Aleochara

Aleochara curtula, copyright U. Schmidt.

Belongs within: Aleocharini.

Aleochara is a genus of aleocharine staphylinids characterised by a partially concealed head and five-segmented tarsi with the first segment much longer than the second. They are commonly associated with decaying matter (Olliff 1886).

Characters (from Olliff 1886): Mentum very short, narrowed and slightly emarginate in front. Ligula very short, broad, bifid. Maxillary palpi 5-segmented, the second and third segment subequal, the fifth very small. Labial palpi 4-segmented, the first segment longer than the others, the fourth very small. Mandibles unarmed. Labrum transverse, truncate in front. Head ovate, retracted, not narrowed behind. Antennae short, thickened towards the extremity, 1st, 2nd and 3rd segments elongate, the following, with the exception of the 11th, transverse. Prothorax broadly transverse, narrowed in front, rounded at the sides. Elytra slightly rounded behind. Abdomen either parallel-sided or narrowed behind. Legs rather short; intermediate coxae somewhat remote; tarsi 5:5:5, first segment of posterior pair distinctly longer than 2nd.

Aleochara Gravenhorst 1802 O86
    |--A. actae Olliff 1886 O86
    |--A. analis (Macleay 1871) [=Oxypoda analis] O86
    |--A. asiatica O86
    |--A. baliola Olliff 1886 O86
    |--A. bilineata P91
    |--A. bipunctata C01
    |--A. bisulcata (Redt. 1867) [=Oxypoda bisulcata] O86
    |--A. brachialis Jekel 1873 O86
    |--A. crassiuscula G89
    |--A. croceipennis Motschulsky 1858 [incl. A. maculipennis Kraatz 1859, A. sanguinipennis Kraatz 1859] O86
    |--A. curtula (Goeze 1777) M04
    |--A. fuscipes O86
    |--A. haemorrhoidalis O86
    |--A. insuavis Olliff 1886 O86
    |--A. lata Gravenhorst 1802 M04
    |--A. marginata Fauvel 1877 O86
    |--A. mastersi Macleay 1871 O86
    |--A. moesta C01
    |--A. nitida C01
    |--A. (Isochara) perturbans Fenyes 1921 F21
    |--A. puberula Klug 1883 [incl. A. armitagei Wollaston 1854, A. decorata Aubé 1850, A. dubia Fauvel 1863] O86
    |--A. punctum Fauvel 1878 O86
    |--A. scutellaris Lucas 1846 E12
    |--A. semirubra Fauvel 1877 O86
    |--A. speculifera Erichson 1842 O86
    |--A. subaenea ED04
    |--A. tristis C01
    `--A. vicina Olliff 1886 O86

*Type species of generic name indicated

REFERENCES

[C01] Csiki, E. 1901. Bogarak [Coleopteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] Band II. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 75–120. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[E12] Evenhuis, N. L. 2012. Publication and dating of the Exploration Scientifique de l’Algérie: Histoire Naturelle des Animaux Articulés (1846–1849) by Pierre Hippolyte Lucas. Zootaxa 3448: 1–61.

[ED04] Ewers, R. M., & R. K. Didham. 2004. A first assessment of New Zealand beetles restricted to large forest areas. Weta 27: 33–40.

[F21] Fenyes, A. 1921. New genera and species of Aleocharinae with a polytomic synopsis of the tribes. Bulletin of the Museum of Comparative Zoology 65 (2): 15–36.

[G89] Gestro, R. 1889. Viaggio ab Assab nel Mar Rosso dei signori G. Doria ed O. Beccari con il R. Avviso «Esploratore» dal 16 Novembre 1879 al 26 Febbraio 1880.—IV. Coleotteri. Annali del Museo Civico di Storia Naturale di Genova, Serie 2^a, 7: 5–72.

[M04] Maruyama, M. 2004. Redescription of the genus Creochara (Coleoptera: Staphylinidae: Aleocharinae: Aleocharini) and its systematic position. Canadian Entomologist 136: 621–637.

[O86] Olliff, A. S. 1886. A revision of the Staphylinidae of Australia. Part I. Proceedings of the Linnean Society of New South Wales, series 2, 1 (2): 403–473, pl. 7.

[P91] Petrushov, A. Z. 1991. Selection and mechanisms of ambush resistance in Metaseiulus occidentalis N. (Acarina: Phytoseiidae). In: Dusbábek, F., & V. Bukva (eds) Modern Acarology: Proceedings of the VIII International Congress of Acarology, held in České Budĕjovice, Czechoslovakia, 6–11 August 1990 vol. 2 pp. 741–748. SPB Academic Publishing: The Hague.

