Austrostipa

Soft speargrass Austrostipa mollis, from here.


Belongs within: Poaceae.

Austrostipa, speargrass, is a genus of grasses endemic to Australasia.

Characters (from New South Wales Flora Online): Perennials (rarely annuals) of various habit. Auricles often present, sometimes one-sided, grading into an usually membranous ligule. Blade rolled or folded in bud, convolute, terete or flat, usually scabrous or hairy. Inflorescence an open or contracted panicle, axis persistent or falling as a whole. Spikelets borne singly, pedicellate, disarticulating above the glumes; 1 bisexual floret. Glumes equal or unequal. 1–5-nerved, longer than the floret; a long or short bearded callus below each floret. Lemma terete, hardened, with convolute or involute margins enclosing the palea, usually 5-nerved, with or without 1–2 minute lobes, awned from the tip or between the lobes; awn with once or twice geniculate twisted column and a straight or curved bristle. Palea membranous. 0–2-nerved, longer or shorter than the lemma. Lodicles 3 or 2.

<==Austrostipa
    |--A. acrociliata G04b
    |--A. compressa RL05
    |--A. crinita KM08
    |--A. elegantissima KM08
    |--A. eremophila B00
    |--A. flavescens SM06
    |--A. hemipogon OS04
    |--A. juncifolia GK00
    |--A. mollis OS04
    |--A. nitida KM08
    |--A. platychaeta G04b
    |--A. pycnostachya GK00
    |--A. scabra B00
    |--A. semibarbata B00
    |--A. tenuifolia OS04
    |--A. trichophylla G04a
    `--A. variabilis OS04

*Type species of generic name indicated

REFERENCES

[B00] Braby, M. F. 2000. Butterflies of Australia: their identification, biology and distribution vol. 1. CSIRO Publishing: Collingwood (Victoria).

[G04a] Gibson, N. 2004a. Flora and vegetation of the Eastern Goldfields Ranges: part 6. Mt Manning Range. Journal of the Royal Society of Western Australia 87 (2): 35–47.

[G04b] Gibson, N. 2004b. Flora and vegetation of the Eastern Goldfields Ranges: part 7. Middle and South Ironcap, Digger Rock and Hatter Hill. Journal of the Royal Society of Western Australia 87 (2): 49–62.

[GK00] Gibson, N., & G. J. Keighery. 2000. Flora and vegetation of the Byenup-Muir reserve system, south-west Western Australia. CALMScience 3 (3): 323–402.

[KM08] Keighery, G. J., & W. Muir. 2008. Vegetation and vascular flora of Faure Island, Shark Bay, Western Australia. Records of the Western Australian Museum Supplement 75: 11–19.

[OS04] Obbens, F. J., & L. W. Sage. 2004. Vegetation and flora of a diverse upland remnant of the Western Australian wheatbelt (Nature Reserve A21064). Journal of the Royal Society of Western Australia 87 (1): 19–28.

[RL05] Rafferty, C., & B. B. Lamont. 2005. Selective feeding by macropods on vegetation regenerating following fire. Journal of the Royal Society of Western Australia 88 (4): 155–165.

[SM06] Semeniuk, C. A., L. A. Milne, P. Ladd & V. Semeniuk. 2006. Pollen in the surface sediments of wetlands in the Becher Point area, southwestern Australia: a baseline for use in interpreting Holocene sequences. Journal of the Royal Society of Western Australia 89 (1): 27–43.

Deyeuxia

Reed bent-grass Deyeuxia quadriseta, from here.


Belongs within: Pooideae.

Deyeuxia is a genus of grasses found in temperate habitats.

Characters (from Cheeseman 1906): Annual or perennial grasses. Leaves flat or involute ; ligules membranous. Spikelets small, 1-flowered, arranged in effuse or contracted or spike-like panicles with capillary whorled branches; rhachilla disarticulating above the 2 outer glumes, produced beyond the flower into a silky bristle. Glumes 3; 2 outer equal or subequal, persistent, empty, keeled, acute, not awned, usually 1-nerved; 3rd or flowering glume shorter than the empty glumes or equalling them, thin and hyaline or rigidly membranous or almost coriaceous, 5-nerved, entire or 2-4-dentate, callus at the base silky; awn generally present, straight or twisted, inserted above or below the middle of the glume. Palea more than half as long as the flowering glume or almost equalling it, thin, 2-nerved or 2-keeled. Stamens 3. Styles distinct, short; stigmas plumose. Grain oblong or obovoid, enclosed within the flowering glume and palea.

<==Deyeuxia
    |--D. aucklandica W91
    |--D. avenoides [=Agrostis avenoides] C06
    |    |--D. a. var. avenoides C06
    |    `--D. a. var. brachyantha Hack. in Cheeseman 1906 C06
    |--D. billardieri [=Agrostis billardieri] C06
    |    |--D. b. var. billardieri C06
    |    `--D. b. var. tenuis Petrie in Cheeseman 1906 C06
    |--D. filiformis A27
    |--D. forsteri [=Agrostis forsteri; incl. Ag. aemula, Avena filiformis, Ag. solandri] C06
    |    |--D. f. var. forsteri C06
    |    |--D. f. var. humilior Hack. in Cheeseman 1906 [incl. Agrostis striata Col. 1889] C06
    |    |--D. f. var. littoralis Hack. in Cheeseman 1906 C06
    |    |--D. f. var. lyallii Hack. in Cheeseman 1906 [=Agrostis lyallii] C06
    |    |--D. f. var. micrathera Hack. in Cheeseman 1906 C06
    |    |--D. f. var. pilosa Cheeseman 1906 [=Agrostis pilosa] C06
    |    `--D. f. var. semiglabra Hack. in Cheeseman 1906 C06
    |--D. holciformis O88
    |--‘Agrostis’ leptostachya C06
    |--‘Calamagrostis’ lilloi Hackel in Stuckert 1906 S06
    |--‘Calamagrostis’ malamalensis Hackel in Stuckert 1906 S06
    |--D. nepalensis O88
    |--D. nivicola O88
    |--D. pulchella O88
    |--D. quadriseta [=Agrostis quadriseta, Avena quadriseta] C06
    |--D. scabra [=Agrostis scabra] C06
    |--D. scabrescens O88
    |--D. setifolia [=Agrostis setifolia] A27
    |--‘Calamagrostis’ spiciformis Hackel in Stuckert 1906 S06
    `--D. youngii [=Agrostis youngii] C06
         |--D. y. var. youngii C06
         `--D. y. var. petriei [=Calamagrostis petriei Hack. 1903] C06

*Type species of generic name indicated

REFERENCES

[A27] Andersen, J. C. 1927. Popular names of New Zealand plants. Transactions and Proceedings of the New Zealand Institute 57: 905–977.

[C06] Cheeseman, T. F. 1906. Manual of the New Zealand Flora. John Mackay, Government Printer: Wellington.

[O88] Ohba, H. 1988. The alpine flora of the Nepal Himalayas: an introductory note. In: Ohba, H., & S. B. Malla (eds) The Himalayan Plants vol. 1. The University Museum, University of Tokyo, Bulletin 31: 19–46.

[S06] Stuckert, T. 1906. Segunda contribución al conocimiento de las gramináceas Argentinas. Anales del Museo Nacional de Buenos Aires, serie 3, 6: 408–555.

[W91] Williams, P. A. 1991. Subalpine and alpine vegetation of granite ranges in western Nelson, New Zealand. New Zealand Journal of Botany 29: 317–330.

Triodia

Triodia pungens, copyright Mark Marathon.


Belongs within: Chloridoideae.

Triodia, spinifex, is a genus of grasses with rigid, spine-like leaves.

Characters (from Cheeseman 1906): Perennial grasses, of very various habit. Leaves narrow, rigid. Spikelets 2- to many-flowered, arranged in a lax or narrow panicle; rhachilla disarticulating above the two outer glumes and between the flowering glumes. Two outer glumes longer or shorter than the flowering glumes, somewhat rigid, empty, keeled, acute, awnless. Flowering glumes more or less imbricated, rounded on the back at the base, coriaceous or chartaceous, often hairy on the margins and callus, 3-nerved, 3-lobed or 3-toothed at the apex, the lobes equal or the central one produced into a short awn or mucro. Palea broad, thin, with 2 almost marginal keels. Lodicules 2. Stamens 3. Styles short, distinct; stigmas plumose. Grain usually compressed on the back, free within the flowering glume and palea.

