |           `--A. unifasciatus Fairm. 1860 G89
         |--Orphilinae B14
         |    |--Orphilus subnitidus LeConte 1861 MW15, B14
         |    `--Orphilodes B14
         `--+--Dermestes MW15
            `--Megatominae MW15

*Type species of generic name indicated

REFERENCES

[A61] Andreae, H. 1961. Coleoptera: Cossyphodidae. In: Hanström, B., P. Brinck & G. Rudebeck (eds) South African Animal Life: Results of the Lund University Expedition in 1950–1951 vol. 8 pp. 198–216. Almqvist & Wiksell: Uppsala.

[AY04] Austin, A. D., D. K. Yeates, G. Cassis, M. J. Fletcher, J. La Salle, J. F. Lawrence, P. B. McQuillan, L. A. Mound, D. J. Bickel, P. J. Gullan, D. F. Hales & G. S. Taylor. 2004. Insects ‘Down Under’—diversity, endemism and evolution of the Australian insect fauna: examples from select orders. Australian Journal of Entomology 43 (3): 216–234.

[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature's gems. Ivy Press: Lewes (United Kingdom).

[B70] Britton, E. B. 1970. Coleoptera (beetles). In: CSIRO. The Insects of Australia: A textbook for students and research workers pp. 495–621. Melbourne University Press.

[C01] Csiki, E. 1901. Bogarak [Coleopteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 75–120. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[E12] Evenhuis, N. L. 2012. Publication and dating of the Exploration Scientifique de l’Algérie: Histoire Naturelle des Animaux Articulés (1846–1849) by Pierre Hippolyte Lucas. Zootaxa 3448: 1–61.

[G89] Gestro, R. 1889. Viaggio ab Assab nel Mar Rosso dei signori G. Doria ed O. Beccari con il R. Avviso «Esploratore» dal 16 Novembre 1879 al 26 Febbraio 1880.—IV. Coleotteri. Annali del Museo Civico di Storia Naturale di Genova, Serie 2a, 7: 5–72.

[L02] Latreille, P. A. 1802. Histoire Naturelle, générale et particulière des crustacés et des insectes vol. 3. Familles naturelles des genres. F. Dufart: Paris.

[LB91] Lawrence, J. F., & E. B. Britton. 1991. Coleoptera (beetles). In: CSIRO. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 2 pp. 543–683. Melbourne University Press: Carlton (Victoria).

[MW15] McKenna, D. D., A. L. Wild, K. Kanda, C. L. Bellamy, R. G. Beutel, M. S. Caterino, C. W. Farnum, D. C. Hawks, M. A. Ivie, M. L. Jameson, R. A. B. Leschen, A. E. Marvaldi, J. V. McHugh, A. F. Newton, J. A. Robertson, M. K. Thayer, M. F. Whiting, J. F. Lawrence, A. Ślipiński, D. R. Maddison & B. D. Farrell. 2015. The beetle tree of life reveals that Coleoptera survived end-Permian mass extinction to diversify during the Cretaceous terrestrial revolution. Systematic Entomology 40 (4): 835–880.

[OM91] Okamoto, M., & K. Matsumoto. 1991. Observations on the behavior of the hypopus of Lardoglyphus konoi. In: Dusbábek, F., & V. Bukva (eds) Modern Acarology: Proceedings of the VIII International Congress of Acarology, held in České Budĕjovice, Czechoslovakia, 6–11 August 1990 vol. 2 pp. 555–559. SPB Academic Publishing: The Hague.

[YHT05] Yates, C. J., S. D. Hopper & R. H. Taplin. 2005. Native insect flower visitor diversity and feral honeybees on jarrah (Eucalyptus marginata) in Kings Park, an urban bushland remnant. Journal of the Royal Society of Western Australia 88 (4): 147–153.

Scydmaeninae

Microscydmus, copyright Edenis.


Belongs within: Staphylinidae.
Contains: Euconnini.

The Scydmaeninae, ant-like stone beetles, are a group of minute beetles found in habitats such as leaf litter or moss, or in association with ants or termites (O'Keefe 2001).

Characters (from O'Keefe 2001, as Scydmaenidae): Body elongate, slender to ovoid; slightly to very convex, 0.6–2.7 mm long; often distinctly constricted between head and pronotum and between pronotum and elytra, usually light to dark brown, sometimes black in color; often densely pubescent on head, pronotum, and elytra, pubescence usually long. Head short and broad to ovoid; deflexed; usually distinctly constricted between vertex and occiput; occiput formed into a “neck”; clypeofrontal region moderate in length, slightly narrowed anteriorly. Eyes anterior, median, or posterior; moderate in size or absent. Antennae inserted anterior to eyes, antennal insertions adjacent to widely separated; antennae composed of scape, pedicel and nine flagellomeres; antennal club composed of distal three to five antennomeres, from indistinct to distinct in form. Mandibles usually planar, subtriangular with large basal area, narrowed to acute incisor, subapical teeth usually absent. Maxillary palpus composed of four palpomeres; palpomere III large, clavate; palpomere IV variable, distinctly smaller than III; apices of lacinea and galea densely covered with long setae. Labial palpus composed of three palpomeres; mentum large. Thorax with pronotum distinctly wider than head, usually with fovea or transverse furrow along posterior margin. Scutellum small, visible, occasionally hidden. Elytra ovoid, convex to rectangular, flat; humeri varied in form with 0-2 basal foveae in broad basal impression; elytral apices usually entire, occasionally truncate or fused. Hind wings, if present, well-developed. Legs relatively short to moderate in length; procoxae projecting beyond prosternum, procoxal cavities open; mesocoxal cavities closed; trochanters relatively large and sublinear or rounded triangular; femora strongly clavate in distal half; protibiae and mesotibiae usually with dense patches of setae at distal end. Prosternum varied from subquadrate to very narrow before conical, contiguous procoxae, pubescence varied. Mesosternum usually raised, sometimes densely setose, bearing mid-ventral carina separating otherwise contiguous mesocoxae. Metasternum long, broad, occasionally carinate or medially excavate; metacoxae round to transverse, conical, widely separate to contiguous. Abdomen with six visible sternites. Male genitalia with median lobe bulbous, large, to elongate, to reduced, lightly sclerotised, distinctly curved, parameres long, slender, articulated on dorsal surface of aedeagus, occasionally absent. Ovipositor composed of paired, fused or unfused dorsolateral paraprocts, triangular dorsal proctiger, paired elongate ventral valvifers, paired elongate gonocoxae; gonostyli absent; spermatheca present, variable in shape from elongate, slender to spherical, accessory gland sometimes present.