Abrotelina

Abroteles beaumonti, from Seevers (1957).

Belongs within: Corotocini.

The Abrotelina are a group of distinctively frail-bodied physogastric staphylinid beetles found in association with nasutitermitine termites in the Neotropical region (Seevers 1957).

Characters (from Seevers 1957): Frail-bodied, frequently with very thin sclerites; head and thorax slender, abdomen ovoidal, weakly to strongly physogastric. Entire clypeus very weakly sclerotised or not sclerotised; very little sclerotisation anterior to tentorial pit level. Pronotum small, its surface not impressed. Tergites relatively narrow, about three-fifths as broad as abdomen; inner paratergites large, about one-third as broad as tergites. Elytra generalized, contiguous. Wings normal. Antennae 11-segmented. Tarsi 4, 4, 4-segmented or 5, 5, 5-segmented. Abdominal segments 7-9 distinctive.

<==Abrotelina S57
    |--Oecidiophilus Silvestri 1946 S57
    |    `--*O. mimellus Silvestri 1946 [=O. oglobinii ms] S57
    |--Parvidolum Silvestri 1946 S57
    |    `--*P. microsomatis Silvestri 1946 [=P. microsamatis] S57
    |--Mormellus Silvestri 1946 S57
    |    `--*M. bicolor Silvestri 1946 S57
    |--Termitosius Silvestri 1901 S57
    |    `--*T. pauciseta Silvestri 1901 S57
    |--Termitoiceus Silvestri 1901 S57
    |    |--*T. anastrephoproctus Silvestri 1901 [=T. anastrephoroctus (l. c.)] S57
    |    `--T. simulans Silvestri 1946 [=T. anastrephoproctus simulans] S57
    |--Perlinctus Silvestri 1946 S57
    |    |--*P. quaesitus Silvestri 1946 S57
    |    `--P. foveicollis Seevers 1957 S57
    |--Eburniogaster Seevers 1938 S57
    |    |--*E. termitocola Seevers 1938 S57
    |    |--E. anahuaci Seevers 1957 S57
    |    `--E. texana (Brues 1902) [=Termitogaster texana] S57
    |--Fonsechellus Silvestri 1946 [incl. Trianellus Silvestri 1946] S57
    |    |--*F. diversicolor Silvestri 1946 S57
    |    |--F. bicolor Silvestri 1946 [=F. (*Trianellus) bicolor] S57
    |    `--F. fragilis Seevers 1957 S57
    |--Termitonidia Seevers 1938 S57
    |    |--*T. lunata Seevers 1938 S57
    |    |--T. michoacani Seevers 1957 S57
    |    |--T. jaliscensis Seevers 1957 S57
    |    `--T. tarascani Seevers 1957 S57
    `--Abroteles Casey 1890 [incl. Termitothymus Silvestri 1901] S57
         |--*A. beaumonti Casey 1890 S57
         |--A. badia Seevers 1957 S57
         |--A. bisetosus Seevers 1937 S57
         |--A. compacticornis Borgmeier 1950 S57
         |--A. lobata Seevers 1957 S57
         |--A. philetaerus (Silvestri 1901) (n. d.) [=*Termitothymus philetaerus] S57
         |--A. pubicollis Seevers 1957 S57
         `--A. rurrenabaquensis Seevers 1946 S57

*Type species of generic name indicated

REFERENCES

[S57] Seevers, C. H. 1957. A monograph on the termitophilous Staphylinidae (Coleoptera). Fieldiana Zoology 40: 1–334.

Corotocini

Thyreoxenus brevitibialis, copyright Taro Eldredge.

Belongs within: Staphylinidae.
Contains: Abrotelina, Termitogastrina.

The Corotocini are a group of physogastric staphylinid beetles found mostly in association with nasutitermitine termites, with a very few species found with the termitine genus Anoplotermes (Seevers 1957).

See also: The Corotocini in their gut-swollen glory.

Characters (from Seevers 1957): Mentum fused to submentum; mesocoxal acetabula not margined; metepimera abbreviated; hind coxae triangular; physogastric.