Triodia
    `--T. subg. Rhombolytrum C06
         |--T. australis Petrie 1890 C06
         |    |--T. a. var. australis C06
         |    `--T. a. var. mucronulata Hack. in Cheeseman 1906 C06
         |--T. exigua Kirk 1882 C06
         `--T. pumila Hack. in Cheeseman 1906 [=Atropis pumila Kirk 1882] C06

Triodia incertae sedis:
  T. acuminata S06
  T. acutispicula LK14
  T. aeria LK14
  T. angusta P-JO04
  T. basedowii B05
  T. bitextura EF04
  T. brizoides EF04
  T. burbidgeana LK14
  T. bynoei LK14
  T. caelestialis LK14
  T. claytonii LK14
  T. decumbens C06
  T. epactia LK14
  T. irritans LF10
  T. lanigera HD04
  T. longiceps HD04
  T. longiloba LK14
  T. melvillei B05
  T. microstachya WF01
  T. mitchellii B00
  T. paraguayensis S06
  T. plurinervata KM08
  T. pungens HD04
  T. rigidissima G04
  T. scariosa CWB06
  T. schinzii B05
  T. secunda HD04
  T. stenostachya LK14
  T. wiseana BS05

*Type species of generic name indicated

REFERENCES

[B05] Beard, J. S. 2005. Drainage evolution in the Lake Disappointment Catchment, Western Australia—a discussion. Journal of the Royal Society of Western Australia 88 (2): 57–64.

[B00] Braby, M. F. 2000. Butterflies of Australia: their identification, biology and distribution vol. 1. CSIRO Publishing: Collingwood (Victoria).

[BS05] Butcher, R., & L. W. Sage. 2005. Tetratheca fordiana (Elaeocarpaceae), a new species from the Pilbara of Western Australia. Journal of the Royal Society of Western Australia 88 (2): 73–76.

[C06] Cheeseman, T. F. 1906. Manual of the New Zealand Flora. John Mackay, Government Printer: Wellington.

[CWB06] Craig, M. D., P. C. Withers & S. D. Bradshaw. 2006. Patterns of diet and microhabitat use by four species of sympatric Ctenotus lizards: do they reveal foraging specialisation? Journal of the Royal Society of Western Australia 89 (1): 1–5.

[EF04] Etten, E. J. B. van, & J. E. D. Fox. 2004. Vegetation classification and ordination of the central Hamersley Ranges, Western Australia. Journal of the Royal Society of Western Australia 87 (2): 63–79.

[G04] Gibson, N. 2004. Flora and vegetation of the Eastern Goldfields Ranges: part 6. Mt Manning Range. Journal of the Royal Society of Western Australia 87 (2): 35–47.

[HD04] How, R. A., & J. Dell. 2004. Reptile assemblage of the Abydos Plain, north-eastern Pilbara, Western Australia. Journal of the Royal Society of Western Australia 87 (3): 85–95.

[KM08] Keighery, G. J., & W. Muir. 2008. Vegetation and vascular flora of Faure Island, Shark Bay, Western Australia. Records of the Western Australian Museum Supplement 75: 11–19.

[LF10] Langlands, P. R., & V. W. Framenau. 2010. Systematic revision of Hoggicosa Roewer, 1960, the Australian 'bicolor' group of wolf spiders (Araneae: Lycosidae). Zoological Journal of the Linnean Society 158: 83–123.

[LK14] Lyons, M. N., G. J. Keighery, L. A. Gibson & T. Handasyde. 2014. Flora and vegetation communities of selected islands off the Kimberley coast of Western Australia. Records of the Western Australian Museum Supplement 81: 205–244.

[P-JO04] Pruett-Jones, S., & E. O'Donnell. 2004. Land birds on Barrow Island: status, population estimates, and responses to an oil-field development. Journal of the Royal Society of Western Australia 87 (3): 101–108.

[S06] Stuckert, T. 1906. Segunda contribución al conocimiento de las gramináceas Argentinas. Anales del Museo Nacional de Buenos Aires, serie 3, 6: 408–555.

[WF01] Woinarski, J. C. Z., A. Fisher, K. Brennan, I. Morris & R. Chatto. 2001. Patterns of bird species richness and composition on islands off Arnhem Land, Northern Territory, Australia. Austral Ecology 26: 1–13.

Sejoidea

Sejus sp., copyright Pierre-Marc Brousseau.


Belongs within: Mesostigmata.

The Sejoidea are a group of densely setose mesostigmatan mites with fragmented dorsal and ventral plates (Lindquist et al. 2009).

Characters (from Lindquist et al. 2009): Body with moderate to extensive hypertrichy, numerous gland openings, often densely tuberculate. Adult female dorsally with podonotal and pygidial shields (sometimes extensive) and usually with mesonotal or mesolateral shields; secondary hardening of the cuticle often obscures shield extent and number. Peritremes and peritrematic shields usually well developed, the latter fused anteriorly to the podonotal shield and sometimes included in a process over the gnathosoma (vertex); lateral setae often on small platelets. Venter of female with large ventrianal shield (sometimes developed dorsolaterally and fused to pygidial shield); metapodal shields usually free and well developed; epigynal shield large, usually bearing 6 to numerous setae (rarely 1–2 pairs) and covering the opening to a large, saclike spermatheca. Sternal area not fully sclerotised, but with seta-bearing platelets (sometimes obscured by genital shield) and often a shield bearing two pairs of setae. Males with more consolidated body shields; male genital aperture on an intercoxal shield, covered by a subcircular valve. Tarsus I ofen with an acrotarsus, with or without claws; tarsus IV with 20 or more setae, including setae av4 and pv4 on well-defined intercalary sclerite. Tritosternum with well-developed base and separate laciniae. Hypostomatic setae h1 often flattened, scalelike; corniculi entire or bifid. Palp apotele 2-tined. Chelicerae chelate-dentate, slender and with setiform pilus dentilis, to massive and without pilus dentilis; movable digit without arthrodial brush or excrescences. Phoretic deutonymph with a cylindrical anal process and 1–2 dorsal shields.

Sejoidea [Liroaspina, Sejida, Sejina]
    |--Uropodella Berlese 1888 FH93 [Uropodellidae LKW09]
    |    |--U. laciniata Berlese 1888 [incl. U. camini Hirschmann & Zierngiebl-Nicol 1984] FH93
    |    `--U. krantzi Hirschmann & Zirngiebl-Nicol 1984 FH93
    |--Ichthyostomatogasteridae LKW09
    |    |--Japanoasternolaelaps LKW09
    |    |--Archaeopodella Athias-Henriot 1977 LKW09, H98
    |    |    `--A. scopulifera Athias-Henriot 1977 H98
    |    `--Asternolaelaps Berlese 1923 [incl. Ichthystomatogaster Sellnick 1953] ET79
    |         |  i. s.: A. australis Womersley & Domrow 1959 H98
    |         |--*A. (Asternolaelaps) fecundus Berlese 1923 [incl. Ichthystomatogaster nyhleni Sellnick 1953] ET79
    |         `--A. (Parasternolaelaps Athias-Henriot 1972) LKW09
    |              `--A. (*P.) castrii LKW09
    `--Sejidae [Liroaspidae, Liroaspoidea, Seiidae] LKW09
         |--Epicroseius Berlese 1905 FH93
         |--Adenosejus krantzi LKW09
         |--Africasejus LKW09
         |--Zuluacarus LKW09
         |--Liroaspis Banks 1902 H98
         |    `--L. acanthurus (Canestrini 1884) [=Sejus acanthurus, Seius acanthurus, Zercon acanthurus] H98
         `--Sejus Koch 1836 ET79 (see below for synonymy)
              |--*S. togatus Koch 1836 ET79 [=Liroaspis togatus S61]
              |--S. aciculatus Evans & Hyatt 1960 FH93
              |--S. americanus (Banks 1902) [=Liroaspis americana] FH93
              |--S. armatus (Fox 1947) [=Liroaspis armatus] FH93
              |--S. baloghi LKW09
              |--S. borealis (Berlese 1904) K91
              |--S. insulanus Trägårdh 1931 T31
              |--‘Seius’ italicus Berlese 1916 H98
              |--S. necorniger (Oudemans 1903) K91
              |--S. paricornis Berlese 1916 H98
              |--S. porosus (Domrow 1957) [=Epicroseius porosus] H98
              |--S. sanborni Packard 1888 FH93
              |--S. serratus (Halbert 1915) K91 [=*Microcheles serratus S58, Zercon serratus S58]
              `--‘Willmannia’ sejiformis QA91

Sejus Koch 1836 ET79 [incl. Dwigubskyia Oudemans 1936 ET79, Microcheles S58, Willmannia Balogh 1938 FH93]

*Type species of generic name indicated

REFERENCES

[ET79] Evans, G. O., & W. M. Till. 1979. Mesostigmatic mites of Britain and Ireland (Chelicerata: Acari-Parasitiformes). An introduction to their external morphology and classification. Transactions of the Zoological Society of London 35: 139–270.