<==Scydmaeninae [Anisosphaeridae, Scydmaenidae, Scydmaenides] B14
    |  i. s.: Adrastia MW15
    |         Phagonophana B70
    |         Heterognathus myrmecophilus B70
    |         Syndicus [Syndicini] LB91
    |         Palaeostigus Newton 1982 GN09
    |           `--P. bifoveolatus (Boheman 1851) GN09
    |         Palaeoleptochromus GN09
    |--Hapsomela [Hapsomelinae, Hapsomelitae] GN09
    |--Mastigitae [Mastiginae] GN09
    |    |--Leptomastacini GN09
    |    |--Mastigini GN09
    |    `--Clidicini GN09
    |         |--Clidicus AY04
    |         `--Papusus Casey 1897 O01
    |              `--P. macer O01
    `--Scydmaeinitae GN09
         |  i. s.: Cephenniini GN09
         |           |--Coatesia LB91
         |           |--Neseuthia LB91
         |           |--Chelonoidum Strand 1935 [=Chelonoides Croissandeau 1894] O01
         |           |    `--C. corporosum O01
         |           `--Cephennium Müller & Kunze 1822 O01
         |                |--C. anophthalmicum O01
         |                `--C. thoracicum (Müller & Kunze 1822) B14
         |         Plaumanniolini GN09
         |         Chevrolatia Jacquelin du Val 1850 O01 [Chevrolatiini GN09]
         |           |--C. amoena LeConte 1866 B14
         |           `--C. occidentalis B14
         |         Leptoscydmus Casey 1897 O01 [Leptoscydmini GN09]
         |           `--L. caseyi O01
         |--Scydmaenini LB91
         |    |--Palaeoscydmaenus LB91
         |    |--Ceramphis Casey 1897 O01
         |    |    `--C. deformata O01
         |    `--Scydmaenus Latreille 1802 GN09 (see below for synonymy)
         |         |--S. angustatus Lucas 1846 E12
         |         |--S. hellwigii [=Pselaphus hellwigii] L02
         |         |--S. motschulskii O01
         |         |--S. myrmecophilus LB91
         |         |--S. nanus Lucas 1846 E12
         |         `--S. schaumii Lucas 1846 E12
         `--+--+--Euconnini GN09
            |  `--+--Cephennodes Reitter 1884 GN09
            |     `--Eutheiini O01
            |          |--Eutheia Stephens 1830 [=Euthia Agassiz 1847; incl. Euthiopsis Müller 1925] O01
            |          |--Euthiconus Reitter 1881 [incl. Ascydmus Casey 1897, Euthiodes Brendel 1893] O01
            |          |    `--E. lata O01
            |          `--Veraphis Casey 1987 GN09
            |               `--V. cristata O01
            `--Cyrtoscydmini MW01
                 |--Neuraphus MW01
                 |--Parascydmus Casey 1897 O01
                 |--Brachycepsis Brendel 1889 O01
                 |    `--B. subpunctatus O01
                 |--Catalinus Casey 1897 O01
                 |    `--C. angustus O01
                 |--Lophioderus Casey 1897 O01
                 |    `--L. arcifer O01
                 |--Taphroscydmus Casey 1897 O01
                 |    `--T. californicus O01
                 |--Stenichnus Thomson 1859 GN09 [incl. Cyrtoscydmus Motschulsky 1870 O01]
                 |    |--S. perforatus O01
                 |    `--S. turbatus Casey 1897 GN09
                 `--Microscydmus Saulcy & Croissandeau 1893 O01
                      |--M. (Microscydmus) [incl. Opresus Casey 1897] O01
                      |    `--M. misellus O01
                      |--M. (Delius Casey 1897) O01
                      |    `--M. (D.) robustulus O01
                      `--M. (Neladius Casey 1897) O01
                           `--M. (N.) tenuis O01

Scydmaenus Latreille 1802 GN09 [incl. Acholerops Casey 1897 O01, Eumicrus LaPorte 1840 O01, Microstemma Motschulsky 1858 O01]

*Type species of generic name indicated

REFERENCES

[AY04] Austin, A. D., D. K. Yeates, G. Cassis, M. J. Fletcher, J. La Salle, J. F. Lawrence, P. B. McQuillan, L. A. Mound, D. J. Bickel, P. J. Gullan, D. F. Hales & G. S. Taylor. 2004. Insects ‘Down Under’—diversity, endemism and evolution of the Australian insect fauna: examples from select orders. Australian Journal of Entomology 43 (3): 216–234.

[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature's gems. Ivy Press: Lewes (United Kingdom).

[B70] Britton, E. B. 1970. Coleoptera (beetles). In: CSIRO. The Insects of Australia: A textbook for students and research workers pp. 495–621. Melbourne University Press.

[E12] Evenhuis, N. L. 2012. Publication and dating of the Exploration Scientifique de l’Algérie: Histoire Naturelle des Animaux Articulés (1846–1849) by Pierre Hippolyte Lucas. Zootaxa 3448: 1–61.

[GN09] Grebennikov, V. V., & A. F. Newton. 2009. Good-bye Scydmaenidae, or why the ant-like stone beetles should become megadiverse Staphylinidae sensu latissimo (Coleoptera). European Journal of Entomology 106: 275–301.

[L02] Latreille, P. A. 1802. Histoire Naturelle, générale et particulière des crustacés et des insectes vol. 3. Familles naturelles des genres. F. Dufart: Paris.

[LB91] Lawrence, J. F., & E. B. Britton. 1991. Coleoptera (beetles). In: CSIRO. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 2 pp. 543–683. Melbourne University Press: Carlton (Victoria).