<==Corotocini S57
    |  i. s.: Termitochara Wasmann 1893 [Termitocharina] S57
    |           `--*T. kraatzi Wasmann 1893 S57
    |--Timeparthenina S57
    |    |--Termituncula Borgmeier 1950 S57
    |    |    `--*T. gracilipes Borgmeier 1950 S57
    |    `--+--Termitozophilus Silvestri 1901 [incl. Corymbogaster Mann 1923] S57
    |       |    |--*T. laetus Silvestri 1901 S57
    |       |    `--T. mirandus (Mann 1923) [=*Corymbogaster miranda] S57
    |       `--+--Autuoria Silvestri 1946 S57
    |          |    `--*A. elegantulum Silvestri 1946 [incl. A. orthocephali ms] S57
    |          `--+--Ptocholellus Silvestri 1946 S57
    |             |    `--*P. mimus Silvestri 1946 S57
    |             `--Timeparthenus Silvestri 1901 S57
    |                  |--*T. regius Silvestri 1901 S57
    |                  `--T. oglobini Silvestri 1946 [=Timeparthemus (l. c.) oglobini] S57
    `--+--+--Abrotelina S57
       |  `--Termitogastrina S57
       `--Corotocina S57
            |  i. s.: Eutermitoptochus Silvestri 1921 S57
            |           `--*E. novaehollandiae Silvestri 1921 S57
            |--Corotoca Schiødte 1853 S57
            |    |--*C. melantho Schiødte 1854 S57
            |    |--C. araujoi Seevers 1957 S57
            |    |--C. guyanae Mann 1923 S57
            |    `--C. phylo Schiødte 1854 S57
            `--+--+--Termitomimus Trägärdh 1907 S57
               |  |    |--*T. entendveniensis Trägärdh 1907 S57
               |  |    |--T. emersoni Seevers 1957 S57
               |  |    `--T. latipes Seevers 1957 S57
               |  `--+--Termitopullus Reichensperger 1922 [incl. Termitoscapha Bernhauer 1938] S57
               |     |    `--*T. sociusculus Reichensperger 1922 [incl. *Termitoscapha gestroi Bernhauer 1938] S57
               |     |--Termitoptocinus Silvestri 1921 S57
               |     |    `--*T. australiensis Silvestri 1921 S57
               |     `--Eburniola Mann 1923 S57
               |          |--*E. leucogaster Mann 1923 S57
               |          |--E. gastrovittata Seevers 1937 S57
               |          `--E. lujae Seevers 1957 S57
               `--+--Spirachtha Schiødte 1853 S57
                  |    |--*S. eurymedusa Schiødte 1854 S57
                  |    |--S. mirabilis Mann 1923 S57
                  |    `--S. schioedtei Mann 1923 S57
                  `--+--Oideprosoma Silvestri 1920 S57
                     |    `--*O. mirandum Silvestri 1920 S57
                     `--Thyreoxenus Mann 1923 S57
                          |--*T. parviceps Mann 1923 S57
                          |--T. albidus Seevers 1946 S57
                          |--T. autuorii Silvestri 1946 S57
                          |--T. boliviae Seevers 1946 S57
                          |--T. brevitibialis Seevers 1946 S57
                          |--T. convexinotus Seevers 1946 S57
                          |--T. cucullatus Seevers 1946 S57
                          |--T. major S57
                          |--T. pulchellus Mann 1923 S57
                          `--T. solomonensis Seevers 1937 S57

*Type species of generic name indicated

REFERENCES

[S57] Seevers, C. H. 1957. A monograph on the termitophilous Staphylinidae (Coleoptera). Fieldiana Zoology 40: 1–334.

Termitonannini

Termitonannus vagans, from Seevers (1957).

Belongs within: Staphylinidae.

The Termitonannini are a group of mostly limuloid staphylinid beetles found in association with nasutitermitine termites (Seevers 1957).

Characters (from Seevers 1957): Body usually non-physogastric and limuloid, very rarely physogastric. Head either prognathous or hypognathous; antennae 9-11-segmented, terminal antennal segments bearing one or more pairs of polytrichous coeloconic sensilla; mentum free except in certain derived genera. Mesocoxal cavities margined; hind coxae and metepimera generalised except in certain derived genera. Tarsi 4, 4, 4-segmented or 4, 4, 5-segmented.