[FH93] Farrier, M. H., & M. K. Hennessey. 1993. Soil-inhabiting and free-living Mesostigmata (Acari-Parasitiformes) from North America: an annotated checklist with bibliography and index. North Carolina Agricultural Research Service, North Carolina State University, Technical Bulletin 302: i–xvi, 1–408.

[H98] Halliday, R. B. 1998. Mites of Australia: A checklist and bibliography. CSIRO Publishing: Collingwood.

[K91] Koehler, H. H. 1991. A five year study on the secondary succession of Gamasina on a ruderal site: the influence of recultivation. In: Dusbábek, F., & V. Bukva (eds) Modern Acarology: Proceedings of the VIII International Congress of Acarology, held in České Budĕjovice, Czechoslovakia, 6–11 August 1990 vol. 1 pp. 373–383. SPB Academic Publishing: The Hague.

[LKW09] Lindquist, E. E., G. W. Krantz & D. E. Walter. 2009. Order Mesostigmata. In: Krantz, G. W. & D. E. Walter (eds) A Manual of Acarology 3rd ed. pp. 124–232. Texas Tech University Press.

[QA91] Quintero, M. T., & H. A. Acevedo. 1991. Studies on deep litter mites on farms in Mexico. In: Dusbábek, F., & V. Bukva (eds) Modern Acarology: Proceedings of the VIII International Congress of Acarology, held in České Budĕjovice, Czechoslovakia, 6–11 August 1990 vol. 1 pp. 443–448. SPB Academic Publishing: The Hague.

[S61] Schweizer, J. 1961. Die Landmilben der Schweiz (Mittelland, Jura und Alpen): Parasitiformes Reuter, mit 246 Arten und Varietäten und 268 meist kombinierten Originalzeichnungen. Denkschriften der Schweizerischen Naturforschenden Gesellschaft [Mémoires de la Société Helvétique des Sciences Naturelles] 84: i–vii, 1–207.

[S58] Sellnick, M. 1958. Die familie Zerconidae Berlese. Acta Zoologica Academiae Scientiarum Hungaricae 3: 313–368.

[T31] Trägårdh, I. 1931. Acarina from the Juan Fernandez Islands. Natural History of Juan Fernandez and Easter Islands 3 (4): 553–628.

Haemaphysalis

Bush tick Haemaphysalis longicornis, copyright Commonsource.


Belongs within: Ixodidae.

Haemaphysalis is a genus of ticks feeding on birds and mammals primarily found in the Old World (Keirans 2009).

Characters (from Keirans 2009): Ornate; eyes absent; palpi short, with segment II as broad as long and usually much broader than long, obviously extended laterally, basis capituli dorsally rectangular.

<==Haemaphysalis Koch 1844 [Haemaphysalinae] KDB02
    |--H. (Haemaphysalis) FKK91
    |    |--H. (H.) concinna Koch 1844 FKK91, VR91
    |    |--H. (H.) flava FKK91
    |    |--H. (H.) japonica FKK91
    |    `--H. (H.) megaspinosa FKK91
    |--H. (Aboimisalis) punctata Canestrini & Fanzago 1877 FKK91, CF77
    |--H. (Alloceraea) kitaokai Hoogstraal 1969 KDB02
    |--H. (Gonixodes) leporispalustris F91
    |--H. (Kaiseriana) FKK91
    |    |--H. (K.) bancrofti Nuttall & Warburton 1915 FKK91, H98 [incl. H. leachi var. australis Neumann 1905 H98]
    |    |--H. (K.) cornigera FKK91
    |    |    |--H. c. cornigera FKK91
    |    |    `--H. c. ias FKK91
    |    |--H. (K.) hystricis FKK91
    |    |--H. (K.) longicornis Neumann 1901 FKK91, KDB02 (see below for synonymy)
    |    |--H. (K.) mageshimaensis FKK91
    |    `--H. (K.) yeni FKK91
    `--H. (Ornithophysalis) humerosa Warburton & Nuttall 1909 FKK91, T13

Haemaphysalis incertae sedis:
  H. bremneri Roberts 1963 H98
  H. doenitzi Warburton & Nuttall 1909 H98
  H. erinacei Pavesi 1884 [incl. H. numidiana Neumann 1897] A56
  H. inermis Birula 1895 VR91
  H. konigsbergeri Warburton & Nuttall 1909 H98
  H. lagostrophi Roberts 1963 H98
  H. leachi K09
  H. novaeguineae Hirst 1914 [=H. spinigera var. novaeguineae] H98
  H. petrogalis Roberts 1970 H98
  H. ratti Kohls 1948 H98
  H. rhinolophi Canestrini & Fanzago 1877 CF77
  H. sulcata Canestrini & Fanzago 1877 CF77

Haemaphysalis (Kaiseriana) longicornis Neumann 1901 FKK91, KDB02 [=H. concinna var. longicornis H98; incl. H. neumanni Dönitz 1905 H98]

*Type species of generic name indicated

REFERENCES

[A56] Anastos, G. 1956. The ticks (Acarina: Ixodoidea) of the J. Klapperich Afghanistan Expedition, 1952 and 1953. Journal of the Washington Academy of Sciences 46 (1): 18–19.

[CF77] Canestrini, G., & F. Fanzago. 1877. Intorno agli Acari Italiani. Atti del Reale Istituto Veneto di Scienze, Lettere ed Arti, Serie 5, 4: 69–208, pls 2–7.

[F91] Filippova, N. A. 1991. A hypothesis for the palaeogenesis of the distribution of the main vectors for Lyme disease. In: Dusbábek, F., & V. Bukva (eds) Modern Acarology: Proceedings of the VIII International Congress of Acarology, held in České Budĕjovice, Czechoslovakia, 6–11 August 1990 vol. 1 pp. 109–118. SPB Academic Publishing: The Hague.

[FKK91] Fujisaki, K., T. Kamio & S. Kawazu. 1991. Theileria sergenti cannot be regarded as the same species as T. buffeli and T. orientalis because of its transmissibility only by Kaiseriana ticks. In: Dusbábek, F., & V. Bukva (eds) Modern Acarology: Proceedings of the VIII International Congress of Acarology, held in České Budĕjovice, Czechoslovakia, 6–11 August 1990 vol. 1 pp. 233–237. SPB Academic Publishing: The Hague.

[H98] Halliday, R. B. 1998. Mites of Australia: A checklist and bibliography. CSIRO Publishing: Collingwood.

[K09] Keirans, J. E. 2009. Order Ixodida. In: Krantz, G. W., & D. E. Walter (eds) A Manual of Acarology 3rd ed. pp. 111–123. Texas Tech University Press.

[KDB02] Klompen, H., S. J. Dobson & S. C. Barker. 2002. A new subfamily, Bothriocrotoninae n. subfam., for the genus Bothriocroton Keirans, King & Sharrad, 1994 status amend. (Ixodida: Ixodidae), and the synonymy of Aponomma Neumann, 1899 with Amblyomma Koch, 1844. Systematic Parasitology 53: 101–107.

[T13] Taylor, C. K. 2013. Annotated bibliography for Barrow Island terrestrial invertebrates. Records of the Western Australian Museum Supplement 83: 135–144.

[VR91] Výrosteková, V., J. Řeháček, D. Guryčová & E. Kocianová. 1991. Prevalence of Francisella tularensis in ticks of Slovakia. In: Dusbábek, F., & V. Bukva (eds) Modern Acarology: Proceedings of the VIII International Congress of Acarology, held in České Budĕjovice, Czechoslovakia, 6–11 August 1990 vol. 2 pp. 55–59. SPB Academic Publishing: The Hague.

Uropodoidea

Dinychus sp., copyright Scott Justis.


Belongs within: Uropodina.
Contains: Trichocylliba, Oplitis, Discourella, Trachyuropoda, Nenteria, Uroobovella, Trichouropoda, Uropoda.

The Uropodoidea are a diverse group of mites found in soil, humus, animal nests or decaying organic matter (Lindquist et al. 2009).

