Merizodinae

Merizodus soledadinus, from Lebouvier et al. (2020).

Belongs within: Conjunctae.

The Merizodinae are a group of small ground beetles with a disjunct distribution in southern South America, southern Australasia and northeastern Asia (Roig-Juñent & Cicchino 2001).

Characters (from Roig-Juñent & Cicchino 2001, as Zolini): Maxillary palpomere 3 plurisetose; elytron with apical plica, recurrent stria joined with apical portion of stria 5; tarsomeres dorsally sulcate.

<==Merizodinae [Merizodini, Oopterini, Zolini] R-JC01
    |--Sinozolina R-JC01
    |--Chaltenia Roig-Juñent & Cicchino 2001 [Chalteniina] R-JC01
    |    `--*C. patagonica Roig-Juñent & Cicchino 2001 R-JC01
    `--+--Oopterus Guérin-Ménéville 1841 [incl. Zolus Sharp 1886; Oopterina, Zolina] R-JC01
       |    `--‘Zolus’ helmsi H15
       `--Merizodus [Merizodina] R-JC01
            |--M. angustiocollis R-JC01
            `--M. soledadinus R-JC01

Merizodinae incertae sedis:
  Idacarabus Lea 1910 LB91, LM87
    |--I. cordicollis Moore 1967 LM87
    |--I. longicollis Moore 1978 LM87
    `--I. troglodytes Lea 1910 LM87
  Pterocyrtus Sloane 1920 B02, LM87
    |--*P. globosus Sloane 1920 LM87
    |--P. rubescens Sloane 1920 LM87
    |--P. striatulus Sloane 1920 LM87
    |--P. tasmanicus (Castelnau 1867) [=Drimostoma tasmanica] LM87
    `--P. truncaticollis Sloane 1923 LM87
  Sloaneana Csiki 1933 B02, LM87 (see below for synonymy)
    |--S. lamingtonensis Baehr 2002 B02
    |--S. similis Baehr 2002 B02
    `--S. tasmaniae (Sloane 1915) [=Brachydema tasmaniae; incl. B. victoriae Sloane 1915] LM87

Sloaneana Csiki 1933 B02, LM87 [=Brachydema Sloane 1915 non Fairmaire 1881 LM87, Sloanella Csiki 1928 non Jeannel 1927 LM87]

*Type species of generic name indicated

REFERENCES

[B02] Baehr, M. 2002. Two new species of Sloaneana Csiki from southern Queensland (Coleoptera, Carabidae, Merizodinae). Memoirs of the Queensland Museum 48: 9–15.

[H15] Hedley, C. 1915. Presidential address. Journal and Proceedings of the Royal Society of New South Wales 49 (1): 1–77, pls 1–7.

[LB91] Lawrence, J. F., & E. B. Britton. 1991. Coleoptera (beetles). In: CSIRO. The Insects of Australia: A textbook for students and research workers 2^nd ed. vol. 2 pp. 543–683. Melbourne University Press: Carlton (Victoria).

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

[R-JC01] Roig-Juñent, S., & A. C. Cicchino. 2001. Chaltenia patagonica, new genus and species belonging to Chalteniina, a new subtribe of Zolini (Coleoptera: Carabidae). Canadian Entomologist 133: 651–670.

Bembidiini

Notaphus varius, copyright Udo Schmidt.

Belongs within: Conjunctae.
Contains: Elaphropus, Pericompsus, Tachys, Philipis, Bembidion, Peryphus.

The Bembidiini are a group of carabid beetles with reduced terminal segments on the labial and maxillary palps, and two unequally dilated segments in the tarsi of males. The majority of bembidiins are surface-dwelling hygrophiles though some are xerophilous and others are endogeous (Ball & Bousquet 2001). Members of the genus Micratopus have truncate elytra and a single pair of supraorbital setae whereas other Bembidiini possess two pairs of supraorbital setae and the elytra are not truncate (Bell 1990). Members of the subtribe Tachyina are characterised by an apicolateral notch on the protibiae, and usually an apical recurrent groove on the elytra.

<==Bembidiini [Bembidiinae]
    |--Gouleta [Xystosomina] R-JC01
    |    `--G. cayennensis R-JC01
    |--Tachyina R-JC01
    |    |--Pericompsus R-JC01
    |    |--Tachys BB01
    |    `--Philipis B02
    `--Bembidiina R-JC01
         |--Bembidion LM87
         |--Notaphus Dejean 1821 R-JC01, BB01
         |    |--N. varius (Olivier 1795) LM87 (see below for synonymy)
         |    |--N. brullei (Gemminger & Harold 1868) (see below for synonymy) LM87
         |    `--N. convergens [=Austronotaphus convergens] R-JC01
         `--Peryphus R-JC01

Bembidiini incertae sedis:
  Lymnastis Motschulsky 1862 B08
    |--*L. indicus (Motschulsky 1851) [=Lymnaeum indicum] B08
    |--L. atricapillus B08
    |--L. brooksi Baehr 2008 B08
    |--L. coomani B08
    |--L. decorsei B55
    |--L. inops B08
    |--L. macrops B55
    |--L. niloticus B55
    |--L. pilosus Bates 1892 [incl. Tachys setiger Sloane 1903] LM87
    `--L. pullulus B08
  Brachytachys Basilewsky 1953 B55
    |--B. curtulus Basilewsky 1953 B55
    `--B. villiersi Basilewsky 1953 B55
  Sphaerotachys B55
    |--S. fumigatus B55
    `--S. haemorrhoidalis B55
  Micratopus Casey 1914 [Micratopini] BB01
    `--M. aenescens (LeConte 1848) BB01
  Anillinus Casey 1918 [incl. Troglanillus Jeannel 1963] BB01
    `--A. valentinei (Jeannel 1963) BB01
  Asaphidion des Gozis 1886 BB01
    |--A. cyanicorne F05
    |    |--A. c. cyanicorne F05
    |    `--A. c. atlanticum Ortuño & Toribio 2005 F05
    |--A. flavipes (Linnaeus 1761) [=Cicindela flavipes] B55
    `--A. stierlini (Heyden 1880) VPB11
  Mioptachys Bates 1882 [incl. Tachymenis Motschulsky 1862 (preoc.)] BB01
    `--M. flavicauda (Say 1823) B14
  Tachyta Kirby 1837 B90, LM87
    |  i. s.: T. subvirens B55
    `--T. (Tachyta) LM87
         |--T. (T.) nana LM87
         |    |--T. n. nana LM87
         |    `--T. n. inornatus (Say 1823) [incl. *T. picipes Kirby 1837] LM87
         `--T. (T.) brunnipennis (Macleay 1871) [=Bembidium brunnipenne] LM87
  Porotachys Netolitzky 1914 BB01
    `--P. bisulcatus (Nicolai 1822) BB01
  Illaphanus Macleay 1865 LB91, LM87
    |--*I. stephensii Macleay 1865 LM87
    `--I. macleayi Lea 1906 LM87
  Phrypeus Casey 1924 BB01
    `--P. rickseckeri (Hayward 1897) BB01
  Elaphropus BB01
  Polyderis Motschulsky 1862 [incl. Microtachys Casey 1918] BB01
    |--*P. brevicornis (Chaudoir 1846) [=Tachys brevicornis] LM87
    |--P. antigua Erwin 1971 BB01
    |--P. laevis (Say 1823) BB01
    `--P. leleupi Basilewsky 1953 B55
  Paratachys Casey 1918 BB01
    `--*P. austinicus Casey 1918 LM87
  Micranillodes Jeannel 1963 BB01
    `--M. depressus Jeannel 1963 BB01
  Anillodes Jeannel 1963 BB01
  Serranillus Barr 1995 BB01
    `--S. jeanneli Barr 1995 BB01
  Anillaspis Casey 1918 BB01
  Horologion Valentine 1932 BB01
    `--H. speokoites Valentine 1932 BB01

Notaphus brullei (Gemminger & Harold 1868) [=Bembicidium brullei, Bembidion (Notaphus) brullei, Bembidion variegatum Brullé 1838 non Say 1823] LM87

Notaphus varius (Olivier 1795) LM87 [=Carabus varius LM87, Bembidion (*Notaphus) varium LM87, Elaphrus (Bembidion) varius G20]

*Type species of generic name indicated

REFERENCES

[B02] Baehr, M. 2002. New species and additional records of the tachyine Philipis Erwin (Coleoptera: Carabidae: Bembidiinae). Memoirs of the Queensland Museum 48: 1–7.