<==Termitonannini S57
    |--Termitonannina S57
    |    |  i. s.: Termitocomes Seevers 1941 S57
    |    |           |--*T. wasmanni Seevers 1941 S57
    |    |           `--T. trinidadensis Seevers 1957 S57
    |    |--+--Eunannodes Silvestri 1946 S57
    |    |  |    `--*E. reconditi Silvestri 1946 S57
    |    |  `--Nannellus Silvestri 1946 S57
    |    |       `--*N. anoplotermitis Silvestri 1946 S57
    |    `--+--Termitonilla Borgmeier 1950 S57
    |       |    `--*T. luteola Borgmeier 1950 S57
    |       `--+--Macrotrichuris Silvestri 1946 S57
    |          |    |--*M. brasiliensis Silvestri 1946 S57
    |          |    `--M. notabilis Silvestri 1946 S57
    |          `--Termitonannus Wasmann 1902 [incl. Tetraphilus Silvestri 1946] S57
    |               |--*T. schmalzi Wasmann 1902 S57
    |               |--T. brachycerus Silvestri 1946 [=T. (*Tetraphilus) brachycerus] S57
    |               |--T. domunculi Silvestri 1946 S57
    |               |--T. echinoides Seevers 1946 S57
    |               |--T. gatuni Seevers 1941 S57
    |               |--T. major Wasmann 1902 S57
    |               |--T. microsomatis Silvestri 1946 S57
    |               |--T. parvulus Silvestri 1946 S57
    |               |--T. proximatus Silvestri 1946 S57
    |               |--T. setosus Seevers 1941 S57
    |               |--T. silvestrii Wasmann 1902 S57
    |               `--T. vagans Silvestri 1946 S57
    `--Perinthina [Perinthini] S57
         |--Termitopelta Borgmeier 1950 S57
         |    `--*T. fulgens Borgmeier 1950 S57
         `--+--Termitonicus Mann 1926 S57
            |    |--*T. mahout Mann 1926 S57
            |    `--T. uroclaviger Silvestri 1946 S57
            `--+--Macrognathellus Silvestri 1946 S57
               |    `--M. crassicornis (Wasmann 1902) (see below for synonymy) S57
               `--+--Poduroides Mann 1926 S57
                  |    `--*P. bovingi Mann 1926 S57
                  `--+--Perinthus Casey 1890 S57
                     |    |--*P. dudleyanus Casey 1890 S57
                     |    |    |--P. d. dudleyanus S57
                     |    |    |--P. d. major Seevers 1957 S57
                     |    |    `--P. d. wasmanni Mann 1923 S57
                     |    |--P. bolivari Seevers 1946 S57
                     |    |--P. guatemalae Seevers 1946 S57
                     |    |--P. hageni Seevers 1937 S57
                     |    |--P. mayae S57
                     |    |--P. tarsatus Mann 1923 S57
                     |    `--P. xenocostalis Seevers 1937 S57
                     `--+--Termitocola Seevers 1937 S57
                        |    |--*T. cylindricornis Seevers 1937 S57
                        |    `--T. silvestrii (Wasmann 1902) [=Perinthus silvestrii] S57
                        `--+--Lauella Mann 1921 S57
                           |    |--*L. vitiensis Mann 1921 [incl. Hetairotermes leai Cameron 1927] S57
                           |    |--L. javana Seevers 1957 S57
                           |    `--L. palauensis Seevers 1957 S57
                           `--+--Eutermitophila Cameron 1939 S57
                              |    `--*E. fletcheri Cameron 1939 S57
                              `--+--Perinthodes Seevers 1957 S57
                                 |    `--*P. africanus Seevers 1957 S57
                                 `--Paraperinthus Seevers 1957 S57
                                      `--*P. pauciseta Seevers 1957 S57

Macrognathellus crassicornis (Wasmann 1902) [=Perinthus crassicornis; incl. *M. paraguayensis Silvestri 1946] S57

*Type species of generic name indicated

REFERENCES

[S57] Seevers, C. H. 1957. A monograph on the termitophilous Staphylinidae (Coleoptera). Fieldiana Zoology 40: 1–334.

Termitogastrina

Termitogaster insolens, from Seevers (1957).

Belongs within: Corotocini.

The Termitogastrina are a lineage of physogastric staphylinids found in association with nasutitermitine termites of the genera Nasutitermes, Trinervitermes and Grallatotermes. Though difficult to characterise as a whole due to the derived morphology of some genera, but all carry the variously inflated abdomen recurved over the foreparts. In less derived species, the tarsi are 5, 5, 5-segmented, albeit with the fifth segment partially fused and not movable relative to the fourth segment; in some derived species the tarsi have become 4, 4, 4-segmented. These species have the sixth and seventh tergites characteristically enlarged with the seventh tergite shield-shaped (Seevers 1957).