Characters (from Lindquist et al. 2009): Coxae I broad, covering or partially covering narrow tritosternal base, vertex and camerostome well developed, pedofossae often present; palpgenu with 4–5 setae. Dorsal shield of adults smooth or variously ornamented, sometimes with deep pits and ridges, entire or with a posterior pygidial element, free or contiguous with holoventral shield, fused anteriorly to peritrematic elements and often bordered by marginal scutella or coalesced marginal shields; pygidial shield, when present, may be narrow to broad, sometimes subdivided, with or without setae. Chelicerae elongate, sometimes appearing four-segmented, without lyrifissure, terminal digits usually small and often bearing elaborate sensory structures, pilus dentilis present or absent, levator tendon with or without a sclerotized node. Peritremes well developed and often convoluted or serpentine, tritosternal laciniae fused basally and often subdivided distally. Femur I with 8–9 setae, genua III–IV without (or occasionally with 1) ventral setae, tibia I with only 3 dorsal and 2 ventral setae (typically 1 1/1, 2/1 1). Female genital opening covered by a single nude epigynal shield, paraanal elements absent. Male with holoventral shield, genital opening between coxae II–IV (rarely postcoxal), covered by 1–2 valves; protonymph with separate or fused podonotal and mesonotal elements; phoretic deutonymphs may attach to host via strand produced by anal pedicel.