[B08] Baehr, M. 2008. A new cavernicolous and eyeless species of the genus Lymnastis from Christmas Island, Australia (Insecta: Coleoptera: Carabidae: Bembidiinae). Records of the Western Australian Museum 24 (2): 199–203.

[BB01] Ball, G. E., & Y. Bousquet. 2001. Carabidae Latreille, 1810. In: Arnett, R. H., Jr & M. C. Thomas (eds) American Beetles vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia pp. 32–132. CRC Press: Boca Raton.

[B55] Basilewsky, P. 1955. Carabidae (Coleoptera) de l’Angola (première partie). Diamang Publicações Culturais 27: 91–137.

[B90] Bell, R. T. 1990. Insecta: Coleoptera Carabidae adults and larvae. In: Dindal, D. L. (ed.) Soil Biology Guide pp. 1053–1092. John Wiley & Sones: New York.

[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature's gems. Ivy Press: Lewes (United Kingdom).

[F05] Fernández, J. 2005. Noticia de nuevos táxones para la ciencia en el ámbito Íbero-Balear y Macaronésico. Nuevos táxones animales descritos en la península Ibérica y Macaronesia desde 1994 (IX). Graellsia 61 (2): 261–282.

[G20] Goldfuss, G. A. 1820. Handbuch der Naturgeschichte vol. 3. Handbuch der Zoologie pt 1. Johann Leonhard Schrag: Nürnberg.

[LB91] Lawrence, J. F., & E. B. Britton. 1991. Coleoptera (beetles). In: CSIRO. The Insects of Australia: A textbook for students and research workers 2^nd ed. vol. 2 pp. 543–683. Melbourne University Press: Carlton (Victoria).

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

[R-JC01] Roig-Juñent, S., & A. C. Cicchino. 2001. Chaltenia patagonica, new genus and species belonging to Chalteniina, a new subtribe of Zolini (Coleoptera: Carabidae). Canadian Entomologist 133: 651–670.

[VPB11] Varet, M., J. Pétillon & F. Burel. 2011. Comparative responses of spider and carabid beetle assemblages along an urban-rural boundary gradient. Journal of Arachnology 39 (2): 236–243.

Last updated: 20 December 2020.

Elaphrinae

Elaphrus lapponicus, from here.

Belongs within: Geadephaga.

The Elaphrinae are a group of carabid beetles characterised by the presence of a setigerous puncture in each scrobe, and two setigerous punctures above each eye (Ball & Bousquet 2001).

Characters (from Ball & Bousquet 2001): Size moderate to large (total length ca. 6-18 mm); head capsule ventrally without antennal grooves; antennal insertions laterad, each between base of mandible and anterior margin of eye; antenna filiform, antennomeres 2-4 not nodose, 2-6 without unusually long setae, eleven antennomeres; labrum with six setae; mandible with seta in scrobe; terminal palpomeres about cylindrical; pterothorax with mesonotal scutellum dorsally visible, not concealed beneath elytra; front coxal cavities closed by insertion of tip of proepimeron each side in cavity of lateral projection of prosternum, middle and hind coxal cavities disjunct-confluent; elytra extended to apex of abdominal sternum VIII, with apical margins subsinuate, striate or not dorsally; metathoracic wings large; front tibia anisochaete, antenna cleaner sulcate, grade B; abdomen with six (II-VII) pregenital sterna normally exposed, sterna III and IV separated by suture; male genitalia with parameres similar in form, and both moderately long, setose preapically and apically; ovipositor with gonocoxae rotated 90° in repose, without ramus, gonocoxa with two articles, gonocoxite 2 without accessory lobe; and reproductive tract without spermatheca 1, with spermatheca 2; secretions of pygidial glands higher saturated acids, discharge by oozing.

<==Elaphrinae [Elaphrini, Elaphritae] B14
    |--Diacheila Motschulsky 1844 BB01
    |--Nebriosoma Castelnau 1867 LM87
    |    `--*N. fallax Castelnau 1867 LM87
    |--Calyptogonia Sloane 1920 LM87
    |    `--*C. atra Sloane 1920 [=C. ater] LM87
    |--Decogmus Sloane 1915 LM87
    |    `--*D. chalybeus Sloane 1915 LM87
    |--Belthisa Bonelli 1810 BB01
    |    |--B. catenaria Brown 1944 BB01
    |    `--B. multipunctata BB01
    |         |--B. m. multipunctata BB01
    |         `--B. m. aurata Fischer von Waldheim 1828 BB01
    `--Elaphrus Fabricius 1775 BB01
         |  i. s.: E. aquaticus L02
         |         E. californicus OM08
         |         E. cupreus C01
         |         E. riparius C01
         |         E. sugai Nakane 1987 I92
         |         E. uliginosus C01
         |         E. viridis Horn 1878 B14
         |--E. (Arctelaphrus Semenov 1926) BB01
         |    `--E. (A.) lapponicus Gyllenhal 1810 BB01
         |--E. (Elaphroterus Semenov 1895) BB01
         `--E. (Neoelaphrus Hatch 1951) BB01

*Type species of generic name indicated

REFERENCES

[BB01] Ball, G. E., & Y. Bousquet. 2001. Carabidae Latreille, 1810. In: Arnett, R. H., Jr & M. C. Thomas (eds) American Beetles vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia pp. 32–132. CRC Press: Boca Raton.

[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature's gems. Ivy Press: Lewes (United Kingdom).

[C01] Csiki, E. 1901. Bogarak [Coleopteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 75–120. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[I92] Iwahashi, J. (ed.) 1992. Reddo Deeta Animaruzu: a pictorial of Japanese fauna facing extinction. JICC: Tokyo.

[L02] Latreille, P. A. 1802. Histoire Naturelle, générale et particulière des crustacés et des insectes vol. 3. Familles naturelles des genres. F. Dufart: Paris.

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

[OM08] Ober, K. A., & D. R. Maddison. 2008. Phylogenetic relationships of tribes within Harpalinae (Coleoptera: Carabidae) as inferred from 28S ribosomal DNA and the wingless gene. Journal of Insect Science 8: 63.

Megacephala (Pseudotetracha)

Female Megacephala blackburni, from Mal et al. (2012).

Belongs within: Cicindelinae.

The subgenus Pseudotetracha of the genus Megacephala is a group of bright metallic-coloured tiger beetles found on Australian salt flats.

<==Megacephala (Pseudotetracha Fleutiaux 1894) G07
    |--M. (*P.) cylindrica Macleay 1863 [incl. M. frenchi Sloane 1893] LM87
    |--M. (P.) australis (Chaudoir 1865) G07 [=Tetracha australis LM87; incl. T. waterhousii Castelnau 1867 LM87]
    |--M. (P.) blackburni (Fleutiaux 1895) G07 [=Tetracha blackburni LM87]
    |--M. (P.) castelnaui Sloane 1906 G07
    |--M. (P.) cuprascens Sumlin 1997 G07
    |--M. (P.) greyanus (Sloane 1901) G07, LM87 [=Tetracha greyanus LM87]
    |--M. (P.) helmsi (Blackburn 1892) G07 [=Tetracha helmsi LM87]
    |--M. (P.) ion Sumlin 1997 G07
    |--M. (P.) marginicollis Sloane 1906 G07
    |--M. (P.) mendacia Sumlin 1997 G07
    |--M. (P.) oleadorsa Sumlin 1992 G07
    `--M. (P.) pulchra (Brown 1869) G07 [=Tetracha pulchra LM87]
         |--M. p. pulchra LM87
         `--M. p. pseudoblackburni Horn 1923 LM87

*Type species of generic name indicated

REFERENCES

[G07] Golding, M. R. 2007. A Pictorial Field Guide to the Beetles of Australia. Part 2: Cicindelidae. Ocean Publishing.