Termitogastrina S57
    |--Termitopithus Seevers 1957 S57
    |    `--*T. crassiusculus Seevers 1957 S57
    |--Termitophya Wasmann 1902 S57
    |    |--*T. heyeri Wasmann 1902 S57
    |    |--T. amica Mann 1923 S57
    |    |--T. araujoi Borgmeier 1950 S57
    |    |--T. ecuadoriensis Seevers 1937 S57
    |    |--T. emersoni Seevers 1937 S57
    |    |--T. flaviventris Mann 1923 S57
    |    |--T. holmgreni Wasmann in Wasmann & Holmgren 1911 [incl. T. wasmanni Holmgren in Wasmann & Holmgren 1911] S57
    |    |--T. inornata Seevers 1937 S57
    |    |--T. piliventris Seevers 1937 S57
    |    `--T. punctata Mann 1923 S57
    `--+--+--Termitomorpha Wasmann 1894 [incl. Termitosomus Seevers 1939, Thaxteria Fenyes 1921] S57
       |  |    |--*T. meinerti Wasmann 1894 (see below for synonymy) S57
       |  |    |--T. costaricensis (Seevers 1946) [=Termitosomus costaricensis] S57
       |  |    |--T. fissipennis (Casey 1890) (see below for synonymy) S57
       |  |    |--T. huachii Seevers 1946 S57
       |  |    `--T. manni Seevers 1946 S57
       |  `--+--Trachopeplus Mann 1923 S57
       |     |    |--*T. setosus Mann 1923 S57
       |     |    `--T. disjunctipennis Seevers 1939 S57
       |     `--+--Termitoides Seevers 1939 S57
       |        |    `--*T. marginatus Seevers 1939 S57
       |        `--+--Xenopelta Mann 1923 S57
       |           |    |--*X. cornuta Mann 1923 S57
       |           |    `--X. tricornis (Mann 1923) [=Ceratoxenus tricornis] S57
       |           `--Xenogaster Wasmann 1891 S57
       |                |--*X. inflata Wasmann 1891 S57
       |                |--X. fossulata Mann 1923 S57
       |                |--X. glabriventris Seevers 1957 S57
       |                |--X. inquilina Borgmeier 1950 S57
       |                |--X. nana Seevers 1957 S57
       |                |--X. pilosula Seevers 1957 S57
       |                |--X. reichenspergeri Seevers 1957 S57
       |                `--X. subnuda Seevers 1957 S57
       `--+--+--Termitosynodes Seevers 1957 S57
          |  |    `--*T. williamsi Seevers 1957 S57
          |  `--Termitosyne Seevers 1957 S57
          |       `--*T. platygastra Seevers 1957 S57
          |--+--Neotermitogaster Seevers 1939 S57
          |  |    `--*N. colonus Seevers 1939 S57
          |  `--Termitogaster Casey 1889 S57
          |       |--*T. insolens Casey 1889 S57
          |       |--T. beniensis Seevers 1946 S57
          |       |--T. bicolor Seevers 1937 S57
          |       |--T. brevis Mann 1923 S57
          |       |--T. chavantinae Seevers 1957 S57
          |       |--T. diversicollis Seevers 1957 S57
          |       |--T. emarginatus Seevers 1946 S57
          |       |--T. emersoni Mann 1923 S57
          |       |--T. impressicollis Seevers 1937 S57
          |       |--T. magdalenae Seevers 1946 S57
          |       |--T. nigricollis (Silvestri 1901) [=Xenogaster nigricollis] S57
          |       |--T. puncticeps Seevers 1946 S57
          |       |--T. testaceus Seevers 1946 S57
          |       |--T. wasmanni (Holmgren in Wasmann & Holmgren 1911) [=Xenogaster wasmanni] S57
          |       `--T. wenzeli Seevers 1957 S57
          `--+--Termitotima Wasmann 1916 S57
             |    `--*T. assmuthi Wasmann 1916 S57
             `--+--Idiogaster Wasmann 1912 S57
                |    `--*I. escherichi Wasmann 1912 S57
                `--+--+--Affinoptochus Kemner 1925 S57
                   |  |    `--*A. exclusus Kemner 1925 S57
                   |  `--+--Termitella Wasmann 1911 (see below for synonymy) S57
                   |     |    |--*T. lujae Wasmann 1911 S57
                   |     |    |--T. foveolata (Reichensperger 1922) [=*Termitissa foveolata] S57
                   |     |    |--T. gridelli (Bernhauer 1938) [=*Termitoctesis gridelli] S57
                   |     |    `--T. rubricollis Seevers 1957 S57
                   |     `--+--Termitellodes Seevers 1957 S57
                   |        |    `--*T. lativentris Seevers 1957 S57
                   |        `--Idioptochus Seevers 1957 S57
                   |             `--*I. lehmensicki Seevers 1957 S57
                   `--+--Paracorotoca Warren 1920 S57
                      |    `--*P. akermani (Warren 1914) [=Corotoca akermani] S57
                      |--Millotoca Paulian 1948 S57
                      |    |--*M. mirotermitidis Paulian 1948 S57
                      |    `--M. tanganyuikae Seevers 1957 S57
                      `--Termitoptochus Silvestri 1910 S57
                           |--*T. indicus Silvestri 1910 S57
                           |--T. ceylonicus Silvestri 1921 S57
                           |--T. luzonicus Silvestri 1921 S57
                           |--T. peninsularis Silvestri 1921 S57
                           |--T. philippinus Silvestri 1921 S57
                           `--T. sumatranus Silvestri 1921 S57