<==Uropodoidea [Prodinychini] LKW09
    |--Clausiadinychus Sellnick 1930 KA90, FH93 [Clausiadinychidae FH93]
    |    `--C. cristatus Sellnick 1930 FH93
    |--Thrichocyllibidae [Circocyllibidae, Coxequesomidae, Planodiscidae, Trichocyllibidae] FH93
    |    |--Trichocylliba ET79
    |    |--Antennequesoma Sellnick 1926 FH93
    |    `--Hirtitrichocyllibula Hirschmann 1979 FH93
    |--Metagynella Berlese in Trouessart & Berlese 1919 FH93 (see below for synonymy)
    |    |--*M. paradoxa Berlese 1919 ET79
    |    |--M. lindquisti Hirschmann 1979 FH93
    |    |--M. mexicana Hirschmann 1979 FH93
    |    |--M. moseri Hirschmann 1975 FH93
    |    `--M. parvula Camin 1953 FH93
    |--Eutrachytidae FH93
    |    |--Dentibaiulus Schuster 1962 FH93
    |    |--Dicornutophorus Hirschmann 1979 FH93
    |    |--Mayaphorus Hirschmann 1979 FH93
    |    `--Eutrachytes Berlese 1914 S61, FH93
    |         `--E. gaieri Schweizer 1961 S61
    |--Prodinychidae FH93
    |    |--Allodinychus Tragardh 1943 FH93
    |    |--Trachyxenura Leitner 1947 FH93
    |    `--Prodinychus Berlese 1917 ET79, FH93
    |         |--*P. fimicolus (Berlese 1903) [=Dinychus fimicolus] ET79
    |         |--P. americanus (Banks 1904) (see below for synonymy) FH93
    |         |--P. flagellifer (Berlese 1910) FH93 [=Dinychus flagellifer FH93, Allodinychus flagellifer S61]
    |         |--P. hummelincki Sellnick 1963 FH93
    |         `--P. tetraphyllus (Berlese 1903) FH93 [=Dinychus tetraphyllus FH93, Phyllodinychus tetraphyllus S61]
    |--Macrodinychidae FH93
    |    |--Baloghmacrodinychus Hirschmann 1979 FH93
    |    |--Bregetovamacrodinychus Hirschmann 1979 FH93
    |    |--Loksamacrodinychus Hirschmann 1979 FH93
    |    `--Macrodinychus Berlese 1917 FH93
    |         |  i. s.: M. andrassyi Hirschmann 1975 H98
    |         |         M. baloghi Hirschmann 1975 H98
    |         |         M. parallelepipidus KGG07
    |         |--M. (Macrodinychus) VKC07
    |         |    |--M. (M.) paraguayensis VKC07
    |         |    `--M. (M.) sellnicki Hirschmann & Zirngiebl-Nicol 1974 FH93
    |         `--M. (Monomacrodinychus) multispinosus Sellnick 1973 FH93
    |--Oplitidae LKW09
    |    |--Oplitis ET79
    |    |--Berleseoplitis Hirschmann 1979 FH93
    |    |--Cariboplitis Sellnick 1963 FH93
    |    |--Chelonuropoda Sellnick 1954 FH93
    |    |--Marginura Sellnick 1926 FH93
    |    |--Sellnickoplitis Hirschmann 1979 FH93
    |    |--Stammeroplitis Hirschmann 1979 FH93
    |    |--Uropectinia Kneissl 1908 FH93
    |    |--Uroplitana Sellnick 1926 FH93
    |    |--Woelkeoplitis Hirschmann 1979 FH93
    |    `--Womersleyoplitis Hirschmann 1979 FH93
    |--Uroactiniidae [Uroactiniinae] LKW09
    |    |--Chiropturopoda Sellnick 1958 FH93
    |    |--Centrouropoda Berlese 1916 FH93
    |    |    `--C. rackae Hirschmann 1975 FH93
    |    `--Uroactinia Hirschmann & Zirngiebl-Nicol 1964 H98
    |         |--*U. consanguinea [=Uropoda consanguinea] D81
    |         |--U. agitans (Banks in Newell 1908) LKW09, FH93 [=Uropoda agitans] FH93
    |         |--U. anchor (Trouessart 1902) [=Uropoda anchor; incl. Fuscuropoda hippocrepoides Vitzthum 1939] FH93
    |         |--U. centroamericana (Stoll 1893) [=Uropoda centroamericana] FH93
    |         |--U. domrowi Hirschmann 1990 H98
    |         |--U. hippocrepea (Berlese 1918) [=Fuscuropoda hippocrepea] FH93
    |         |--U. hirschmanni Hiramatsu 1978 FH93
    |         |--U. ovata (Fox 1948) [=Fuscuropoda ovata] FH93
    |         `--U. rarosae VKC07
    |--Discourellidae FH93
    |    |--Discourella LKW09
    |    |--Baloghurella Hirschmann 1979 FH93
    |    |--Capitodiscus Vitzthum 1931 FH93
    |    |--Clivosurella Hirschmann 1979 FH93
    |    |--Comydinychus Berlese 1917 FH93
    |    |--Cosmogynurella Hirschmann 1979 FH93
    |    |--Formosaurella Hirschmann 1979 FH93
    |    |--Gerlachurella Hirschmann 1979 FH93
    |    |--Longicarinaturella Hirschmann 1979 FH93
    |    |--Reticulaturella Hirschmann 1979 FH93
    |    |--Rotundurella Hirschmann 1979 FH93
    |    `--Stammerurella Hirschmann 1979 FH93
    |--Trachyuropodidae [Phymatodiscidae, Trachyuropodidae, Trachyuropodina, Trachyuropodini, Trachyuropodoidea] LKW09
    |    |--Phymatodiscus Berlese 1917 VKC07, FH93
    |    |--Trachyuropoda ET79
    |    |--Arculatatrachys Hirschmann 1979 FH93
    |    |--Bostocktrachys Hirschmann 1979 FH93
    |    |--Castritrachys Hirschmann 1979 FH93
    |    |--Cephalojanetia Willmann 1951 FH93
    |    |--Cephalouropoda Berlese 1903 FH93
    |    |--Cristicepstrachys Hirschmann 1979 FH93
    |    |--Excavatatrachys Hirschmann 1979 FH93
    |    |--Foliatrachys Hirschmann 1979 FH93
    |    |--Graecatrachys Hirschmann 1979 FH93
    |    |--Lindquisttrachys Hirschmann 1979 FH93
    |    |--Magnatrachys Hirschmann 1979 FH93
    |    |--Origmatrachys Hirschmann 1979 FH93
    |    |--Trogulotrachys Hirschmann 1979 FH93
    |    |--Leonardiella Berlese 1904 FH93
    |    |    `--*L. canestriniana (Berlese 1891) ET79, FH93 [=Uropoda canestriniana FH93]
    |    `--Urojanetia Berlese 1913 [incl. Janetiella Berlese 1904 (preoc.)] FH93
    |         `--U. cristipes S61
    |--Nenteriidae FH93
    |    |--Nenteria KA90
    |    |--Calurodiscus Radford 1950 FH93
    |    |--Castrinenteria Hirschmann 1979 FH93
    |    |--Stylinenteria Hirschmann 1979 FH93
    |    |--Austrodinychus Trägårdh 1952 FH93
    |    |    |--A. canadiensis (Hirschmann 1978) [=Nenteria canadiensis] FH93
    |    |    |--A. chiapasus (Hirschmann 1978) [=Nenteria chiapasa] FH93
    |    |    |--A. fungivorus (Hirschmann 1978) [=Nenteria fungivora] FH93
    |    |    |--A. howdeni (Hirschmann 1978) [=Nenteria howdeni] FH93
    |    |    |--A. martini (Hirschmann 1978) [=Nenteria martini] FH93
    |    |    |--A. mexicanus (Hirschmann 1978) [=Nenteria mexicana] FH93
    |    |    |--A. moseri (Hirschmann 1972) [=Nenteria moseri] FH93
    |    |    `--A. sellnicki (Hirschmann & Wiśniewski 1985) [=Nenteria sellnicki] FH93
    |    `--Ruhmnenteria Hirschmann 1979 FH93
    |         |--R. bretonensis (Hirschmann 1978) [=Nenteria bretonensis] FH93
    |         |--R. chihuahuaensis (Hirschmann 1978) [=Nenteria chihuahuaensis] FH93
    |         |--R. durangoensis (Hirschmann 1978) [=Nenteria durangoensis] FH93
    |         |--R. eutamiasae (Hirschmann 1978) [=Nenteria eutamiasae] FH93
    |         |--R. microti (Hirschmann 1978) [=Nenteria microti] FH93
    |         |--R. orghidani (Huţu 1977) [=Nenteria orghidani] FH93
    |         |--R. potosi (Hirschmann 1978) [=Nenteria potosi] FH93
    |         `--R. saltoensis (Hirschmann 1978) [=Nenteria saltoensis] FH93
    |--Dinychidae [Dinychini, Urodiaspididae] LKW09
    |    |--Uroobovella LKW09
    |    |--Dipolyaspis Berlese 1916 FH93
    |    |    `--*P. sansonei (Berlese 1916) [=Polyaspis (*Dipolyaspis) sansonei] B16
    |    |--Diurodinychus FH93
    |    |--Phyllodinychus Trägårdh 1943 FH93
    |    |--Lindquistidiaspis Hirschmann 1984 FH93
    |    |    `--L. lindquisti (Hirschmann 1979) [=Urodiaspis lindquisti] FH93
    |    |--Urodiaspis Berlese 1916 ET79
    |    |    |--*U. tecta (Kramer 1876) [=Notaspis tectus] ET79
    |    |    |--U. pannonica Willmann 1951 K07b
    |    |    `--U. walkeri Hirschmann & Zirngiebl-Nicol 1969 FH93
    |    |--Crinitodiscus Sellnick 1931 FH93
    |    |    `--C. (Oriendiscus) EU03
    |    |         |--C. (O.) pawlolowski EU03
    |    |         `--C. (O.) rafalskii EU03
    |    `--Dinychus Kramer 1886 FH93
    |         |--*D. perforatus Kramer 1882 ET79
    |         |--D. greensladeae Bloszyk & Halliday 1995 H98
    |         |--D. inermis (Koch 1841) [=Sejus inermis; incl. D. vartismithi Hull 1918] FH93
    |         |--D. micropunctatus Evans 1955 FH93
    |         |--D. ornatus (Fox 1957) [=Phyllodynchus ornatus[ FH93
    |         |--D. ovatus (Ewing 1909) [=Polyaspis ovatus] FH93
    |         |--D. ruseki Athias-Binche, Błoszyk & Olszanowski 1989 FH93
    |         `--D. septentrionalis (Trägårdh 1943) [=Phyllodinychus septentrionalis] FH93
    |--Trematuridae [Trematurellidae, Trematurini] LKW09
    |    |--Trichouropoda LKW09
    |    |--Trematurella Trägårdh 1942 FH93
    |    |--Dentidinychus Sellnick 1926 FH93
    |    |--Gitodinychus Berlese 1917 FH93
    |    |--Trematuroides Cooreman 1960 FH93
    |    |--Trichodinychus Berlese 1917 FH93
    |    |--Trichointerstructura Hirschmann 1979 FH93
    |    |--Trichosociata Hirschmann 1979 FH93
    |    |--Urolaelaps Berlese 1916 FH93
    |    |--Urospina Sellnick 1931 FH93
    |    |--Trematura Berlese 1917 ET79
    |    |    |--*T. patavina (Can. 1885) [=Uropoda patavina] ET79
    |    |    `--T. jacksoni ET79
    |    |--Leiodinychus Berlese 1917 FH93
    |    |    |--L. orbicularis LO91
    |    |    `--L. parasititucs SH01
    |    `--Oodinychus Berlese 1917 FH93
    |         |--*O. janeti (Berlese 1904) [=Urodinychus janeti] ET79
    |         |--O. granulatus Sellnick 1963 FH93
    |         |--O. jurassicus Schweizer 1961 S61
    |         |--O. karawaiewi Berlese 1903 S61
    |         |--O. margaritaensis Sellnick 1963 FH93
    |         |--O. ovalis (Koch 1839) MS91 [=Uropoda ovalis S61]
    |         `--O. venezolanus Sellnick 1963 FH93
    |--Urodinychidae [Rotundabaloghiidae, Urodinychini] VKC07
    |    |--Angulobaloghia Hirschmann 1979 FH93
    |    |--Appendiculobovella Hirschmann 1979 FH93
    |    |--Calouropoda Berlese 1916 FH93
    |    |--Dinychopsis Berlese 19126 FH93
    |    |--Formosaobovella Hirschmann 1979 FH93
    |    |--Indotrachytes Deb & Raychandhuri 1965 FH93
    |    |--Olouropoda Berlese 1916 FH93
    |    |--Paradinychus Berlese 1916 FH93
    |    |    `--*P. venustus Berlese 1916 B16
    |    |--Phaulotrachytes Oudemans 1913 FH93
    |    |--Polyaspidiella Berlese 1910 FH93
    |    |--Pulchellaobovella Hirschmann 1979 FH93
    |    |--Styluropoda Trägåardh 1952 FH93
    |    |--Urocyclella Berlese 1913 FH93
    |    |--Urocyclellopsis Willmann 1953 FH93
    |    |--Dinychura Berlese 1913 FH93
    |    |    `--D. cordieri Berlese 1916 [=Discourella cordieri] ET79
    |    |--Vinicoloraobovella Hirschmann 1979 FH93
    |    |    `--V. americana (Hirschmann 1972) (see below for synonymy) FH93
    |    |--Dendrouropoda Willmann 1959 FH93
    |    |    `--D. vallei S79
    |    |--Neoseius Oudemans 1903 FH93
    |    |    `--N. novus WP99
    |    |--Sellnickiobovella Hirschmann 1984 FH93
    |    |    |--S. hilli (Sellnick 1970) [=Fuscuropoda hilli] FH93
    |    |    `--S. marmorea (Fox 1948) [=Fuscuropoda marmorea] FH93
    |    |--Urosternella Berlese 1903 FH93
    |    |    |--U. cylindrica (Berlese 1913) [=Urodinychus cylindricus] FH93
    |    |    `--U. neptuni S79
    |    |--Urodinychus Berlese 1903 ET79
    |    |    |--*U. carinatus (Berlese 1888) [=Uropoda carinata] ET79
    |    |    |--U. karawaiewi S22
    |    |    `--U. ovalis S22
    |    `--Rotundabaloghia Hirschmann 1975 VKC07, K07a
    |         |--R. benyovszkyi Kontschán 2007 K07a
    |         |--R. liriformes Hirschmann 1992 VKC07
    |         `--R. madagascarensis Kontschán 2007 K07a
    `--Uropodidae [Castriidinychidae, Cillibidae, Deraiophoridae, Phaulodinychidae, Uropoden, Uropodini] LKW09
         |--Habeogula LKW09
         |--Uropoda LKW09
         |--Australiuropoda Hirschmann 1979 FH93
         |--Ceyloniphorus Hirschmann 1979 FH93
         |--Discotrachytes Berlese 1916 FH93
         |--Foveolaturopoda Hirschmann 1979 FH93
         |--Gibbauropoda Hirschmann 1979 FH93
         |--Heterodinychus Berlese 1912 FH93
         |--Interrupturopoda Hirschmann 1979 FH93
         |--Jerzywisniewskia Hirschmann 1979 FH93
         |--Karguropoda Hirschmann 1979 FH93
         |--Laqueaturopoda Hirschmann 1979 FH93
         |--Manuleatophorus Hirschmann 1979 FH93
         |--Neobirophorus Hirschmann 1979 FH93
         |--Multiporuropoda Hirschmann 1979 FH93
         |--Odonturopoda Marais 1977 FH93
         |--Penicillaturopoda Hirschmann 1979 FH93
         |--Spiculaturopoda Hirschmann 1979 FH93
         |--Spinosissuropoda Hirschmann 1979 FH93
         |--Dentatadinychus Hirschmann 1979 FH93
         |--Marginalidinychus Hirschmann 1979 FH93
         |--Toplaidinychus Hirschmann 1979 FH93
         |--Uropolyaspis Berlese 1903 ET79
         |    `--*U. hamulifera (Michael 1894) [=Uropoda hamulifera] ET79
         |--Cyclacarus Ewing 1933 FH93
         |    `--C. aberrans Ewing 1933 FH93
         |--Kaszabjbaloghia Hirschmann 1973 [Kaszabjbaloghiidae] H98
         |    `--K. hirschmanni Hiramatsu 1978 H98
         |--Olodiscus Berlese 1917 ET79
         |    |--*O. integer (Berlese 1910) [=Discopoma integra] ET79
         |    `--O. minima ET79
         |--Neodiscopoma Vitzthum 1943 [incl. Discopoma Berlese 1904 non Canestrini & Canestrini 1882] ET79
         |    |--*N. splendida (Kramer 1882) ET79 (see below for synonymy)
         |    `--N. pulcherrima (Berlese 1904) [=Discopoma pulcherrima] S61
         |--Metadinychus Berlese 1916 FH93
         |    |--M. argasiformis Berlese 1916 FH93
         |    |--M. exilis (Hiramatsu & Hirschmann 1979) [=Uropoda (Metadinychus) exilis] FH93
         |    `--M. pura (Hiramatsu & Hirschmann 1979) [=Uropoda (Metadinychus) pura] FH93
         |--Tuberdinychus Schweizer 1961 FH93
         |    |--*T. subterranus (Schweizer 1922) [=Urodinychus subterranus] S61
         |    |--T. fumicolus Schweizer 1961 S61
         |    `--T. parvus Schweizer 1961 S61
         |--Castriidinychus Hirschmann 1973 H98
         |    |--C. baloghi Hirschmann 1975 H98
         |    |--C. kaszabi Hirschmann 1975 H98
         |    `--C. mahunkai Hirschmann 1975 H98
         |--Diphaulocylliba Vitzthum 1925 FH93
         |    |--*D. amplior (Berlese 1923) [=Phaulocylliba amplior] V25
         |    |--D. debilior Vitzthum 1925 V25
         |    |--D. ingens Vitzthum 1925 V25
         |    `--D. permagna (Canestrini 1897) [=Discopoma permagna] V25
         |--Phaulocylliba Berlese 1903 FH93
         |    |--*P. ventricosa Berlese 1903 V25
         |    |--P. inflata Berlese 1920 V25
         |    |--P. littoralis (Trouessart 1902) [=Discopoma littorale; incl. P. berlesi Halbert 1915] V25
         |    |--P. misella (Berlese 1916) [=Cyllibano (Microcylliba) misella] V25
         |    |--P. orbicularis (Müller 1882) S61 (see below for synonymy)
         |    `--P. virgata (Hull 1918) [=Cillibano virgata] V25
         |--Deraiophorus Canestrini 1897 LKW09, FH93
         |    |--D. australis Hirschmann 1973 H98
         |    |--D. maya (Krantz 1969) LKW09, FH93 [=Eutrachytes maya FH93]
         |    |--D. neobiroi Hirschmann 1973 H98
         |    |--D. novaehollandiae (Domrow 1957) [=Urodiaspis novaehollandiae] H98
         |    |--D. simplicior (Domrow 1957) [=Eutrachytes simplicior] H98
         |    `--D. truncatus (Berlese 1888) [=Celaeno truncata] FH93
         `--Piracarus crozetensis P94