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

Cicindela (Rivacindela)

Rivacindela vannidekiana, copyright Mark Golding.

Belongs within: Cicindelinae.

The subgenus Rivacindela of the genus Cicindela is a group of tiger beetles found on Australian salt flats, characterised by the presence of two or three teeth on the labrum and usually with extensive patches of white on the elytra.

<==Cicindela (Rivacindela Brouerius van Nidek 1973) LM87
    |--C. (*R.) blackburni Sloane 1906 LM87
    |--‘Rivacindela’ aurifodina Sumlin 1997 G07
    |--‘Rivacindela’ avita Sumlin 1997 G07
    |--C. (R.) browni Sloane 1913 LM87
    |--‘Rivacindela’ eburneola Sumlin 1997 G07
    |--C. (R.) gairdneri Freitag 1979 LM87
    |--C. (R.) igneicollis Bates 1874 LM87
    |--‘Rivacindela’ igneicolloides Sumlin 1992 G07
    |--‘Rivacindela’ pseudotrepida Sumlin 1997 G07
    |--C. (R.) saetigera Horn 1893 [incl. C. jungi Blackburn 1901] LM87
    |--‘Rivacindela’ salicursoria Sumlin 1987 G07
    |--‘Rivacindela’ trepida Sumlin 1996 G07
    |--‘Rivacindela’ vannidekiana Sumin 1988 G07
    `--‘Rivacindela’ velox Sumlin 1987 G07

*Type species of generic name indicated

REFERENCES

[G07] Golding, M. R. 2007. A Pictorial Field Guide to the Beetles of Australia. Part 2: Cicindelidae. Ocean Publishing.

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

Cicindela (Myriochile)

Cicindela semicincta, copyright Victor W. Fazio III.

Belongs within: Cicindelinae.

The subgenus Myriochile of the genus Cicindela is a group of tiger beetles with completely metallic-coloured legs bearing hooked setae along the posterior margin, and moderately abundant appressed setae on the venter of the body.

<==Cicindela (Myriochile Motschulsky 1861) [=Myriochila (l. c.)] LM87
    |--‘Myriochile’ melancholica (Fabricius 1798) B55, LM87 (see below for synonymy)
    |--‘Myriochile’ angusticollis B55
    |--‘Myriochile’ fastidiosa B55
    |--C. (M.) mastersi Castelnau 1867 LM87
    |    |--C. m. mastersi [incl. C. curvicollis Sloane 1905, C. pseudoplebeja Mandl 1960] LM87
    |    |--C. m. catoptriola Horn 1901 LM87
    |    `--C. m. plebeia Sloane 1905 LM87
    |--‘Myriochile’ octoguttata B55
    |--C. (M.) parasemicincta Freitag 1979 LM87
    |--C. (M.) semicincta Brullé 1834 (see below for synonymy) LM87
    `--‘Myriochile’ vicina B55

Cicindela (Myriochile) semicincta Brullé 1834 [incl. C. circumcincta Castelnau 1867, C. hemicycla Montrouzier 1855, C. interrupta Fabricius 18701 non Fabricius 1775, C. menestriesi Gistl 1837] LM87

‘Myriochile’ melancholica (Fabricius 1798) B55, LM87 [=Megacephala melancholica LM87; incl. Cicindela aegyptiaca Klug 1832 LM87, *Myriochile aegyptiaca LM87]

*Type species of generic name indicated

REFERENCES

[B55] Basilewsky, P. 1955. Carabidae (Coleoptera) de l’Angola (première partie). Diamang Publicações Culturais 27: 91–137.

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

Broscinae

Apotomus rufus, copyright elleelle.

Belongs within: Conjunctae.
Contains: Broscini.

The Broscinae are a group of ground beetles characterised by a pedunculate connection between the pro- and mesothorax, and a single supraorbital seta. Members of the genus Apotomus, which have elongate palps and an unmargined prothorax, are found on mudflats adjacent to rivers (Bouchard 2014).

<==Broscinae [Broschidae, Broscitae]
    |--+--Broscini R-JC01
    |  `--Melaenitae [Melaeninae] R-JC01
    |       |--Melaenus [Melaenini] R-JC01
    |       |    `--M. piger R-JC01
    |       `--Cymbionotum B14 [Cymbionotini R-JC01]
    |            `--C. (Procoscinia) B14
    |                 |--C. (P.) fernandezi Ball & Shpeley 2005 B14
    |                 `--C. (P.) negrei B14
    `--Apotomus Illiger 1807 R-JC01, LM87 [Apotominae, Apotomini, Apotomitae]
         |--*A. rufus (Rossi 1790) [=Scarites rufus] LM87
         |--A. australis Castelnau 1867 LM87
         |--A. fuscus B89
         |--A. hirsutulus R-JC01
         |--A. mastersii Macleay 1871 LM87
         |--A. novaehollandiae Castelnau 1867 LM87
         |--A. reichardti Erwin 1980 B14
         `--A. xanthotelus Bates 1874 B89

Broscinae incertae sedis:
  Brithysternum Macleay 1873 LM87
    |--*B. calcaratum Macleay 1873 LM87
    |--B. macleayi Sloane 1910 LM87
    `--B. nodosum Sloane 1910 LM87
  Adotela Castelnau 1867 LM87
    |--A. carenoides Putzeys 1873 LM87
    |--A. concolor Castelnau 1867 LM87
    |--A. esmeralda Castelnau 1867 LM87
    |--A. frenchi Sloane 1890 LM87
    |--A. nigerrima Macleay 1873 LM87
    |--A. striolata Putzeys 1873 LM87
    `--A. viridis (Macleay 1871) [=Promecoderus viridis] LM87
  Parroa Castelnau 1867 LM87
    |--P. apicalis Sloane 1893 LM87
    |--P. atronitens (Sloane 1890) [=Adotela atronitens] LM87
    |--P. australis (Sloane 1890) [=Adotela australis] LM87
    |--P. bicolor Castelnau 1867 LM87
    |--P. carbonaria Castelnau 1867 LM87
    |--P. grandis Castelnau 1867 LM87
    |--P. howittii Castelnau 1867 LM87
    |--P. laevigata Sloane 1893 LM87
    |--P. noctis Sloane 1893 LM87
    `--P. violacea Castelnau 1867 LM87
  Cerotalis Castelnau 1867 LM87
    |--C. amabilis Sloane 1890 LM87
    |--C. brachypleura Sloane 1898 LM87
    |--C. longipes Sloane 1898 LM87
    |--C. majuscula (Putzeys 1868) [=Promecoderus majusculus] LM87
    |--C. semiviolacea Castelnau 1867 LM87
    |--C. substriata Castelnau 1867 LM87
    `--C. versicolor Castelnau 1867 LM87
  Gnathoxys Westwood 1842 LM87
    |--G. barbatus Macleay 1864 LM87
    |--G. cicatricosus Reiche 1842 LM87
    |--G. crassipes Sloane 1898 LM87
    |--G. foveatus Macleay 1866 LM87
    |--G. granularis Westwood 1842 [incl. G. blissii Macleay 1866] LM87
    |--G. humeralis M80
    |--G. insignitus Macleay 1864 LM87
    |--G. irregularis Westwood 1842 LM87
    |--G. macleayi Putzeys 1868 LM87
    |--G. murrumbidgensis Macleay 1865 LM87
    |--G. obscurus Reiche 1842 LM87
    |--G. punctipennis Macleay 1873 LM87
    |--G. submetallicus Macleay 1864 LM87
    |--G. sulcicollis Sloane 1910 LM87
    |--G. tesselatus Macleay 1864 LM87
    `--G. westwoodi Putzeys 1868 LM87
  Chylnus Sloane 1920 [=Lychnus Putzeys 1868 non Matheson 1832] LM87
    `--*C. ater (Putzeys 1868) (see below for synonymy) LM87
  Percolestus Sloane 1892 LM87
    `--*P. blackburni Sloane 1892 LM87
  Percosoma Schaum 1858 LM87
    |--*P. carenoides (White 1846) (see below for synonymy) LM87
    |--P. concolor Sloane 1892 LM87
    |--P. montanum (Castelnau 1867) [=Mecodema montanum] LM87
    |--P. substriatum Moore 1960 LM87
    `--P. sulcipenne Bates 1878 LM87

*Chylnus ater (Putzeys 1868) [=*Lychnus ater; incl. L. strangulatus Bates 1878, L. striatulus Bates 1878] LM87

*Percosoma carenoides (White 1846) [=Broscus carenoides; incl. B. hopei Sturm 1843 (n. n.), Mecodema percoides Castelnau 1867] LM87

*Type species of generic name indicated

REFERENCES

[B89] Bates, H. W. 1889. Viaggio di Leonardo Fea in Birmani e regioni vicine. XVI.—On some Carabidae from Burma collected by Mr. L. Fea. Annali del Museo Civico di Storia Naturale di Genova, Serie 2^a, 7: 100–111.