[MW15] McKenna, D. D., A. L. Wild, K. Kanda, C. L. Bellamy, R. G. Beutel, M. S. Caterino, C. W. Farnum, D. C. Hawks, M. A. Ivie, M. L. Jameson, R. A. B. Leschen, A. E. Marvaldi, J. V. McHugh, A. F. Newton, J. A. Robertson, M. K. Thayer, M. F. Whiting, J. F. Lawrence, A. Ślipiński, D. R. Maddison & B. D. Farrell. 2015. The beetle tree of life reveals that Coleoptera survived end-Permian mass extinction to diversify during the Cretaceous terrestrial revolution. Systematic Entomology 40 (4): 835–880.

[MW01] Molleman, F., & D. E. Walter. 2001. Niche segregation and can-openers: scydmaenid beetles as predators of armoured mites in Australia. In: Halliday, R. B., D. E. Walter, H. C. Proctor, R. A. Norton & M. J. Colloff (eds) Acarology: Proceedings of the 10th International Congress pp. 283–288 Publishing: Melbourne.

[O01] O’Keefe, S. T. 2001. Scydmaenidae Leach, 1915. In: Arnett, R. H., Jr & M. C. Thomas (eds) American Beetles vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia pp. 259–267. CRC Press: Boca Raton.

Lomechusini

Drusilla canaliculata, copyright Udo Schmidt.


Belongs within: Aleocharinae.
Contains: Zyras.

The Lomechusini are a diverse but poorly defined group of staphylinid beetles that have been recognised in the past (under the name Myrmedoniini) as including almost all Aleocharinae with 4,5,5-segmented tarsi. Many species possess mouthparts with extensive maxillary sinuses that extend caudally beyond the cardo, and an elongated galea and lacinia (Seevers 1957). Many, but not all, species are associated with ants and members of the subtribe Lomechusina possess prominent tufts of golden-yellow trichomes on the abdomen (Newton et al. 2001).

Characters (from Newton et al. 2001): Tarsi 4-5-5; metasternal process distinctly longer than very short mesosternal process; galea and lacinia of maxilla usually moderately to greatly elongated, length of galea equal to or greater than distance to base of galea from cardo; mesocoxae moderately to very broadly separated by broad meso- and metasternal processes.

Lomechusini [Myrmedoniini, Zyrasini] NT01
    |--Schistogeniae S57
    |--Termitonda Seevers 1957 [Termitondina] S57
    |    |--*T. liberiae Seevers 1957 S57
    |    `--T. tachyporoides Seevers 1957 S57
    |--Lomechusina NT01
    |    |--Lomechusa strumosa A71
    |    `--Xenodusa Wasmann 1894 [incl. Pseudolomechusa Mann 1914] NT01
    |         |--X. cava B14
    |         `--X. reflexa (Walker 1866) B14
    |--Feldina S57
    |    |--Felda Blackwelder 1952 [=Asticta Wasmann 1916 (preoc.)] S57
    |    |    `--*F. butteli (Wasmann 1916) [=*Asticta butteli] S57
    |    |--Zunia Blackwelder 1952 [=Disticta Wasmann 1916 (preoc.)] S57
    |    |    `--*Z. capritermitis (Wasmann 1916) [=*Disticta capritermitis] S57
    |    `--Termitobiella Wasmann 1916 S57
    |         `--*T. setipes Wasmann 1916 S57
    |--Termitozyrina S57
    |    |--Termitophagus Silvestri 1945 S57
    |    |    `--*T. synterminus Silvestri 1945 S57
    |    |--Termitonusa Borgmeier 1950 S57
    |    |    `--*T. sequax Borgmeier 1950 S57
    |    |--Iheringocantharus Bernhauer 1912 S57
    |    |    `--*I. ypiranganus Bernhauer 1912 S57
    |    |--Termitosymbia Seevers 1957 S57
    |    |    `--*T. nitida Seevers 1957 S57
    |    `--Termitozyras Seevers 1957 S57
    |         |--*T. adamsoni Seevers 1957 S57
    |         |--T. boliviae Seevers 1957 S57
    |         `--T. guianae Seevers 1957 S57
    `--Myrmedoniina [Myrmedoniae, Zyrasina] NT01
         |--Atemeles S57
         |--Zyras NT01
         |--Termozyras Cameron 1930 S57
         |    `--*T. politus Cameron 1930 S57
         |--Urodonia Silvestri 1946 S57
         |    `--*U. notabilis Silvestri 1946 S57
         |--Apalonia Casey 1906 NT01
         |    `--A. seticornis NT01
         |--Meronera Sharp 1887 [incl. Merona Sharp 1883 non Norman 1865] NT01
         |    `--M. venustula NT01
         |--Tetradonia Wasmann 1894 [incl. Chlorotusa Casey 1906] NT01
         |    `--T. megalops (Casey 1906) NT01
         |--Trachyota Casey 1906 NT01
         |    `--T. lativentris NT01
         `--Drusilla Leach 1819 [incl. Astilbus Dillwyn 1829, Myrmedonia Erichson 1837] NT01
              |--*D. canaliculata (Fabricius 1787) S57, NT01 [=Astilbus canaliculatus C01]
              |--D. cavicollis Casey 1906 NT01
              |--‘Myrmedonia’ clavigera Fauvel 1878 O86
              |--‘Myrmedonia’ humeralis C01
              |--D. impressicollis [incl. D. sculpticollis] S57
              |--D. incola S57
              |--‘Myrmedonia’ insignicornis Fauvel 1878 O86
              |--D. planaticollis S57
              `--‘Myrmedonia’ tristis Lucas 1846 E12

Lomechusini incertae sedis:
  Dinocoryna Casey 1893 [incl. Ecitonusa Wasmann 1897] NT01
    `--D. carolinensis NT01
  Ecitocala Frank 1981 NT01
    `--E. rugosa Frank 1981 NT01
  Ecitonidia Wasmann 1900 NT01
    `--E. wheeleri Wasmann 1900 NT01
  Ecitoxenidia Wasmann 1909 NT01
    `--E. brevicornis NT01
  Microdonia Casey 1893 NT01
    `--M. nitidiventris NT01
  Xesturida Casey 1906 NT01
    `--X. laevis Casey 1906 NT01

*Type species of generic name indicated

REFERENCES

[A71] Askew, R. R. 1971. Parasitic Insects. Heinemann Educational Books: London.

[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature's gems. Ivy Press: Lewes (United Kingdom).