Metagynella Berlese in Trouessart & Berlese 1919 FH93 [incl. Metagynura Balogh 1943 FH93, Metagynellidae, Metagynuridae LKW09]

*Neodiscopoma splendida (Kramer 1882) ET79 [=Discopoma splendida ET79; incl. D. splendidae var. porticensis Berlese 1904 S61]

Phaulocylliba orbicularis (Müller 1882) S61 [incl. Discopoma romana Canestrini & Canestrini 1882 V25, P. romana S61]

Prodinychus americanus (Banks 1904) [=Dinychus americanus; incl. Uroobovella sellnickiamericana Hirschmann 1979] FH93

Vinicoloraobovella americana (Hirschmann 1972) [=Uroobovella americana, Fuscuropoda ameriana Moser & Roton 1971 (n. n.); incl. U. neoamericana Hirschmann & Zirngiebl-Nicol 1972] FH93

*Type species of generic name indicated

REFERENCES

[B16] Berlese, A. 1916. Centuria terza di Acari nuovi. Redia 12: 289–338.

[D81] Domrow, R. 1981. A small lizard stifled by phoretic deutonymphal mites (Uropodina). Acarologia 22 (3): 247–252.

[EU03] Ekiz, A. N., & R. Urhan. 2003. A new record of Uroobovella Berlese, 1903 (Acari: Uropodina) from Turkey with some ecological notes. Zoology in the Middle East 29: 105–111.

[ET79] Evans, G. O., & W. M. Till. 1979. Mesostigmatic mites of Britain and Ireland (Chelicerata: Acari-Parasitiformes). An introduction to their external morphology and classification. Transactions of the Zoological Society of London 35: 139–270.

[FH93] Farrier, M. H., & M. K. Hennessey. 1993. Soil-inhabiting and free-living Mesostigmata (Acari-Parasitiformes) from North America: an annotated checklist with bibliography and index. North Carolina Agricultural Research Service, North Carolina State University, Technical Bulletin 302: i–xvi, 1–408.

[H98] Halliday, R. B. 1998. Mites of Australia: A checklist and bibliography. CSIRO Publishing: Collingwood.

[K07a] Kontschán, J. 2007a. Two new Rotundabaloghia Hirschmann, 1975 species from Madagascar (Acari: Mesostigmata: Uropodina). Annales Historico-Naturales Musei Nationalis Hungarici 99: 171–176.

[K07b] Kontschán, J. 2007b. Some new records of Uropodina mites (Acari: Mesostigmata) from Croatia, Serbia and Montenegro with descriptions of two new species. Annales Historico-Naturales Musei Nationalis Hungarici 99: 177–188.

[KA90] Krantz, G. W., & B. D. Ainscough. 1990. Acarina: Mesostigmata (Gamasida). In: Dindal, D. L. (ed.) Soil Biology Guide pp. 583–665. John Wiley & Sones: New York.

[KGG07] Krantz, G. W., L. A. Gómez & V. E. Gonzalez. 2007. Parasitism in the Uropodina: a case history from Colombia. In: Morales-Malacara, J. B., V. M. Behan-Pelletier, E. Ueckermann, T. M. Pérez, E. G. Estrada-Venegas & M. Badii (eds) Acarology XI: Proceedings of the International Congress pp. 29–38. Instituto de Biología and Faculdad de Ciencias, Universidad Nacional Autónoma de México, Sociedad Latinoamericana de Acarología: México.

[LO91] Lehtinen, P. T., & I. Oksala. 1991. Astigmatic and prostigmatic mites of grain stores, mills and sawmills in Finland. In: Schuster, R., & P. W. Murphy (eds) The Acari: Reproduction, development and life-history strategies pp. 519–520. Chapman & Hall: London.

[LKW09] Lindquist, E. E., G. W. Krantz & D. E. Walter. 2009. Order Mesostigmata. In: Krantz, G. W., & D. E. Walter (eds) A Manual of Acarology 3rd ed. pp. 124–232. Texas Tech University Press.

[MS91] Miko, L., & M. Stanko. 1991. Small mammals as carriers of non-parasitic mites (Oribatida, Uropodina). In: Dusbábek, F., & V. Bukva (eds) Modern Acarology: Proceedings of the VIII International Congress of Acarology, held in České Budĕjovice, Czechoslovakia, 6–11 August 1990 vol. 1 pp. 395–402. SPB Academic Publishing: The Hague.

[P94] Pugh, P. J. A. 1994. Non-indigenous Acari of Antarctica and the sub-Antarctic islands. Zoological Journal of the Linnean Society 110: 207–217.

[S79] Schuster, R. 1979. Soil mites in the marine environment. Recent Advances in Acarology 1: 593–602.