[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature's gems. Ivy Press: Lewes (United Kingdom).

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

[M80] Matthews, E. G. 1980. A Guide to the Genera of Beetles of South Australia. Part 1. Archostemata and Adephaga. South Australian Museum: Australia.

[R-JC01] Roig-Juñent, S., & A. C. Cicchino. 2001. Chaltenia patagonica, new genus and species belonging to Chalteniina, a new subtribe of Zolini (Coleoptera: Carabidae). Canadian Entomologist 133: 651–670.

Last updated: 3 November 2018.

Promecoderus

Promecoderus viridiaeneus, copyright Lynne Forster.

Belongs within: Broscini.

Promecoderus is an Australian genus of broscin ground beetles characterised by the presence of a median tooth on the mentum.

<==Promecoderus Dejean 1829 [incl. Acallistus Sharp 1886, Anheterus Putzeys 1868, Cnemacanthus Gray 1832] LM87
    |--*P. brunnicornis Dejean 1829 [incl. P. degener Guérin-Méneville 1841, P. morosus Putzeys 1868] LM87
    |--P. albaniensis Castelnau 1867 LM87
    |--P. ambiguus Sloane 1892 LM87
    |--P. anguliceps Sloane 1898 LM87
    |--P. bassii Castelnau 1867 LM87
    |--P. blackburni Sloane 1890 LM87
    |--P. castelnaui Sloane 1892 LM87
    |--P. clivinoides Guérin-Méneville 1841 [incl. P. minutus Castelnau 1867] LM87
    |--‘Cnemacanthus’ coerulescens B49
    |--P. comes Sloane 1890 LM87
    |--P. concolor Germar 1848 (see below for synonymy) LM87
    |--P. cordicollis Sloane 1908 LM87
    |--P. cuprescens Sloane 1920 LM87
    |--P. curvipes Sloane 1920 LM87
    |--P. distinctus Sloane 1890 LM87
    |--P. dorsalis Macleay 1873 LM87
    |--P. dyschirioides Guérin-Méneville 1841 LM87
    |--P. elegans Castelnau 1867 LM87
    |--P. gibbosus (Gray 1832) [=*Cnemacanthus gibbosus] LM87
    |--P. gracilis Germar 1848 [=*Anheterus gracilis; incl. P. parvulus Macleay 1873] LM87
    |--P. hunteriensis Macleay 1873 LM87
    |--P. inornatus Macleay 1873 [incl. P. puncticollis Macleay 1873] LM87
    |--P. insignis Sloane 1890 LM87
    |--P. intermedius Sloane 1898 LM87
    |--P. interruptus Macleay 1873 LM87
    |--P. leai Sloane 1898 LM87
    |--P. longus Sloane 1920 LM87
    |--P. lucidicollis Castelnau 1867 [incl. P. politus Sloane 1890] LM87
    |--P. lottini Brullé 1834 LM87
    |--P. maritimus Castelnau 1867 LM87
    |--P. mastersii Macleay 1873 LM87
    |--P. modestus Castelnau 1867 LM87
    |--P. neglectus Castelnau 1867 [incl. P. puella Putzeys 1868] LM87
    |--P. nigellus Sloane 1890 LM87
    |--P. nigricornis Castelnau 1867 LM87
    |--P. oblongus Castelnau 1867 LM87
    |--P. olivaceus Macleay 1873 LM87
    |--P. ovicollis Castelnau 1867 LM87
    |--P. ovipennis Sloane 1898 LM87
    |--P. pacificus Sloane 1890 LM87
    |--P. plebius Sloane 1920 LM87
    |--P. pygmaeus Castelnau 1867 LM87
    |--P. riverinae Macleay 1873 LM87
    |--P. scauroides Castelnau 1867 LM87
    |--P. semistriatus Castelnau 1867 LM87
    |--P. sloanei Blackburn 1901 LM87
    |--P. striatopunctatus Castelnau 1867 LM87
    |--P. subdepressus Guérin-Méneville 1841 LM87
    |--P. tasmanicus Castelnau 1867 [incl. *Acallistus simplex Sharp 1886] LM87
    |--P. viridiaeneus Sloane 1915 LM87
    `--P. wilcoxii Castelnau 1867 LM87

Promecoderus concolor Germar 1848 [incl. P. anthracinus Macleay 1873, P. howittii Castelnau 1867, P. lucidus Putzeys 1868, P. concolor namoyensis Sloane 1892, P. suturalis Castelnau 1867] LM87

*Type species of generic name indicated

REFERENCES

[B49] Britton, E. B. 1949. The Carabidae (Coleoptera) of New Zealand. Part III. A revision of the tribe Broscini. Transactions and Proceedings of the Royal Society of New Zealand 77 (4): 533–581.

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

Broscini

Broscus cephalotes, copyright Udo Schmidt.

Belongs within: Broscinae.
Contains: Mecodema, Promecoderus.

The Broscini are a group of large ground beetles with a pedunculate thorax, most diverse in the Australasian region (Britton 1949). The genus Broscus, species of which are relatively large and have two pairs of setae on the pronotum and eight or nine setae laterally on the elytra, is found in the Palaearctic region, though the species B. cephalotes has also been introduced to eastern Canada (Ball & Bousquet 2001).

Characters (from Britton 1949): Size large; colour dark brown or black, occasionally slightly aeneous; mandibles usually with scrobal seta; labrum with angles rounded, with six setae on front margin and row of small setae arising just beneath anterolateral margins; prothorax connected to mesothorax by peduncle formed by mesothoracic projection bearing scutellum; pronotum without setiferous puncture at posterior angle; border base of elytra not complete, elytra fused along suture, hind wings vestigial; anterior coxal cavities with single perforation; mesepimera not reaching border of middle coxal cavities; anterior tarsi not widened in male, without modified setae ventrally; male genitalia with parameres prolonged into long styles at apices, right-hand style bearing fringe of long setae, base of aedeagus closed on dorsal side so orifice is circular.