[C01] Csiki, E. 1901. Bogarak [Coleopteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 75–120. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[E12] Evenhuis, N. L. 2012. Publication and dating of the Exploration Scientifique de l’Algérie: Histoire Naturelle des Animaux Articulés (1846–1849) by Pierre Hippolyte Lucas. Zootaxa 3448: 1–61.

[NT01] Newton, A. F., M. K. Thayer, J. S. Ashe & D. S. Chandler. 2001. Staphylinidae Latreille, 1802. In: Arnett, R. H., Jr & M. C. Thomas (eds) American Beetles vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia pp. 272–418. CRC Press: Boca Raton.

[O86] Olliff, A. S. 1886. A revision of the Staphylinidae of Australia. Part I. Proceedings of the Linnean Society of New South Wales, series 2, 1 (2): 403–473, pl. 7.

[S57] Seevers, C. H. 1957. A monograph on the termitophilous Staphylinidae (Coleoptera). Fieldiana Zoology 40: 1–334.

Ptomaphagini

Habitus of Adelopsis tandapi, from Salgado (2005). Scale bar = 1 mm.


Belongs within: Leiodidae.

The Ptomaphagini are a group of soil-dwelling scavenging beetles, some of which are found in subterranean habitats such as animal burrows or caves (Peck 2001).

Characters (from Peck 2001): Head with occipital carina, resting against front of pronotum when head is in repose; cervical sclerites present; elytra with glossy surface, setal bases arranged in strigae; procoxal cavities internally closed; posterior coxae contiguous; tarsi five-segmented; hind margin of hind tibia with two inner long spines and outer row or comb of short and equal spines only.

Ptomaphagini S05
    |--Adelopsis Portevin 1907 P90
    |    |--A. bioforestae Salgado 2002 S05
    |    |--A. brasiliensis Jeannel 1936 S05
    |    |--A. coronaria Gnaspini & Peck 1996 S05
    |    `--A. tandapi Salgado 2005 S05
    `--Ptomaphagus Illiger 1798 (see below for synonymy) P01
         |  i. s.: P. consobrinus P90
         |         P. hirtus (Tellkampf 1844) B14
         |         P. shapardi P90
         `--P. (Adelops Tellkampf 1884) S05, P01
              |--P. (A.) bordoni (Jeannel 1964) [=Adelopsis bordoni] S05
              `--P. (A.) sciakyi (Zoia 1992) [=Adelopsis sciakyi] S05

Ptomaphagus Illiger 1798 [incl. Appaladelopsis Gnaspini 1996, Echinocoleus Horn 1885, Merodiscus Jeannel 1934, Tupania Szymczakowski 1961] P01

*Type species of generic name indicated

REFERENCES

[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature's gems. Ivy Press: Lewes (United Kingdom).

[P90] Peck, S. B. 1990. Insecta: Coleoptera Silphidae and the associated families Agyrtidae and Leiodidae. In: Dindal, D. L. (ed.) Soil Biology Guide pp. 1113–1136. John Wiley & Sones: New York.

[P01] Peck, S. B. 2001. Leiodidae Fleming, 1821. In: Arnett, R. H., Jr & M. C. Thomas (eds) American Beetles vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia pp. 250–258. CRC Press: Boca Raton.

[S05] Salgado, J. M. 2005. Cholevinae (Coleoptera, Leiodidae) from Ecuador: new data and two new species. Graellsia 61 (1): 51–60.

Hydraenidae

Ochthebius sp., copyright Guy A. Hanley.


Belongs within: Staphylinoidea.

The Hydraenidae are a group of small beetles characterised by maxillary palps as long as or longer than their antennae, that feed on algae in various aquatic or damp habitats (Lawrence & Britton 1991).

Characters (from Lawrence & Britton 1991): Small with relatively inconspicuous antennae, often concealed within grooves beneath head and cavities between prostemum and pronotal hypomera, and well-developed maxillary palps which may be as long as or longer than antennae; antennae eight- or nine-segmented with five- or six-segmented club, segment preceding club usually forming cupule; fore coxae transverse or globular, trochantins exposed, fore coxal cavities externally and internally open or closed; mid coxal cavities narrowly to moderately separated, laterally open; tarsi pentamerous or tetramerous; abdomen with six or seven ventrites, none connate. Larvae usually elongate, campodeiform, with well-developed legs, articulated urogomphi, and pair of hooks on segment 10.

Hydraenidae [Limnebiidae]
    |--Hydraeninae LB91
    |    |--Limnebiini P01
    |    |    |--Laeliaena Sahlberg 1900 P01
    |    |    `--Limnebius Leach 1915 P01
    |    |         |--L. discolor P01
    |    |         `--L. piceus P01
    |    `--Hydraenini P01
    |         |--Adelphydraena P01
    |         `--Hydraena Kugelann 1794 P01
    |              |  i. s.: H. anisonycha Perkins 1980 B14
    |              |         H. luridipennis LB91
    |              |         H. riparia L02
    |              `--H. (Hydraena) F05
    |                   |--H. (H.) gavarrensis Jäch, Díaz & Martinoy 2005 F05
    |                   `--H. (H.) marinae Castro 2004 F05
    `--Ochthebiinae LB91
         |  i. s.: Tympanogaster LB91
         |         Hughleechia giulianii LB91
         |         Meropathus zealandicus Ordish 1984 LB91, E02
         `--Ochthebiini P01
              |--Neochthebius Orchymont 1932 P01
              |    `--N. vandykei P01
              |--Enicocerus Stephens 1829 P01
              |    `--E. benefossus P01
              |--Gymnochthebius Orchymont 1943 P01
              |    |--G. laevipennis P01
              |    |--G. maureenae P01
              |    `--G. nitidus P01
              `--Ochthebius Leach 1815 P01
                   |--O. arenicolus P01
                   |--O. aztecus Sharp 1887 B14
                   |--O. bisinuatus P01
                   |--O. californicus P01
                   |--O. discretus P01
                   |--O. gruwelli P01
                   |--O. lejolisi PP64
                   |--O. lineatus P01
                   |--O. marinus [=Elophorus marinus] G89
                   |--O. puncticollis P01
                   `--O. subinteger PP64

*Type species of generic name indicated

REFERENCES

[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature's gems. Ivy Press: Lewes (United Kingdom).

[E02] Emberson, R. M. 2002. The beetle (Coleoptera) fauna of the Chatham Islands: additions and corrections. New Zealand Entomologist 25: 69–77.