Termitella Wasmann 1911 [incl. Termitissa Reichensperger 1912, Termitoctesis Bernhauer 1938] S57

Termitomorpha fissipennis (Casey 1890) [=Termitogaster fissipennis, *Termitosomus fissipennis; incl. Termitogaster simopelta Mann 1923] S57

*Termitomorpha meinerti Wasmann 1894 [incl. *Thaxteria insularis Fenyes 1921, Termitogaster simulans Mann 1923, Termitosomus simulans] S57

*Type species of generic name indicated

REFERENCES

[S57] Seevers, C. H. 1957. A monograph on the termitophilous Staphylinidae (Coleoptera). Fieldiana Zoology 40: 1–334.

Termitopaediini

Lateral and dorsal views of Termitobia burgeoni, from Seevers (1957).

Belongs within: Staphylinidae.

The Termitopaediini is a group of physogastric staphylinid beetles found in association with fungus-culturing termites in Africa and southern Asia (Seevers 1957).

Characters (from Seevers 1957): Maxillary acetabula moderately sized; galea and lacinia moderate in length, galea usually foliate and membranous; terminal antennal segments without coeloconic sensilla; mesocoxal acetabula large, shallow, faintly margined externally or not at all; broad mesosternal process rounded or angulate; middle coxae narrowly or moderately widely separated; hind coxae subtriangular, not produced beyond lateral articulation; metepimera obliquely truncated posteriorly, not prolonged beyond coxal articulation; tarsi 4, 5, 5-segmented.

<==Termitopaediini S57
    |  i. s.: Termitobaena Bernhauer 1915 S57
    |           `--*T. bryanti Bernhauer 1915 S57
    |         Termitotropha Wasmann 1899 S57
    |           `--*T. oneili Wasmann 1899 S57
    |--+--Termitotecna Wasmann 1912 S57
    |  |    `--*T. braunsi Wasmann 1912 S57
    |  |--Protermitobia Seevers 1957 S57
    |  |    |--*P. kirbyi Seevers 1957 S57
    |  |    `--P. comes Seevers 1957 S57
    |  `--Termitobia Wasmann 1891 S57
    |       |--*T. physogastra Wasmann 1891 [=T. gastrophysa (l. c.)] S57
    |       |--T. burgeoni Cameron 1930 S57
    |       |--T. paolii Gridelli 1928 S57
    |       `--T. rhodesiae Seevers 1957 S57
    `--+--Termitopaedia Wasmann 1911 S57
       |    `--*T. kohli Wasmann 1911 S57
       |--Termitolinus Wasmann 1911 S57
       |    `--*T. natalensis Wasmann 1911 S57
       |--Neodioxeuta Seevers 1957 S57
       |    |--*N. oudemansi (Franssen 1932) [=Jacobsonella oudemansi] S57
       |    `--N. burmae Seevers 1957 S57
       |--Termitopulex Fauvel 1899 [incl. Silvestrinus Bernhauer 1932] S57
       |    |--*T. grandicornis Fauvel 1899 [incl. *Silvestrinus erythraeanus Bernhauer 1932] S57
       |    |--T. natalensis Wasmann 1912 S57
       |    `--T. sjoestedti Eichelbaum in Sjöstedt 1908 S57
       `--Dioxeuta Sharp 1899 [incl. Jacobsonella Silvestri 1910] S57
            |--*D. microps Sharp 1899 [incl. *Jacobsonella termitobia Silvestri 1910] S57
            |--D. flavescens (Cameron 1939) [=Jacobsonella flavescens] S57
            `--D. indosinensis (Silvestri 1946) [=Jacobsonella indosinensis] S57

*Type species of generic name indicated

REFERENCES

[S57] Seevers, C. H. 1957. A monograph on the termitophilous Staphylinidae (Coleoptera). Fieldiana Zoology 40: 1–334.