<==Broscini [Cnemacanthides] B49
    |--Axonya [Axonyina] R-JC01
    |    `--A. championi R-JC01
    `--+--Nothobroscus [Nothobroscina] R-JC01
       |    `--N. chilensis R-JC01
       `--Broscina R-JC01
            |--Zacotus LeConte 1869 BB01
            |    `--Z. matthewsii LeConte 1869 BB01
            |--Miscodera Eschscholtz 1830 BB01
            |    `--M. arctica (Paykull 1800) BB01
            |--Broscodera Lindroth 1961 BB01
            |    |--B. (Broscodera) insignis (Mannerheim 1852) BB01
            |    `--B. (Sinobrosculus Deuve 1990) BB01
            `--Broscus Panzer 1813 B49
                 |--B. cephalotes (Linnaeus 1758) B14
                 |--B. insularis B14
                 |--B. laevigatus G89
                 |--B. nobilis B14
                 |--B. politus B14
                 `--B. punctatus G89

Broscini incertae sedis:
  Rawlinsius R-JC01
  Broscodes R-JC01
  Diglymma Sharp 1886 [incl. Snofru Broun 1908] B49
    |--*D. clivinoides (Castelnau 1867) (see below for synonymy) B49
    |--D. castigatum Broun 1909 B49
    |--D. marginale Broun 1914 B49
    `--D. obtusum (Broun 1893) (see below for synonymy) B49
  Oregus Putzeys 1868 B49
    |--*O. aereus (White 1846) [=Promecoderus aereus] B49
    `--O. inaequalis (Castelnau 1867) [=Mecodema inaequale] B49
  Mecodema B49
  Metaglymma Bates 1867 [incl. Maoria Castelnau 1867] B49
    |--*M. monilifer Bates 1867 [incl. M. minor Broun 1905, M. rugipenne Broun 1893] B49
    |--M. aberrans Putzeys 1868 (see below for synonymy) B49
    `--M. tibiale (Castelnau 1867) (see below for synonymy) B49
  Brullea Castelnau 1867 B49
    `--*B. antarctica Castelnau 1867 B49
  Promecoderus B49
  Eurylychnus Bates 1891 B49, LM87
    |--E. dyschirioides (Castelnau 1867) LM87 [=Maoria dyschirioides B49; incl. *E. olliffi Bates 1891 LM87]
    |--E. blagravei (Castelnau 1867) [=Mecodema blagravei; incl. Percosoma mastersi Rainbow 1899] LM87
    |--E. cylindricus Sloane 1916 LM87
    |--E. femoralis Sloane 1915 LM87
    |--E. kershawi Sloane 1915 LM87
    |--E. ovipennis Sloane 1915 LM87
    |--E. regularis Sloane 1911 LM87
    `--E. victoriae Sloane 1892 LM87

*Diglymma clivinoides (Castelnau 1867) [=Maoria clivinoides; incl. D. basale Broun 1917, D. dubium Sharp 1886, D. ovipenne Sharp 1886] B49

Diglymma obtusum (Broun 1893) [=Metaglymma obtusum; incl. Snofru aemulator Broun 1908, Diglymma nigripes Broun 1893, D. punctipenne Broun 1893, D. tarsale Broun 1908, D. thoracicum Broun 1917] B49

Metaglymma aberrans Putzeys 1868 [incl. M. rugiceps Broun 1903, M. tersatum Broun 1893, M. thoracicum Broun 1893] B49

Metaglymma tibiale (Castelnau 1867) [=*Maoria tibialis; incl. Mecodema asperum Broun 1893, Mec. calcaratum Broun 1903, Mec. junctum Broun 1893, Mec. rufipes Broun 1886] B49

*Type species of generic name indicated

REFERENCES

[BB01] Ball, G. E., & Y. Bousquet. 2001. Carabidae Latreille, 1810. In: Arnett, R. H., Jr & M. C. Thomas (eds) American Beetles vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia pp. 32–132. CRC Press: Boca Raton.

[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature's gems. Ivy Press: Lewes (United Kingdom).

[B49] Britton, E. B. 1949. The Carabidae (Coleoptera) of New Zealand. Part III. A revision of the tribe Broscini. Transactions and Proceedings of the Royal Society of New Zealand 77 (4): 533–581.

[G89] Gestro, R. 1889. Viaggio ab Assab nel Mar Rosso dei signori G. Doria ed O. Beccari con il R. Avviso «Esploratore» dal 16 Novembre 1879 al 26 Febbraio 1880.—IV. Coleotteri. Annali del Museo Civico di Storia Naturale di Genova, Serie 2^a, 7: 5–72.

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

[R-JC01] Roig-Juñent, S., & A. C. Cicchino. 2001. Chaltenia patagonica, new genus and species belonging to Chalteniina, a new subtribe of Zolini (Coleoptera: Carabidae). Canadian Entomologist 133: 651–670.

Last updated: 3 November 2018.

Mecyclothorax

Mecyclothorax punctipennis, from Liebherr (2012).

Belongs within: Conjunctae.

Mecyclothorax is a widespread genus of ground beetles found in insular south-east Asia and Oceania, characterised by filiform antennae and elytral epipleura that are twisted near the apex.

<==Mecyclothorax Sharp 1903 [incl. Cyclothorax Macleay 1871 non Frauenfeld 1867; Mecyclothoracini] LM87
    |--*M. montivagus (Blackburn 1878) [=Cyclothorax montivagus] LM87
    |--M. ambiguus (Erichson 1842) [=Anchomenus ambiguus] LM87
    |    |--M. a. ambiguus M83
    |    `--M. a. rotundicollis M83
    |--M. blackburni (Sloane 1898) [=Cyclothorax blackburni] LM87
    |--M. cordicollis (Sloane 1900) [=Cyclothorax cordicollis] LM87
    |--M. curtus (Sloane 1895) [=Cyclothorax curtus] LM87
    |--M. eyrensis (Blackburn 1892) [=Cyclothorax eyrensis] LM87
    |--M. fortis (Blackburn 1889) [=Cyclothorax fortis] LM87
    |--M. lateralis (Castelnau 1867) (see below for synonymy) LM87
    |--M. lewisensis Moore 1984 LM87
    |--M. lophoides (Chaudoir 1854) [=Anchomenus lophoides] LM87
    |--M. minutus (Castelnau 1867) [=Phorticosomus minutus, Forticosomus minutus] LM87
    |--M. monteithi Moore 1985 LM87
    |--M. ovalis Sloane 1915 LM87
    |--M. peryphoides (Blackburn 1889) [=Cyclothorax peryphoides] LM87
    |--M. punctatus (Sloane 1895) [=Cyclothorax punctatus] LM87
    |--M. punctipennis (Macleay 1871) [=*Cyclothorax punctipennis; incl. C. obsoletus Blackburn 1889] LM87
    |--M. storeyi Moore 1984 LM87
    `--M. vulcans OM08

Mecyclothorax lateralis (Castelnau 1867) [=Phorticosomus lateralis, Forticosomus lateralis; incl. Cyclothorax cinctipennis Blackburn 1889] LM87

*Type species of generic name indicated

REFERENCES

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

[M83] Martin, N. A. 1983. Miscellaneous observations on a pasture fauna: an annotated species list. DSIR Entomology Division Report 3: 1–98.

[OM08] Ober, K. A., & D. R. Maddison. 2008. Phylogenetic relationships of tribes within Harpalinae (Coleoptera: Carabidae) as inferred from 28S ribosomal DNA and the wingless gene. Journal of Insect Science 8: 63.

Gnathaphanus

Male Gnathaphanus melbournensis, photographed by S. E. Thorpe.

Belongs within: Harpalitae.

Gnathaphanus is a genus of ground beetles found in Asia and Australasia, characterised by conspicuous dorsal punctures on the elytra, and sponge-like soles of adhesive setae on the fore and mid tarsi of males.