[S22] Schweizer, J. 1922. Beitrag zur Kenntnis der terrestrischen Milbenfauna der Schweiz. Verhandl. Naturf. Ges. Basel 33: 23–112, 4 pls.

[S61] Schweizer, J. 1961. Die Landmilben der Schweiz (Mittelland, Jura und Alpen): Parasitiformes Reuter, mit 246 Arten und Varietäten und 268 meist kombinierten Originalzeichnungen. Denkschriften der Schweizerischen Naturforschenden Gesellschaft [Mémoires de la Société Helvétique des Sciences Naturelles] 84: i–vii, 1–207.

[SH01] Sumangala, K., & M. A. Haq. 2001. Survey of the mite fauna associated with Apis spp. in Kerala, southern India. In: Halliday, R. B., D. E. Walter, H. C. Proctor, R. A. Norton & M. J. Colloff (eds) Acarology: Proceedings of the 10th International Congress pp. 565–568. CSIRO Publishing: Melbourne.

[VKC07] Vázquez, M. M., H. Klompen & C. I. Chargoy. 2007. Study of the Uropodina (Acari: Mesostigmata) in tropical forests of the Yucatan Peninsula, Mexico and Belize. In: Morales-Malacara, J. B., V. M. Behan-Pelletier, E. Ueckermann, T. M. Pérez, E. G. Estrada-Venegas & M. Badii (eds) Acarology XI: Proceedings of the International Congress pp. 43–50. Instituto de Biología and Faculdad de Ciencias, Universidad Nacional Autónoma de México, Sociedad Latinoamericana de Acarología: México.

[V25] Vitzthum, H. 1925. Fauna sumatrensis. (Beitrag no. 5). Acarinae. Supplementa Entomologica 11: 1–79.

[WP99] Walter, D. E., & H. C. Proctor. 1999. Mites: Ecology, Evolution and Behaviour. CABI Publishing: Wallingford (UK).

Dermanyssidae

Chicken mite Dermanyssus gallinae, from here.


Belongs within: Gamasina.

The Dermanyssidae are a group of mites ectoparasitic on birds and mammals.

Characters (from Evans & Till 1979): Tritosternum present; chelicerae in nymphal stages and female with second article conspicuously elongated, slender, stylet-like with digits minute; tarsi III and IV of male with seta pv2 modified into short tooth-like projection.

<==Dermanyssidae [Dermanyssida, Dermanyssinae]
    |--Liponyssoides Hirst 1913 H98
    |    |--L. adsonis Domrow 1992 H98
    |    |--L. intermedius M68
    |    |--L. lukoschusi Domrow 1979 H98
    |    |--L. sanguineus R01
    |    `--L. warnekei Domrow 1963 H98
    `--Dermanyssus Dugès 1834 H98
         |  i. s.: D. lacertarum (Contarini 1843) [=Ricinus lacertarum] CF77
         |         D. lanius CF77
         |         D. richiardii Canestrini & Fanzago 1877 CF77
         |--D. (Dermanyssus) M68
         |    |--*D. (D.) gallinae (De Geer 1778) ET79 (see below for synonymy)
         |    |--D. (D.) chelidonis M68
         |    |--D. (D.) grochovskae M68
         |    |--D. (D.) hirsutus M68
         |    |--D. (D.) hirundinis (Hermann 1804) M68, H98 [=Acarus hirundinis CF77]
         |    |--D. (D.) prognephilus M68
         |    |--D. (D.) quintus M68
         |    |--D. (D.) transvaalensis M68
         |    `--D. (D.) triscutatus M68
         `--D. (Microdermanyssus Moss 1967) ET79
              |--D. (*M.) alaudae (Schrank 1781) [=Acarus alaudae] ET79
              |--D. (M.) americanus Ewing 1923 M68, H98
              `--D. (M.) brevis M68

*Dermanyssus (Dermanyssus) gallinae (De Geer 1778) ET79 [=Acarus gallinae CF77, Pulex gallinae CF77, Gamasus gallinae CF77; incl. Dermanyssus avium Dugès 1834 H98]

*Type species of generic name indicated

REFERENCES

[CF77] Canestrini, G., & F. Fanzago. 1877. Intorno agli Acari Italiani. Atti del Reale Istituto Veneto di Scienze, Lettere ed Arti, Serie 5, 4: 69–208, pls 2–7.

[ET79] Evans, G. O., & W. M. Till. 1979. Mesostigmatic mites of Britain and Ireland (Chelicerata: Acari-Parasitiformes). An introduction to their external morphology and classification. Transactions of the Zoological Society of London 35: 139–270.

[H98] Halliday, R. B. 1998. Mites of Australia: A checklist and bibliography. CSIRO Publishing: Collingwood.

[M68] Moss, W. W. 1968. Experiments with various techniques of numerical taxonomy. Systematic Zoology 17 (1): 31–47.

[R01] Russell, R. C. 2001. The medical significance of Acari in Australia. In: Halliday, R. B., D. E. Walter, H. C. Proctor, R. A. Norton & M. J. Colloff (eds) Acarology: Proceedings of the 10th International Congress pp. 535–546. CSIRO Publishing: Melbourne.

Astigmatina

Female Glycycometus sp., from here.


Belongs within: Desmonomata.
Contains: Histiostomatidae, Canestrinioidea, Hemisarcoptoidea, Acaroidea, Hypoderatidae, Psoroptidia, Glycyphagidae.

The Astigmatina are a group of soft-bodied mites lacking respiratory tracheae (Philips 1990). Many have rapid life cycles, and may be significant scavengers of decomposing matter. Most astigmatinans (excluding the vertebrate-associated Psoroptidia) have a modified non-feeding deutonymph (hypopus) stage in the life cycle that is adapted for phoretic dispersal or diapause during adverse conditions. However, if conditions are suitable, protonymphs may moult directly to the tritonymphal stage of the life cycle and bypass the deutonymph instar (OConnor 2009).

The basalmost member of the Astigmatina, Schizoglyphus biroi, is known only from two specimens collected from the tenebrionid beetle Dioedus tibialis; these specimens probably represent deutonymphs due to their lack of chelicerae and ventral attachment organ, but differ from deutonymphs of other astigmatinans in having three rather than two pairs of genital papillae (OConnor 2009).

See also: Life in the fast lane.

Characters (from Philips 1990): Weakly sclerotised, slow-moving, pale; respiratory stigmata and tracheae absent, respiration occuring through cuticle. Chelicerae mostly chelate-dentate. Palps two-segmented. Idiosoma usually divided into propodosoma and hysterosoma by sejugal furrow. Hysterosoma with laterodorsal hysterosomal gland openings normally visible. Primary genital opening of both sexes occuring ventrally between the legs, flanked by two pairs of genital papillae. Oviposition occuring through this opening (oviporus) in female, but sperm received through small circular opening (bursa copulatrix) at posterior end of body. Male aedeagus supported by struts; male anal opening often flanked by pair of copulatory suckers.