Athetini

Atheta fungi, from Insect Foto.

Belongs within: Aleocharinae.

The Athetini are a diverse group of staphylinid beetles, primarily found in humid microhabitats among decaying organic matter. Two genera of Athetini, Termitotelus and Coptotermoecia, are found in association with termites (Seevers 1957).

Characters (from Fenyes 1921; Seevers 1957): Head visible from above; antennae inserted medial to eyes. Antennae 11-segmented; maxillary palpi 4-segmented; labial palpi three-segmented. Tarsi 4, 5, 5-segmented.

Athetini S57
    |--Termitotelus Wasmann 1908 [Termitotelina] S57
    |    `--*T. schultzei Wasmann 1908 S57
    |--Coptotermoecia Oke 1933 [Coptotermoeciina] S57
    |    |--*C. alutacia Oke 1933 S57
    |    `--C. gayi Seevers 1957 S57
    |--Mesaraeus Fenyes 1921 F21
    |    `--*M. laevigatus Fenyes 1921 F21
    |--Falagriota F21
    |--Myrmecopora Saulcy 1864 F21, O86
    |    |--M. senilis Fauvel 1878 O86
    |    `--M. termitophila Cameron 1948 S57
    |--Lipodonta Fenyes 1921 F21
    |    `--*L. veris Fenyes 1921 F21
    |--Lypoglossa F21
    |--Tomoglossa Kraatz 1856 F21, G02 [incl. Noverota Casey 1910 G02]
    |    |--*T. luteicornis (Erichson 1837) [=Homalota luteicornis] G02
    |    |--T. aegyptiaca Scheerpeltz 1963 G02
    |    |--T. arizonica Gusarov 2002 G02
    |    |--T. basicornis (Pace 1985) G02
    |    |--T. brakmani Scheerpeltz 1963 G02
    |    |--T. breitiana (Scheerpeltz 1957) G02
    |    |--T. decora (Casey 1910) [=Noverota decora] G02
    |    |--T. floridana Gusarov 2002 G02
    |    |--T. franzi Pace 1991 G02
    |    |--T. fuliginosa Pace 1999 G02
    |    |--T. heydemanni Lohse 1977 G02
    |    |--T. improvisa (Casey 1910) [=Noverota improvisa] G02
    |    |--T. laeta Eppelsheim 1884 G02
    |    |--T. nepalensis (Pace 1987) G02
    |    |--T. ocularis Pace 1991 G02
    |    |--T. ornatella (Casey 1910) [=*Noverota ornatella] G02
    |    |--‘Noverota’ personata Casey 1910 G02
    |    |--T. subcorticalis (Cameron 1939) G02
    |    `--T. venusta (Pace 1984) G02
    `--Atheta Thomson 1858 [Athetae] G02
         |--A. celata (Erichson 1837) P05
         |--A. convivens Silvestri 1945 S57
         |--A. fungi C01
         |--A. (Datomicra) jantarica Paśnik 2005 P05
         |--A. parva C01
         |--A. silvestrii Seevers 1957 [=A. termitaria Seevers 1945 non Bernhauer 1932] S57
         |--A. syntermitis Silvestri 1945 S57
         |--A. termitaria Bernhauer 1932 S57
         |--A. termitobia Wasmann 1894 S57
         |--A. termitophila Motschulsky 1859 S57
         `--A. triangulum K09

*Type species of generic name indicated

REFERENCES

[C01] Csiki, E. 1901. Bogarak [Coleopteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] Band II. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 75–120. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[F21] Fenyes, A. 1921. New genera and species of Aleocharinae with a polytomic synopsis of the tribes. Bulletin of the Museum of Comparative Zoology 65 (2): 15–36.

[G02] Gusarov, V. I. 2002. A revision of Nearctic species of the genus Tomoglossa Kraatz, 1856 (Coleoptera: Staphylinidae: Aleocharinae). Zootaxa 30: 1–19.

[K09] Köhler, G. 2009. Heuschrecken (Saltatoria) und Ohrwürmner (Dermaptera) im Immissionsgebiet des Düngemittelwerkes Steudnitz/Thüringen—eine Langzeitstudie (1978–2001). Mauritiana 20 (3): 601–646.

[O86] Olliff, A. S. 1886. A revision of the Staphylinidae of Australia. Part I. Proceedings of the Linnean Society of New South Wales, series 2, 1 (2): 403–473, pl. 7.