Gnathaphanus Macleay 1825 [incl. Mirosarus Bates 1878, Pachauchenius Macleay 1864] LM87
    |--*G. vulneripennis (Macleay 1825) [=Trechus (*Gnathaphanus) vulneripennis] LM87
    |--G. aridus Blackburn 1892 LM87
    |--G. froggatti (Macleay 1888) [=Diaphoromerus froggatti] LM87
    |--G. herbaceus Sloane 1900 LM87
    |--G. latus Sloane 1900 LM87
    |--G. licinoides Hope 1842 [incl. Harpalus alternans Castelnau 1867, Catadromus impressus Montrouzier 1860] LM87
    |--G. melbournensis (Castelnau 1867) (see below for synonymy) LM87
    |--G. multipunctatus (Macleay 1888) (see below for synonymy) LM87
    |--G. philippensis (Chevrolat 1841) [=Amblygnathus philippensis; incl. *Pachauchenius laeviceps Macleay 1864] LM87
    |--G. picipes (Macleay 1864) [=Harpalus picipes; incl. Diaphoromerus sulcatulus Macleay 1888] LM87
    |--G. porcatulus (Macleay 1888) [=Diaphoromerus porcatulus] LM87
    |--G. pulcher (Dejean 1829) [=Harpalus pulcher] LM87
    |    |--G. p. pulcher [incl. Harpalus interstitialis Macleay 1864] LM87
    |    `--G. p. speciosus Schauberger 1935 LM87
    |--G. punctifer (Castelnau 1867) [=Harpalus punctiferus; incl. H. montanus Castelnau 1867] LM87
    |--G. rectangulus Chaudoir 1878 LM87
    |--G. riverinae Sloane 1895 LM87
    |--G. sexpunctatus (Macleay 1888) [=Diaphoromerus sexpunctatus] LM87
    `--G. whitei Sloane 1917 LM87

Gnathaphanus melbournensis (Castelnau 1867) [=Harpalus melbournensis; incl. H. adelaidae Castelnau 1867, H. aeneonitens Macleay 1871, H. angustatus Macleay 1871, H. atroviridis Macleay 1871, H. gayndahensis Macleay 1871, *Mirosarus insularis Bates 1878, H. marginicollis Castelnau 1867, H. paroensis Castelnau 1867, H. planipennis Macleay 1871] LM87

Gnathaphanus multipunctatus (Macleay 1888) [=Diaphoromerus multipunctatus; incl. G. darwini Blackburn 1888, Harpalus impressipennis Castelnau 1867 non Dejean 1829, Dioryche upolensis Csiki 1915] LM87

*Type species of generic name indicated

REFERENCES

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

Lecanomerus

Lecanomerus atriceps, copyright Birgit E. Rhode.

Belongs within: Harpalitae.

Lecanomerus is an Australasian genus of small ground beetles with the penultimate segment of the labial palpi conspicuously bisetose, and the fore and mid tarsi of the males ventrally densely pubescent.

Lecanomerus Chaudoir 1850 (see below for synonymy) LM87
    |--L. verticalis (Erichson 1842) (see below for synonymy) LM87
    |--L. aberrans Macleay 1871 LM87
    |--L. ater (Macleay 1871) [=Trechus ater] LM87
    |--L. atriceps (Macleay 1871) [=Trechus atriceps] LM87
    |--L. australis (Blackburn 1888) [=Thenarotes australis] LM87
    |--L. bicolor (Sloane 1900) [=Thenarotes bicolor] lM87
    |--L. carteri Sloane 1911 LM87
    |--L. concolor (Macleay 1871) [=Trechus concolor] LM87
    |--L. curtus Sloane 1911 LM87
    |--L. discoidalis (Blackburn 1888) (see below for synonymy) LM87
    |--L. limbatus Moore 1967 LM87
    |--L. lindi Blackburn 1888 LM87
    |--L. lucidus Sloane 1917 LM87
    |--L. major Blackburn 1892 LM87
    |--L. niger (Darlington 1956) [=*Odontagonum nigrum] LM87
    |--L. obtusus (Sloane 1920) [=Nemaglossa obtusa] LM87
    |--L. recticollis (Macleay 1888) [=Haplaner recticollis] LM87
    |--L. ruficeps Macleay 1871 LM87
    |--L. speluncarius (Moore 1967) [=Thenarotes speluncarius] LM87
    |--L. striatus Blackburn 1892 LM87
    |--L. tasmanicus (Bates 1878) [=*Thenarotes tasmanicus] LM87
    |--L. vestigialis (Erichson 1842) (see below for synonymy) LM87
    `--L. victoriensis (Blackburn 1891) [=Diaphoromerus victoriensis] LM87

Lecanomerus Chaudoir 1850 [incl. Nemaglossa Sloane 1920 non Solier 1849, Odontagonum Darlington 1956, Thenarotes Bates 1878] LM87

Lecanomerus discoidalis (Blackburn 1888) [=Thenarotes discoidalis; incl. T. discoidalis minor Blackburn 1888, T. discoidalis nigricornis Blackburn 1891] LM87

Lecanomerus verticalis (Erichson 1842) [=Harpalus verticalis; incl. L. flavocinctus Blackburn 1888, *L. insidiosus Chaudoir 1850, L. occidentalis Sloane 1898] LM87

Lecanomerus vestigialis (Erichson 1842) [=Harpalus vestigialis; incl. L. labralis Broun 1914, Acupalpus mastersii Macleay 1871, L. nitidus Blackburn 1891, L. stenopus Broun 1886] LM87

*Type species of generic name indicated

REFERENCES

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

Phorticosomus

Phorticosomus gularis, copyright Lily M. Kumpe.

Belongs within: Harpalitae.

Phorticosomus is an Australian genus of ground beetles, members of which have a truncate labrum, a small tooth on the mentum and simple tarsi in both sexes.

Phorticosomus Schaum 1863 LM87
    |--*P. felix Schaum 1863 [incl. P. brunneus Blackburn 1888] LM87
    |--P. castelnaui Sloane 1915 LM87
    |--P. crassus Sloane 1915 [incl. P. crassus brevipennis Sloane 1915] LM87
    |--P. edelii Castelnau 1867 LM87
    |--P. grandis Castelnau 1867 LM87
    |--P. gularis Sloane 1915 LM87
    |--P. horni Sloane 1896 LM87
    |--P. macleayi Sloane 1915 LM87
    |--P. mucronatus Blackburn 1888 LM87
    |--P. nuytsii Castelnau 1867 [incl. P. calcaratus Blackburn 1903] LM87
    |--P. piceus Sloane 1915 LM87
    |--P. randalli Blackburn 1890 LM87
    |--P. robustus Blackburn 1889 LM87
    |--P. rotundatus Moore 1967 LM87
    |--P. rugiceps Macleay 1871 LM87
    |--P. similis Blackburn 1888 LM87
    `--P. zabroides Sloane 1910 LM87

*Type species of generic name indicated

REFERENCES

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

Hypharpax

Hypharpax australis, copyright Jon Sullivan.

Belongs within: Harpalitae.

Hypharpax is a genus of ground beetles found in Indonesia and Australasia, characterised by the presence of a tooth on the mentum, strongly setose abdominal ventrites and a stout hind tarsus.

Hypharpax Macleay 1825 [incl. Sagraemerus Redtenbacher 1867] LM87
    |--*H. lateralis (Macleay 1825) [=Harpalus (*Hypharpax) lateralis] LM87
    |--H. aereus (Dejean 1829) [=Harpalus aereus; incl. H. boisduvalii Castelnau 1867] LM87
    |--H. assimilis (Macleay 1888) [=Haplaner assimilis] LM87
    |--H. australasiae (Dejean 1829) [=Harpalus australasiae] LM87
    |--H. australis (Dejean 1829) (see below for synonymy) LM87
    |--H. bostockii (Castelnau 1867) [=Harpalus bostockii] LM87
    |--H. convexiusculus (Macleay 1871) [=Harpalus convexiusculus] LM87
    |--H. dentipes Wiedemann 1823 [incl. *Sagraemerus javanus Redtenbacher 1867] LM87
    |--H. deyrollei (Castelnau 1867) [=Harpalus deyrolii, Ha. deyrollei] LM87
    |--H. flavitarsis Chaudoir 1878 LM87
    |--H. flindersii (Castelnau 1867) [=Harpalus flindersii] LM87
    |--H. habitans Sloane 1895 LM87
    |--H. interioris Sloane 1895 LM87
    |--H. kingii (Castelnau 1867) [=Harpalus kingii] LM87
    |--H. kreftii (Castelnau 1867) [=Harpalus kreftii; incl. Hy. opacipennis Macleay 1888, Hy. varus Macleay 1888] LM87
    |--‘Harpalus’ mandibularis Castelnau 1867 [=Diaphoromerus queenslandicus Csiki 1932, Hypharpax queenslandicus] lM87
    |--H. moestus (Dejean 1829) (see below for synonymy) LM87
    |--H. nitens Sloane 1911 LM87
    |--H. obsoletus Blackburn 1892 LM87
    |--H. peronii (Castelnau 1867) (see below for synonymy) LM87
    |--H. puncticollis (Macleay 1888) [=Haplaner puncticollis] LM87
    |--H. ranula (Castelnau 1867) [=Harpalus ranula] LM87
    |--H. rotundipennis Chaudoir 1878 LM87
    |--H. sculpturalis (Castelnau 1867) [=Harpalus sculpturalis] LM87
    |--H. sloanei Blackburn 1891 LM87
    |--H. subsericeus (Macleay 1888) [=Haplaner subsericeus] LM87
    `--H. vilis Blackburn 1891 LM87