[F05] Fernández, J. 2005. Noticia de nuevos táxones para la ciencia en el ámbito Íbero-Balear y Macaronésico. Nuevos táxones animales descritos en la península Ibérica y Macaronesia desde 1994 (IX). Graellsia 61 (2): 261–282.

[G89] Gestro, R. 1889. Viaggio ab Assab nel Mar Rosso dei signori G. Doria ed O. Beccari con il R. Avviso «Esploratore» dal 16 Novembre 1879 al 26 Febbraio 1880.—IV. Coleotteri. Annali del Museo Civico di Storia Naturale di Genova, Serie 2a, 7: 5–72.

[L02] Latreille, P. A. 1802. Histoire Naturelle, générale et particulière des crustacés et des insectes vol. 3. Familles naturelles des genres. F. Dufart: Paris.

[LB91] Lawrence, J. F., & E. B. Britton. 1991. Coleoptera (beetles). In: CSIRO. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 2 pp. 543–683. Melbourne University Press: Carlton (Victoria).

[PP64] Peres, J. M., & J. Picard. 1964. Nouveau manuel de bionomie benthique de la mer Mediterranee. Recueil des Travaux de la Station Marine d'Endoume, Bulletin 31 (27): 5–137.

[P01] Perkins, P. D. 2001. Hydraenidae Mulsant, 1844. In: Arnett, R. H., Jr & M. C. Thomas (eds) American Beetles vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia pp. 228–232. CRC Press: Boca Raton.

Julodinae

Brush jewel beetle Julodis viridipes on Othonna arbuscula, copyright Alexey Yakovlev.


Belongs within: Buprestidae.

The Julodinae are a group of jewel beetles whose larvae are free-living soil dwellers, feeding externally on roots (Bouchard 2014).

Julodinae [Amblysternini, Julodides, Julodini] B14
    |--Aaata Semenov-Tian-Shanskij 1906 B85
    |    `--A. finchi (Waterhouse 1884) B14
    |--Julodella Semenov-Tian-Shanskij 1893 [=Julodiella (l. c.)] B85
    |--Neojulodis Kerremans 1902 B85
    |    |  i. s.: N. rufolimbata G08
    |    `--N. (Protojulodis Holm 1979) B85
    |--Amblysterna Thomson 1878 B85
    |    |--A. johnstoni B14
    |    `--A. natalensis (Fåhraeus 1851) B14
    |--Sternocera Eschscholtz 1829 [incl. Charytonia Gistel 1834, Sternoxus] B85
    |    |--S. boucardi G08
    |    |--S. castanea B14
    |    |--S. chrysis (Fabricius 1775) B14
    |    |    |--S. c. chrysis B14
    |    |    `--S. c. nitidicollis B14
    |    |--S. discedens B14
    |    |--S. hildebrandti G08
    |    `--S. hunteri G08
    `--Julodis Eschscholtz 1829 [incl. Phyllis Gistel 1834, Saccosoma Motschulsky 1859] B85
         |--J. arabica G89
         |--J. cirrosa B14
         |    |--J. c. cirrosa B14
         |    |--J. c. hirtiventris Laporte 1835 B14
         |    `--J. c. mellyi B14
         |--J. fascicularis B14
         |--J. fimbriata [=Buprestis fimbriata] G89
         |--J. hirsuta B14
         |--J. sulcicollis B14
         `--J. viridipes B14

*Type species of generic name indicated

REFERENCES

[B85] Bellamy, C. L. 1985. A catalogue of the higher taxa of the family Buprestidae (Coleoptera). Navorsinge van die Nasionale Museum Bloemfontein 4 (15): 405–472.

[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature's gems. Ivy Press: Lewes (United Kingdom).

[G89] Gestro, R. 1889. Viaggio ab Assab nel Mar Rosso dei signori G. Doria ed O. Beccari con il R. Avviso «Esploratore» dal 16 Novembre 1879 al 26 Febbraio 1880.—IV. Coleotteri. Annali del Museo Civico di Storia Naturale di Genova, Serie 2a, 7: 5–72.

[G08] Golding, M. R. 2008. A Pictorial Field Guide to the Beetles of Australia. Part 3: Chalcophorinae. Ocean Publishing.

Dascillidae

Dascillus cervinus, copyright Udo Schmidt.


Belongs within: Elateriformia.

The Dascillidae are a group of elongate beetles with serrate antennae whose larvae live on organic matter in the soil (Lawrence & Britton 1991).

Characters (from Lawrence & Britton 1991): Elongate, more or less parallel-sided, brown and clothed with decumbent grey or yellow pubescence usually forming narrow stripes on elytra; antennae eleven-segmented, serrate; pronotum strongly transverse with complete lateral carinae visible from above; fore coxae transverse, trochantins exposed, fore coxal cavities externally and internally open; mid coxal cavities narrowly separated, laterally open; tarsi pentamerous; abdomen with five ventrites, 0–2 connate. Larva more or less C-shaped, with large head, well-developed legs, and complex epipharyngeal and hypopharyngeal armature; labrum partly fused to clypeus; antennae appearing two-segmented with sensorium at apex; T9 with pair of minute urogomphi.

<==Dascillidae B14
    |  i. s.: Notodascillus sublineatus LB91
    |         Helodes M86
    |           |--H. atkinsoni Waterh. 1877 M86
    |           |--H. australis Erichs. 1841 M86
    |           |--H. distincta Lucas 1846 E12
    |           |--H. maculatus Waterh. 1877 M86
    |           `--H. vicina Lucas 1846 E12
    |         Notiocyphon B70
    |--Dascillus Latreille 1802 L02 [Dascillinae B14]
    |    |--D. brevicornis Macleay 1872 M86
    |    |--D. cervinus [=Atopa cervina] L02
    |    |--D. davidsoni LeConte 1859 B14
    |    `--D. plumbeus B14
    `--Karumiinae B14
         |--Anorus piceus LeConte 1859 B14
         `--Karumia B14
              |--K. estafilinoides B14
              `--K. staphylinus (Semenov & Martynov 1925) B14

*Type species of generic name indicated

REFERENCES

[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature's gems. Ivy Press: Lewes (United Kingdom).