<==Astigmatina [Acaridei, Acaridiae, Acaridida, Astigmata, Diacotricha] O09
    |  i. s.: Myrmolichus greimae W01
    |         Amphicalvolia hurdi O09
    |--Schizoglyphus [Schizoglyphidae, Schizoglyphoidea] O09
    |    `--S. biroi O09
    `--+--Histiostomatoidea O09
       |    |--Histiostomatidae O09
       |    `--Guanolichidae O09
       |         |--Neoguanolichus P90
       |         `--Guanolichus gabonensis O09
       `--+--Canestrinioidea O09
          |--Hemisarcoptoidea O09
          `--+--+--Acaroidea O09
             |  `--+--Hypoderatidae O09
             |     `--Psoroptidia O09
             `--Glycyphagoidea O09
                  |--Aeroglyphidae [Aeroglyphinae] O09
                  |    |--Aeroglyphus robustus O09
                  |    `--Glycycometus Parameswaran Pillai 1957 [incl. Austroglycyphagus Fain & Lowry 1974] H98
                  |         |--G. geniculatus (Vitzthum 1919) (see below for synonymy) H98
                  |         |--‘Austroglycyphagus’ kualalumpurensis Fain & Nadchatram 1980 C-RSV-S88
                  |         |--G. lukoschusi O09
                  |         |--G. malaysiensis (Fain & Nadchatram 1980) C-R02, C-RSV-S88 (see below for synonymy)
                  |         |--‘Austroglycyphagus’ orientalis SH01
                  |         |--‘Austroglycyphagus’ thailandicus SH01
                  |         |--G. troglodytus (Fain & Lowry 1974) [=Austroglycyphagus troglodytus] H98
                  |         `--G. weelawadjiensis (Fain & Lowry 1974) [=Austroglycyphagus weelawadjiensis] H98
                  |--Rosensteiniidae O09
                  |    |--Rosensteinia sieversi O09
                  |    |--Mydopholeus O09
                  |    |--Chiroptoglyphus americanus O09
                  |    |--Micronychites postverrucosus O09
                  |    |--Cheiromelicus trochanteralis O09
                  |    |--Troglotacarus [Troglotacaridae] O09
                  |    |    `--T. hauseri O09
                  |    |--Lophonotacarus [Lophonotacaridae] O09
                  |    |    `--L. minutus O09
                  |    `--Nycteriglyphus Zakhvatkin 1941 H98
                  |         |--N. bifolium O09
                  |         |--N. dewae (Womersley 1963) [=Coproglyphus dewae] H98
                  |         |--N. miniopteri Fain 1963 H98
                  |         |--N. pterophorus O09
                  |         `--N. vespertilio O09
                  |--Euglycyphagidae O09
                  |    |--Euglycyphagus intercalatus O09
                  |    `--Lomelacarus O09
                  |--Pedetopus [Pedetopodidae] O09
                  |    `--P. zumpti O09
                  |--Echimyopodidae [Echimyopinae] O09
                  |    |--Echimyopus dasypus O09
                  |    |--Marmosopus mesoamericanus O09
                  |    |--Blomia Oudemans 1928 H98
                  |    |    `--B. tropicalis van Bronswijk, De Cock & Oshima 1974 H98
                  |    `--Marsupiopus Fain 1967 H98
                  |         |--M. acrobates Fain & Lukoschus 1976 H98
                  |         |--M. antechinus Fain & Lukoschus 1976 H98
                  |         |--M. leporilli Fain 1969 H98
                  |         |    |--M. l. leporilli H98
                  |         |    `--M. l. pseudomys Fain & Lukoschus 1976 H98
                  |         |--M. michaeli Fain 1969 H98
                  |         |--M. myrmecobius Fain & Lukoschus 1976 H98
                  |         |--M. trichosuri Fain 1967 [=Marsupiops (l. c.) trichosuri] H98
                  |         `--M. zyzomys Lukoschus et al. 1979 H98
                  |--Glycyphagidae O09
                  `--Chortoglyphidae [Chortoglyphina] O09
                       |--Chortoglyphus Berlese 1884 H98
                       |    `--C. arcuatus (Troupeau 1879) [=Tyroglyphus arcuatus] H98
                       |--Aplodontopus O09
                       |    |--A. marmotae O09
                       |    `--A. sciuricola Hyland & Fain 1968 M-PV-G01
                       `--Alabidopus Fain 1967 H98
                            |--A. asiaticus O09
                            |--A. hydromys Fain 1967 H98
                            `--A. muris Lukoschus et al. 1979 H98

Glycycometus geniculatus (Vitzthum 1919) [=Glycyphagus geniculatus, Austroglycyphagus geniculatus] H98

Glycycometus malaysiensis (Fain & Nadchatram 1980) C-R02, C-RSV-S88 [=Austroglycyphagus malaysiensis C-RSV-S88]

*Type species of generic name indicated

REFERENCES

[C-R02] Corpuz-Raros, L. A. 2002. Philippine acarine biological control agents: status, bioecology and research prospects. Philippine Agricultural Scientist 85 (2): 137–154.

[C-RSV-S88] Corpuz-Raros, L. A., G. C. Sabio & M. Velasco-Soriano. 1988. Mites associated with stored products, poultry houses and house dust in the Philippines. Philippine Entomologist 7 (3): 311–321.

[H98] Halliday, R. B. 1998. Mites of Australia: A checklist and bibliography. CSIRO Publishing: Collingwood.

[M-PV-G01] Montiel-Parra, G., G. A. Villegas-Guzman, M. Vargas & O. J. Polaco. 2001. Mites associated with nests of Neotoma albigula Hartley, 1894 (Rodentia: Muridae) in Durango, México. In: Halliday, R. B., D. E. Walter, H. C. Proctor, R. A. Norton & M. J. Colloff (eds) Acarology: Proceedings of the 10th International Congress pp. 586–593. CSIRO Publishing: Melbourne.

[O09] OConnor, B. M. 2009. Cohort Astigmatina. In: Krantz, G. W., & D. E. Walter (eds) A Manual of Acarology 3rd ed. pp. 565–657. Texas Tech University Press.

[P90] Philips, J. R. 1990. Acarina: Astigmata (Acaridida). In: Dindal, D. L. (ed.) Soil Biology Guide pp. 757–778. John Wiley & Sones: New York.

[SH01] Sumangala, K., & M. A. Haq. 2001. Survey of the mite fauna associated with Apis spp. in Kerala, southern India. In: Halliday, R. B., D. E. Walter, H. C. Proctor, R. A. Norton & M. J. Colloff (eds) Acarology: Proceedings of the 10th International Congress pp. 565–568. CSIRO Publishing: Melbourne.

[W01] Wurst, E. 2001. The life cycle of Lemanniella minotauri n. sp. and the erection of the new family Lemanniellidae (Acari: Astigmata). Stuttgarter Beiträge zur Naturkunde Serie A (Biologie) 621: 1–34.

Spinicaudata

Limnadia lenticularis, from Sars (1896).


Belongs within: Diplostraca.

The Spinicaudata are a group of clam shrimps characterised by the formation of clasps on the first and second pair of thoracopods of the males.

<==Spinicaudata
    |  i. s.: Caenestheria dictyon TDC06
    |         Caenestheriella TDC06
    |           |--C. mariae T06
    |           `--C. packardi T06
    |         Eocyzicus TDC06
    |--Leptestheria T06 [Leptestheriidae WZK06]
    |    `--L. compleximanus T06
    |--Cyzicoidea G88
    |    |--Cyzicidae WZK06
    |    |--Glyptoasmussiidae G88
    |    `--Asmussiidae G88
    `--Limnadiidae [Limnadioidea] WZK06
         |--Imnadia [Imnadiidae] WZK06
         |--Limnadopsium WZK06
         |--Metalimnadia [Metalimnadiidae, Paraimnadiidae] WZK06
         |    `--M. serratura [incl. Paraimnadia guayanensis] MD01
         |--Limnadopsis [Limnadopsidae] WZK06
         |    |--L. birchii TDC06
         |    |--L. pilbarensis GMD11
         |    `--L. tatei Spencer & Hall 1896 WZK06
         |--Limnetis D01
         |    |--L. mucronatus D01
         |    `--L. zichyi Daday 1901 D01
         |--Eulimnadia WZK06
         |    |--E. dahli Sars 1896 WZK06
         |    |--E. feriensis Dakin 1914 WZK06
         |    `--E. texana T06
         `--Limnadia WZK06
              |--L. badia Wolf 1911 WZK06
              |--L. lenticularis WZK06
              |--L. sordida King 1855 WZK06
              |--L. stanleyana King 1855 WZK06
              `--L. urukhai Webb & Bell 1979 T06

*Type species of generic name indicated

REFERENCES

[D01] Daday, E. 1901. Édesvizi mikroszkópi állatok [Mikroskopische Süsswasserthiere]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 375–470. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[GMD11] George, A. S., N. L. McKenzie & P. Doughty. 2011. A Biodiversity Survey of the Pilbara Region of Western Australia 2002–2007. Records of the Western Australian Museum Supplement 78 (1).

[G88] Gray, J. 1988. Evolution of the freshwater ecosystem: the fossil record. Palaeogeography, Palaeoclimatology, Palaeoecology 62: 1–214.

[MD01] Martin, J. W., & G. E. Davis. 2001. An updated classification of the Recent Crustacea. Natural History Museum Los Angeles County, Science Series 39: 1–124.

[T06] Timms, B. V. 2006. The large branchiopods (Crustacea: Branchiopoda) of gnammas (rock holes) in Australia. Journal of the Royal Society of Western Australia 89 (4): 163–173.

[TDC06] Timms, B. V., B. Datson & M. Coleman. 2006. The wetlands of the Lake Carey catchment, northeast Goldfields of Western Australia, with special reference to large branchiopods. Journal of the Royal Society of Western Australia 89 (4): 175–183.

[WZK06] Weeks, S. C., M. Zofkova & B. Knott. 2006. Limnadiid clam shrimp distribution in Australia (Crustacea: Branchiopoda: Spinicaudata). Journal of the Royal Society of Western Australia 89 (4): 155–161.