Hypharpax australis (Dejean 1829) [=Harpalus australis; incl. Ha. coxii Castelnau 1867, Ha. inornatus Germar 1848, Hypharpax parvus Chaudoir 1878] LM87

Hypharpax moestus (Dejean 1829) [=Harpalus moestus, Diaphoromerus maestus (l. c.); incl. Ha. rana Castelnau 1867, Ha. tasmanicus Castelnau 1867, Ha. yarrae Castelnau 1867] LM87

Hypharpax peronii (Castelnau 1867) [=Harpalus peronii; incl. Hy. inornatus Blackburn 1888 non Ha. inornatus Germar 1848, Hy. latiusculus Chaudoir 1878, Harpalus novaehollandiae Castelnau 1867, Hy. puncticauda Bates 1878] LM87

*Type species of generic name indicated

REFERENCES

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

Stenolophus

Stenolophus quinquepustulatus, from here.

Belongs within: Harpalitae.

Stenolophus is a cosmopolitan genus of ground beetles.

Stenolophus Dejean 1821 LM87
    |  i. s.: S. cyanellus Bates 1889 B89
    |         S. discophorus C01
    |         S. gonidius Bates 1889 B89
    |         S. marginatus B89
    |         S. mixtus C01
    |--‘Carabus’ teutonus Schrank 1781 (see below for synonymy) LM87
    `--S. (Egadroma Motschulsky 1855) LM87
         |--S. (*E.) smaragdulus (Fabricius 1798) [=Carabus smaragdulus] LM87
         |--S. (E.) caeruleus Blackburn 1890 LM87
         |--‘Egadroma’ comptum B55
         |--S. (E.) obscurus (Blackburn 1888) [=Lecanomerus obscurus] LM87
         |--S. (E.) piceus (Guérin-Méneville 1830) (see below for synonymy) LM87
         |--‘Egadroma’ plagiferum [incl. E. plagiferum var. straneoi] B55
         |--S. (E.) quadrimaculatus (Macleay 1888) [=Acupalpus quadrimaculatus; incl. A. bimaculatus Macleay 1888] LM87
         |--‘Egadroma’ quadripustulatum B55
         |    |--E. q. quadripustulatum B55
         |    `--E. q. elegans B55
         |--S. (E.) quinquepustulatus (Wiedemann 1823) [=Badister quinquepustulatus] LM87
         |--‘Egadroma’ relucens B55
         |--S. (E.) robustus Sloane 1907 LM87
         |--‘Egadroma’ scapulare B55
         |--S. (E.) subrobustus (Schauberger 1938) (see below for synonymy) LM87
         |--S. (E.) suturalis Macleay 1888 LM87
         |--‘Egadroma’ terminale B55
         `--‘Egadroma’ vestigialis M80

‘Carabus’ teutonus Schrank 1781 [incl. C. vaporariorum Fabricius 1787 non Linnaeus 1758, *Stenolophus vaporariorum] LM87

Stenolophus (Egadroma) piceus (Guérin-Méneville 1830) [=Acupalpus piceus; incl. Harpalus dingo Castelnau 1867, Stenolophus politus Macleay 1871, S. sexualis Fauvel 1882] LM87

Stenolophus (Egadroma) subrobustus (Schauberger 1938) [=Acupalpus (E.) subrobustus; incl. S. grandiceps Fauvel 1882, Amphibia pallipes Perroud & Montrouzier 1864 non Acupalpus pallipes Dejean 1829, S. smaragdulus Darlington 1968 non Carabus smaragdulus Fabricius 1798] LM87

*Type species of generic name indicated

REFERENCES

[B55] Basilewsky, P. 1955. Carabidae (Coleoptera) de l’Angola (première partie). Diamang Publicações Culturais 27: 91–137.

[B89] Bates, H. W. 1889. Viaggio di Leonardo Fea in Birmani e regioni vicine. XVI.—On some Carabidae from Burma collected by Mr. L. Fea. Annali del Museo Civico di Storia Naturale di Genova, Serie 2^a, 7: 100–111.

[C01] Csiki, E. 1901. Bogarak [Coleopteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 75–120. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

[M80] Matthews, E. G. 1980. A Guide to the Genera of Beetles of South Australia. Part 1. Archostemata and Adephaga. South Australian Museum: Australia.

Notiobia

Notiobia sp., copyright Mike Quinn.

Belongs within: Harpalitae.

Notiobia is a genus of ground beetles known from Africa, Australasia and the Americas.

Notiobia Perty 1830 LM87
    |--*N. (Notiobia) nebrioides Perty 1830 LM87
    `--N. (Anisotarsus Chaudoir 1837) [incl. Diaphoromerus Chaudoir 1843] LM87
         |--N. (*A.) brevicollis (Chaudoir 1837) [=*Anisotarsus brevicollis] LM87
         |--N. (A.) angustulus (Chaudoir 1878) [=Diaphoromerus angustulus] LM87
         |--N. (A.) dampierii (Castelnau 1867) [=Harpalus dampierii; incl. H. versicolor Castelnau 1878] LM87
         |--N. (A.) denisonensis (Castelnau 1867) [=Harpalus denisonensis] LM87
         |--N. (A.) edwardsii (Castelnau 1867) (see below for synonymy) LM87
         |--N. (A.) flavipalpis (Macleay 1864) [=Harpalus flavipalpis] LM87
         |--N. (A.) germari (Castelnau 1867) [=Harpalus germari; incl. H. erichsoni Castelnau 1867] LM87
         |--N. (A.) inaequalipennis (Castelnau 1867) [=Harpalus inaequalipennis] LM87
         |--N. (A.) iridipennis (Chaudoir 1843) (see below for synonymy) LM87
         |--N. (A.) lapeyrousei (Castelnau 1867) [=Harpalus lapeyrousei] LM87
         |--N. (A.) laticollis (Macleay 1888) [=Diaphoromerus laticollis] LM87
         |--N. (A.) lucidicollis (Dejean 1829) [=Harpalus lucidicollis] LM87
         |--N. (A.) melanarius (Dejean 1829) (see below for synonymy) LM87
         |--N. (A.) nigrans (Macleay 1888) [=Diaphoromerus nigrans] LM87
         |--N. (A.) oblongiusculus (Castelnau 1867) [=Harpalus oblongiusculus] LM87
         |--N. (A.) ovatus (Chaudoir 1878) [=Diaphoromerus ovatus] LM87
         |--N. (A.) patrueloides (Castelnau 1867) (see below for synonymy) LM87
         |--N. (A.) perater (Sloane 1920) [=Diaphoromerus perater] LM87
         |--N. (A.) planiusculus (Chaudoir 1878) [=Diaphoromerus planiusculus] LM87
         |--N. (A.) planoimpressus (Castelnau 1867) [=Harpalus planoimpressus; incl. H. femoralis Castelnau 1867] LM87
         |--N. (A.) politus (Macleay 1888) [=Diaphoromerus politus] LM87
         |--N. (A.) quadricollis (Chaudoir 1878) [=Diaphoromerus quadricollis] LM87
         |--N. (A.) rectangulus (Chaudoir 1878) [=Diaphoromerus rectangulus] LM87
         |--N. (A.) rugosipennis (Castelnau 1867) [=Harpalus rugosipennis] LM87
         |--N. (A.) sculptipennis (Castelnau 1867) [=Harpalus sculptipennis] LM87
         |--N. (A.) sericipennis (Macleay 1888) [=Diaphoromerus sericipennis] LM87
         `--N. (A.) viridipennis (Sloane 1920) [=Diaphoromerus viridipennis] LM87

Notiobia (Anisotarsus) edwardsii (Castelnau 1867) [=Harpalus edwardsii; incl. Diaphoromerus edwardsi virescens Sloane 1916] LM87

Notiobia (Anisotarsus) iridipennis (Chaudoir 1843) [=*Diaphoromerus iridipennis; incl. Harpalus infelix Castelnau 1867] LM87

Notiobia (Anisotarsus) melanarius (Dejean 1829) [=Harpalus melanarius; incl. Selenophorus baladicus Fauvel 1862, Ophonus billardieri Montrouzier 1860, Gnathaphanus melanarius subtransversus Chaudoir 1867, Harpalus thouzeti Castelnau 1867, H. wilcoxii Castelnau 1867] LM87

Notiobia (Anisotarsus) patrueloides (Castelnau 1867) [=Harpalus patrueloides; incl. H. amaroides Castelnau 1867, H. illawarrensis Castelnau 1867, H. vandiemenensis Castelnau 1867] LM87

*Type species of generic name indicated

REFERENCES

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

Amblystomus

Amblystomus orpheus, copyright Udo Schmidt.