[B70] Britton, E. B. 1970. Coleoptera (beetles). In: CSIRO. The Insects of Australia: A textbook for students and research workers pp. 495–621. Melbourne University Press.

[L02] Latreille, P. A. 1802. Histoire Naturelle, générale et particulière des crustacés et des insectes vol. 3. Familles naturelles des genres. F. Dufart: Paris.

[LB91] Lawrence, J. F., & E. B. Britton. 1991. Coleoptera (beetles). In: CSIRO. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 2 pp. 543–683. Melbourne University Press: Carlton (Victoria).

[M86] Masters, G. 1886. Catalogue of the described Coleoptera of Australia. Part IV. Proceedings of the Linnean Society of New South Wales, series 2, 1 (2): 259–380.

Chrysochroinae

Capnodis cariosa, copyright Siga.


Belongs within: Buprestidae.
Contains: Chalcophorina.

The Chrysochroinae are a group of jewel beetles whose antennae have pores distributed over both surfaces of the segments rather than being concentrated into a ventral depression.

Chrysochroinae [Chalcophorides, Chalcophorinae] B14
    |--Vadonaxia Descarpentries 1969 [Vadonaxiini] B85
    |--Mendizabalia Cobos 1957 [Mendizabalini] B85
    |--Cinyrini [Paraleptodemini] B85
    |    |--Euleptodema Obenberger 1928 B85
    |    `--Cinyra Laporte & Gory 1837 [=Cynira (l. c.); incl. Paraleptodema Obenberger 1936] B85
    |         `--C. spilota (Hope 1846) [=Cisseis spilota] M86
    |--Epistomentini B85
    |    |--Cyrioxus Hoscheck 1925 B85
    |    |--Araucariana Levey 1978 B85
    |    |--Epistomentis Solier 1850 B85
    |    |    `--E. pictus G08
    |    |--Cyria Solier 1833 [incl. Cyrioides Carter 1920, Xenocyria Obenberger 1947] B85
    |    |    |--C. australis C08 [=Buprestis australis B35; incl. C. gagates C08]
    |    |    |--C. cincta Carter 1908 C08
    |    |    |--C. elateroides Saund. 1872 M86
    |    |    `--C. vittigera C08
    |    `--Diadoxus Saunders 1868 B85, G08
    |         |--D. erythrurus (White 1846) G08 [=Stigmodera erythrura M86; incl. Anthaxia pistacina M86]
    |         |--D. jungi Blackburn 1899 G08
    |         |--D. regius Peterson 1991 G08
    |         `--D. scalaris [incl. Anthaxia erichsoni] M86
    |--Psilopterini [Psilopterides] B85
    |    |--Touzalinia Thery 1922 B85
    |    |--Icarina Alluaud 1896 [incl. Icaria Thomson 1878 non Saussure 1853] B85
    |    |--Asidoptera Obenberger 1923 B85
    |    |--Aurigena Laporte & Gory 1837 B85
    |    |--Ectinogonia Spinola 1837 [incl. Kheilia Obenberger 1926 (preoc.), Kheiliella Obenberger 1940] B85
    |    |    `--E. (Achardella Obenberger 1926) B85
    |    |--Lampetis Spinola 1837 [incl. Pseudolampetis Obenberger 1926, Spinthoptera Casey 1909] B85
    |    |    `--L. gregaria G08
    |    |--Cyphosoma Mannerheim 1837 (see below for synonymy) B85
    |    |    `--C. tartaricum G08
    |    |--Perotis Spinola 1837 B85
    |    |    `--P. unicolor G08
    |    |--Latipalpis Solier 1833 B85
    |    |    `--L. (Palpilatis Bily 1980) B85
    |    |--Polybothris Spinola 1837 (see below for synonymy) B85
    |    |    |--P. angulosa B14
    |    |    |--P. auriventris (Laporte & Gory 1837) B14
    |    |    `--P. sumptuosa G08
    |    |--Psiloptera Solier 1833 B85
    |    |    |  i. s.: P. acroptera B14
    |    |    |         P. attenuata (Fabricius 1793) B14
    |    |    |         P. transversovittata B14
    |    |    |         P. weigelti S12
    |    |    `--P. (Damarsila Thomson 1881) B85
    |    |--Oedisterna Lacordaire 1857 [incl. Lachesis Saunders 1871 nec Daudin 1803 nec Risso 1826] B85
    |    |    |--O. (Embrikillium Obenberger 1936) B85
    |    |    |--O. (Fahraeusia Obenberger 1936) B85
    |    |    |--O. (Monosacra Thomson 1878) B85
    |    |    |--O. (Pseudoperotis Obenberger 1936) B85
    |    |    `--O. (Strandissa Obenberger 1936) B85
    |    `--Capnodis Eschscholtz 1829 B85
    |         |--C. carbonaria B14
    |         |--C. cariosa B14
    |         |--C. excisa G08
    |         |--C. miliaris (Klug 1829) B14
    |         |    |--C. m. miliaris B14
    |         |    `--C. m. metallica B14
    |         `--C. tenebrionis B14
    `--Chrysochroini [Catoxanthini] G08
         |  i. s.: Chrysochroa Solier 1833 G08, B85
         |           |--C. buqueti (Gory 1833) B14
         |           |    |--C. b. buqueti B14
         |           |    |--C. b. kerremansi B14
         |           |    |--C. b. rugicollis B14
         |           |    |--C. b. suturalis B14
         |           |    `--C. b. trimaculata B14
         |           |--C. castelnaudi B14
         |           |--C. fulminans G08
         |           `--C. toulgoeti G08
         |         Megaloxantha Kerremans 1902 B85
         |           |--M. bicolor (Fabricius 1775) B14
         |           |    |--M. b. bicolor B14
         |           |    |--M. b. ohtanii B14
         |           |    `--M. b. ryoi B14
         |           |--M. concolor B14
         |           |--M. gigantea B14
         |           `--M. netscheri B14
         |         Demochroa White 1859 B85
         |           `--D. lacordairi G08
         |         Catoxantha Solier 1833 B14, B85 [incl. Euchloris B85, Pyranthe Gistel 1834 B85]
         |           |--C. bonvouloirii B14
         |           |--C. eburnea B14
         |           |--C. nagaii B14
         |           |--C. opulenta (Gory 1832) B14
         |           |    |--C. o. opulenta B14
         |           |    `--C. o. borneensis B14
         |           |--C. pierrei B14
         |           `--C. purpurea B14
         |         Semenoviella Obenberger 1924 B85
         |         Steraspis Solier 1833 [incl. Phalantha Gistel 1834] B85
         |           |--S. ambigua G08
         |           `--S. squamosa [=Buprestis squamosa] G89
         |         Chrysaspina Thery 1930 [incl. Chrysaspis Kerremans 1893 (preoc.)] B85
         |         Pygichaeta Obenberger 1920 B85
         |         Asamia Thery 1909 B85
         |         Agelia Laporte & Gory 1835 B85
         |           |--A. burmensis B14
         |           |--A. chalybea B14
         |           |--A. fasciata B14
         |           |--A. limbata B14
         |           |--A. lordi B14
         |           |--A. obtusicollis B14
         |           |--A. pectinicornis B14
         |           |--A. petelii (Gory 1840) B14
         |           `--A. theryi B14
         |         Philocteanus Deyrolle 1864 B85
         |         Kolleria Thery 1925 B85
         |         Epidelus Deyrolle 1864 B85
         |         Epidelomorphus Thery 1925 B85
         |         Micropistus Thery 1922 B85
         |           `--M. microcephalus G08
         |         Szentendreya Holynski 1980 B85
         |         Chrysopistus Thery 1922 B85
         |         Callopistus Deyrolle 1864 B85
         |         Pseudocallopistus Obenberger 1942 B85
         |         Cyalithus Thomson 1878 [incl. Aprosopus Deyrolle 1864] B85
         |         Cyalithoides Fisher 1922 B85
         |         Mrazia Obenberger 1928 B85
         `--Chalcophorina B14

Chrysochroinae incertae sedis:
  Cyriodes Carter 1920 G08
    |--C. australis (Boisduval 1835) G08
    |--C. cincta (Carter 1908) G08
    |--C. elateroides (Saunders 1872) G08
    |--C. imperialis (Fabricius 1801) G08
    |--C. sexspilota Carter 1920 G09
    |--C. tridens (Blackburn 1892) G08
    `--C. vittigera (L. & G. 1835) G08
  Calopistus castelnaui G08
  Afrochroa lepida G08
  Chrycestes tripunctata G08

Cyphosoma Mannerheim 1837 [incl. Coeculus Laporte & Gory 1839, Cyphonota Dejean 1836, Phelix Marseul 1865] B85

Polybothris Spinola 1837 [incl. Alampetis Thomson 1878, Amphisbeta Thomson 1878, Apatea Thomson 1878, Aplax Thomson 1878, Carcinias Thomson 1878, Cassidabothris Thomson 1878, Coccinellopsis Thomson 1878, Cornelia Thomson 1878, Enharpya Thomson 1878, Erebodes Thomson 1878, Hemisobothris Thomson 1878, Laconides Thomson 1878, Paleobothris Thomson 1878, Phobetodes Thomson 1878, Pseudophthalma Thomson 1878, Pycnobothris Thomson 1878] B85

*Type species of generic name indicated

REFERENCES

[B85] Bellamy, C. L. 1985. A catalogue of the higher taxa of the family Buprestidae (Coleoptera). Navorsinge van die Nasionale Museum Bloemfontein 4 (15): 405–472.

[B35] Boisduval, J. B. 1835. Voyage de Découvertes de l’Astrolabe. Exécuté par ordre du Roi, pendant les années 1826–1827–1828–1829, sous le commandement de M. J. Dumont d'Urville. Faune entomologique de l'océan Pacifique, avec l'illustration des insectes nouveaux recueillis pendant le voyage vol. 2. Coléoptères et autres ordres. J. Tastu: Paris.

[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature's gems. Ivy Press: Lewes (United Kingdom).

[C08] Carter, H. J. 1908. Revision of the genus Seirotrana, together with descriptions of new species of other Australian Coleoptera. Proceedings of the Linnean Society of New South Wales 33: 392–422.

[G89] Gestro, R. 1889. Viaggio ab Assab nel Mar Rosso dei signori G. Doria ed O. Beccari con il R. Avviso «Esploratore» dal 16 Novembre 1879 al 26 Febbraio 1880.—IV. Coleotteri. Annali del Museo Civico di Storia Naturale di Genova, Serie 2a, 7: 5–72.

[G08] Golding, M. R. 2008. A Pictorial Field Guide to the Beetles of Australia. Part 3: Chalcophorinae. Ocean Publishing.

[G09] Golding, M. R. 2009. A Pictorial Field Guide to the Beetles of Australia. Part 4: Bolboceratinae. Privately published.

[M86] Masters, G. 1886. Catalogue of the described Coleoptera of Australia. Part III. Proceedings of the Linnean Society of New South Wales, series 2, 1 (1): 21–126.

[S12] Schaal, S. 2012. Messel Pit fossil site—the legacy of the environment and life of the Eocene. In: Talent, J. A. (ed.) Earth and Life: Global biodiversity, extinction intervals and biogeographic perturbations through time pp. 225–236. Springer.

Polycestinae

Astraeus fraterculus, copyright Western Australian Museum.


Belongs within: Buprestidae.
Contains: Acmaeodera, Polycesta.

The Polycestinae are a group of jewel beetles that often have strongly punctured and/or carinate elytra, and a sternal cavity for reception of the prosternal process often formed entirely from the mesosternum.