Belongs within: Harpalitae.

Amblystomus is a genus of ground beetles found in Eurasia, Africa and Australasia, characterised by an asymmetrical emarginate clypeus and labrum, quadrate elytra with indistinct striae, and males with two rows of ventral adhesive setae on the fore tarsi.

Amblystomus Erichson 1837 (see below for synonymy) LM87
    |--*A. mauritanicus (Dejean 1829) [=Acupalpus mauritanicus] LM87
    |--A. anthracinus (Sloane 1898) [=Thenarotidius anthracinus] LM87
    |--A. australis Motschulsky 1864 LM87
    |--A. gagatinus (Macleay 1871) [=Bembidium gagatinum, *Thenarotidius gagatinus] LM87
    |--A. gracilis (Blackburn 1888) [=*Notophilus gracilis] LM87
    |--A. indicus (Nietner 1858) [=*Megaristerus indicus] LM87
    |--A. laetus (Blackburn 1888) [=Notophilus laetus] LM87
    |--A. metallicus (Blackburn 1888) [=Thenarotes metallicus] LM87
    |--A. minutus (Castelnau 1867) [=Amblygnathus minutus] LM87
    |--A. montanus (Blackburn 1891) [=Notophilus montanus] LM87
    |--A. nigrinus Csiki 1931 [=Notophilus niger Blackburn 1888 non A. niger Heer 1838] LM87
    |--A. obliquus (Sloane 1898) [=Notophilus obliquus] LM87
    |--A. orpheus B55
    |--A. ovalis (Sloane 1900) [=*Psilonothus ovalis] LM87
    |--A. palustris (Blackburn 1888) [=Notophilus palustris] LM87
    |--A. parvus (Blackburn 1888) [=Notophilus parvus] LM87
    |--A. quadriguttatus (Motschulsky 1858) (see below for synonymy) LM87
    |--A. sloanei Andrewes 1936 [=A. convexus Sloane 1920 non Trechus convexus Macleay 1825] LM87
    `--A. stenolophoides (Nietner 1858) [=Megaristerus stenolophoides] LM87

Amblystomus Erichson 1837 [incl. Megaristerus Nietner 1858, Notophilus Blackburn 1888, Psilonothus Sloane 1900, Thenarotidius Sloane 1898] LM87

Amblystomus quadriguttatus (Motschulsky 1858) [=Hispalis quadriguttatus; incl. Am. (Megaristerus) guttatus Bates 1873, Acupalpus ornatus Macleay 1888] LM87

*Type species of generic name indicated

REFERENCES

[B55] Basilewsky, P. 1955. Carabidae (Coleoptera) de l’Angola (première partie). Diamang Publicações Culturais 27: 91–137.

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

Leiradira

Leiradira violacea, from Kipling Will.

Belongs within: Pterostichitae.

Leiradira is a genus of ground beetles found in eastern Australia, characterised by a deep body, heavy head with relatively long and acuminate mandibles, geniculate antennae, mentum lacking a tooth, and inner lobe of maxilla lined with long slender teeth.

Leiradira Castelnau 1867 LM87
    |--L. (Leiradira) LM87
    |    |--*L. (L.) auricollis Castelnau 1867 (see below for synonymy) LM87
    |    |--L. (L.) alternans Darlington 1953 LM87
    |    |--L. (L.) blandula Tschitschérine 1898 LM87
    |    |--L. (L.) latreillei Castelnau 1867 LM87
    |    |    |--L. l. latreillei [incl. L. liopleura Chaudoir 1868] LM87
    |    |    `--L. l. chaudoiri Straneo 1941 LM87
    |    `--L. (L.) puella Tschitschérine 1898 LM87
    |--L. (Metadira Darlington 1961) LM87
    |    |--L. (*M.) aurifer Darlington 1961 LM87
    |    |--L. (M.) alticola Darlington 1961 LM87
    |    |--L. (M.) opacistriatis (Sloane 1902) [=Notonomus opacistriatis] LM87
    |    `--L. (M.) soror Darlington 1961 LM87
    `--L. (Stomimorphus Straneo 1953) LM87
         |--L. (*S.) violacea (Straneo 1953) [=*Stomimorphus violaceus] LM87
         |--L. (S.) jacobi Darlington 1961 LM87
         `--L. (S.) tenuis Darlington 1961 LM87

*Leiradira (Leiradira) auricollis Castelnau 1867 [incl. L. cupreicollis Tschitschérine 1897, L. purpurascens Tschitschérine 1897] LM87

*Type species of generic name indicated

REFERENCES

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

Darodilia

Darodilia sp., from Will (2011).

Belongs within: Pterostichitae.

Darodilia is an Australian genus of ground beetles with a longitudinally strigose proepisternum and transverse sulci on the abdominal ventrites.

Characters (from Will 2011): Head with clypeo-ocular sulci absent or present (present in some Australian species); mentum very shallowly emarginate, sides divergent, paramedial pits large, deeply impressed; median tooth simple, rounded; paraglossae small, without elongate setae at apex; ligular sclerite with two seta on apical margin; maxillary palpifer with one basal seta; antennae filiform, with three basal segments glabrous. Pronotum orbiculate or cordate, two marginal setae; pro-, meso- and metasterna glabrous; proepisternum smooth or with deep longitudinal strigae; elytra free, border at base, striae 1-4, 8-9 impressed, 5-7 absent or scarcely marked, number of impressed striae in Australian species variable, very short tenth stria at level of plica, apicolateral plica large and visible, parascutellar stria impressed and continuous with stria 1, angular base of stria 1 absent, parascutellar punctures at base of stria 2, no discal punctures, intervals flat or scarcely convex; hind wing full; anterior tarsi of male with three basal segments expanded, ventrally squamous, all tarsi dorsally glabrous. Abdominal ventrites 3-6 transversely sulcate; aedeagus with ostium dorsal, median lobe oriented left side up in repose; right paramere long, tip attenuate, left short rounded at apex; female reproductive tract without dorsolateral bursal lobe, elongate spermatheca broadly attached laterally at base of common oviduct, spermatheca with appended gland, spermatheca with digitiform diverticulum near base, without spermathecal gland duct diverticulum.

Darodilia Castelnau 1867 [incl. Aulacocoelius Chaudoir 1870] LM87
    |--*D. mandibularis Castelnau 1867 LM87
    |--D. castelnaui Macleay 1888 LM87
    |--D. curta Sloane 1915 LM87
    |--D. emarginata Sloane 1898 LM87
    |--D. liopleura (Chaudoir 1870) [=*Aulacocoelius liopleurus] LM87
    |--D. longula Tschitschérine 1902 LM87
    |--D. macilenta Sloane 1895 LM87
    |--D. ovicollis (Macleay 1871) [=Meonis ovicollis] LM87
    |--D. robusta Sloane 1900 LM87
    `--D. rugisternis Sloane 1895 LM87

*Type species of generic name indicated

REFERENCES

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.