Polycestinae [Acmaeoderinae, Acmaeoderites, Polycestides, Ptosimites] B14
    |--Pseudacherusia Kerremans 1904 B85
    |--Perucola Thery 1925 [Perucolini] B85
    |--Prospheres Thomson 1878 [=Phospheres (l. c.); Prospherini] B85
    |--Nothomorpha Thomson 1878 [Nothomorphini] B85
    |--Acmaeoderoides Van Dyke 1942 [Acmaeoderoidini] B85
    |--Xyroscelis Thomson 1878 [Xyroscelini] B85
    |    `--X. crocata [incl. X. melanosticta Hope 1846, X. nodosa Hope 1846] M86
    |--Thrincopyge LeConte 1858 B14, B85 [Thrincopyginae, Thrincopygini]
    |    `--T. alacris LeConte 1858 B14
    |--Tylauchenini B85
    |    |--Tylauchenia Burmeister 1872 [incl. Ocypetes Thomson 1878 non Lesson 1842] B85
    |    |--Neocypetes Cobos 1973 B85
    |    `--Ancylotela Waterhouse 1882 B85
    |--Polyctesini B85
    |    |--Paraxenopsis Cobos 1980 B85
    |    |--Xenopsis Saunders 1867 B85
    |    |--Polyctesis Marseul 1865 [incl. Schoutedenia Obenberger 1924 (preoc.), Schoutendeiastes Burgeon 1941] B85
    |    |--Chrysophana LeConte 1860 B85
    |    `--Beerellus Nelson 1981 B85
    |--Acherusini B85
    |    |--Hayekina Cobos 1980 B85
    |    |--Bulis Laporte & Gory 1837 B85
    |    `--Acherusia Laporte & Gory 1837 B85
    |         |  i. s.: A. childreni M86
    |         `--A. (Nelsonila Cobos 1980) B85
    |--Ptosimini B85
    |    |--Paratrachys Saunders 1873 [Paratrachysae] B85
    |    `--Ptosimae B85
    |         |--Ptosima Solier 1833 B85
    |         `--Sponsor Laporte & Gory 1839 B85
    |              |--S. (Neoptosima Thery 1905) B85
    |              `--S. (Stenianthe Fairmaire 1886) B85
    |--Tyndarini B85
    |    |--Paraancylotela Cobos 1959 B85
    |    |--Bordonia Cobos 1980 B85
    |    `--Tyndaris Thomson 1857 B85
    |         |--T. (Barberia Cobos 1980) B85
    |         |--T. (Knulliella Cobos 1980) B85
    |         |--T. (Paratyndaris Fisher 1919) B85
    |         `--T. (Tucsonia Cobos 1980) B85
    |--Astraeus Laporte & Gory 1837 [Astraeusini] B85
    |    |--A. aberrans B14
    |    |--A. adamsi B14
    |    |--A. caledonicus B14
    |    |--A. carnabyi B14
    |    |--A. dedariensis B14
    |    |--A. flavopictus M86
    |    |--A. fraterculus van de Poll 1889 B14
    |    |--A. mastersi Macleay 1872 M86
    |    |--A. navarchis Thoms. 1856 M86
    |    `--A. samoellei Saund. 1868 M86
    |--Acmaeoderini B85
    |    |--Acmaeodera B14
    |    |--Microacmaeodera Cobos 1966 B85
    |    |--Acmaeoderopsis Barr 1974 B85
    |    |--Anambodera Barr 1974 B85
    |    |--Xantheremia Volkovitsh 1979 B85
    |    |--Cochinchinula Volkovitsh 1984 B85
    |    |    `--*C. quadriareolata [=Acmaeodera quadriareolata] B85
    |    `--Acmaeoderella Cobos 1955 B85
    |         |--A. (Carininota Volkovitsh 1979) B85
    |         |--A. (Euacmaeoderella Volkovitsh 1979) B85
    |         |--A. (Kocheridia Cobos 1958) B85
    |         |--A. (Liogastria Volkovitsh 1979) B85
    |         `--A. (Omphalothorax Volkovitsh 1979) B85
    `--Polycestini B85
         |--Polycesta B85
         |--Pseudocastalia Kraatz 1896 B85
         |--Agenjosiana Cobos 1981 B85
         |--Madecacesta Descarpentries 1975 B85
         |--Cobosesta Holm 1982 B85
         |--Jelinekia Cobos 1981 B85
         |--Kogania Cobos 1981 B85
         |--Madecastalia Descarpentries 1975 B85
         |--Parapolycesta Cobos 1981 B85
         |--Strigopteroides Cobos 1981 B85
         |--Polycestoides Kerremans 1902 B85
         |--Polycestaxis Obenberger 1920 B85
         |--Polycestaxia Cobos 1981 B85
         |--Pseudopolycesta Cobos 1981 B85
         |--Castaliella Cobos 1981 B85
         |--Micropolycesta Cobos 1981 B85
         |--Aldabrica Cobos 1981 B85
         |--Polycestella Kerremans 1902 B85
         |--Thurntaxisia Schatzmayr 1929 [incl. Acassidiocesta Mandl 1972] B85
         |--Paracastalia Kerremans 1902 [incl. Lypocesta Cobos 1981] B85
         |    `--P. (Mattheesta Holm 1982) B85
         |--Polycestina Cobos 1981 B85
         |    `--P. (Damarana Cobos 1981) B85
         |--Neopolycesta Kerremans 1904 [incl. Holmia Cobos 1981] B85
         |    `--N. (Peringueyesta Holm 1982) B85
         `--Strigoptera Dejean 1838 [incl. Castalia Gory & Laporte 1837 non Savigny 1820] B85
              |--S. bimaculata (Linnaeus 1758) B35, L58 (see below for synonymy)
              |--S. bipustulata B35 [=Buprestis bipustulata B35, Castalia bipustulata M86]
              |--‘Castalia’ globithorax Thoms. 1878 M86
              `--‘Castalia’ pacifica M86

Strigoptera bimaculata (Linnaeus 1758) B35, L58 [=Buprestis bimaculata B35, Castalia bimaculata M86; incl. C. impustulata M86]

*Type species of generic name indicated

REFERENCES

[B85] Bellamy, C. L. 1985. A catalogue of the higher taxa of the family Buprestidae (Coleoptera). Navorsinge van die Nasionale Museum Bloemfontein 4 (15): 405–472.

[B35] Boisduval, J. B. 1835. Voyage de Découvertes de l’Astrolabe. Exécuté par ordre du Roi, pendant les années 1826–1827–1828–1829, sous le commandement de M. J. Dumont d'Urville. Faune entomologique de l'océan Pacifique, avec l'illustration des insectes nouveaux recueillis pendant le voyage vol. 2. Coléoptères et autres ordres. J. Tastu: Paris.

[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature's gems. Ivy Press: Lewes (United Kingdom).

[L58] Linnaeus, C. 1758. Systema Naturae per Regna Tria Naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata. Laurentii Salvii: Holmiae.

[M86] Masters, G. 1886. Catalogue of the described Coleoptera of Australia. Part III. Proceedings of the Linnean Society of New South Wales, series 2, 1 (1): 21–126.