[P05] Paśnik, J. 2005. Fossils of Staphylinidae from Baltic amber: A new genus and three new species (Insecta, Coleoptera, Staphylinidae). Senckenbergiana Biologica 85 (1): 97–100.

[S57] Seevers, C. H. 1957. A monograph on the termitophilous Staphylinidae (Coleoptera). Fieldiana Zoology 40: 1–334.

Last updated: 25 May 2017.

Termitodiscini

Termitodiscus heimi, from Seevers (1957).

Belongs within: Staphylinidae.

The Termitodiscini are a group of minute limuloid staphylinid beetles found in association with fungus-growing termites in Africa and Asia (Seevers 1957).

Characters (from Seever 1957): Body-form limuloid; head concealed beneath very large pronotum, hypognathous; antennae inserted in deep fossae medial to eyes, their scapes resting in these fossae; antennae 10- or 11-segmented, their second and third segments small, the fourth to terminal segment forming a compact club; pronotum approximately twice as broad as long, its sides and apex continuously arcuate; pronotal hypomera very large, but not visible from the side; elytra broad, their epipleura very large; prosternum keeled; mesocoxae set in margined acetabula, narrowly or broadly separated; hind coxae broad, produced laterally to meet elongated metepimera; tarsi 4, 5, 5-segmented; abdomen acuminate; aedeagus typically aleocharine in structure.

<==Termitodiscini [Termitodiscinae] S57
    |--Discoxenus Wasmann 1904 S57
    |    |--*D. assmuthi Wasmann 1904 S57
    |    |--D. acuticornis Wasmann 1916 S57
    |    |--D. crassicornis Wasmann 1916 S57
    |    `--D. lepisma Wasmann 1904 S57
    |--Termitogerrus Bernhauer 1932 [incl. Lissodiscus Grassé & Poisson 1940] S57
    |    |--*T. burgeoni (Bernhauer 1932) [=Termitodiscus (*Termitogerrus) burgeoni] S57
    |    |--T. habilis (Silvestri 1947) [=Lissodiscus habilis] S57
    |    |--T. lepidulus (Grassé & Poisson 1940) [=Termitodiscus (*Lissodiscus) lepidulus] S57
    |    |--T. bellicosi (Silvestri 1905) [=Termitodiscus bellicosi, Lissodiscus bellicosi] S57
    |    `--T. quaerens (Silvestri 1947) [=Lissodiscus quaerens] S57
    `--Termitodiscus Wasmann 1899 S57
         |--*T. heimi Wasmann 1899 S57
         |--T. braunsi Wasmann 1912 S57
         |--T. butteli Wasmann 1916 S57
         |--T. escherichi Wasmann in Escherich 1911 [incl. T. escherichi var. piceus Wasmann 1916] S57
         |--T. minutus Cameron 1926 S57
         |--T. splendidus Wasmann 1899 S57
         `--T. transvaalensis Silvestri 1947 S57

*Type species of generic name indicated

REFERENCES

[S57] Seevers, C. H. 1957. A monograph on the termitophilous Staphylinidae (Coleoptera). Fieldiana Zoology 40: 1–334.

Termitohospitina

Termitohospes unicolor, from Seevers (1957).

Belongs within: Staphylinidae.

The Termitohospitina is a South American group of limuloid staphylinid beetles, found in association with termites of the Nasutitermitinae (Seevers 1957).

Characters (from Seevers 1957): Body-form limuloid. Head strongly deflexed; concealed by large, shield-like pronotum except in Paratermitosocius (in which small part of vertex is visible). Pronotum with anterior and lateral margins continuously arcuate; surface not impressed. Anterior margin of submentum straight or slightly arcuate but not emarginate; anterior angles of mentum produced, apex of mentum emarginate. Abdomen slender, acuminate. Ninth tergite bifid, otherwise unmodified.

<==Termitohospitina S57
    |--Termitohospes Seevers 1941 S57
    |    |--*T. miricorniger Seevers 1941 S57
    |    |--T. brasiliana Seevers 1957 S57
    |    |--T. guianae Seevers 1941 S57
    |    |--T. limulus Seevers 1957 S57
    |    |--T. panamensis Seevers 1941 S57
    |    |--T. silvestrii Seevers 1957 S57
    |    |--T. tachyporoides Seevers 1957 S57
    |    `--T. unicolor (Silvestri 1947) [=Termitomus unicolor] S57
    `--+--Paratermitosocius Seevers 1941 S57
       |    `--*P. vestitus (Mann 1923) [=Perinthus vestitus] S57
       |--Termitosocius Seevers 1941 S57
       |    `--*T. microps Seevers 1941 S57
       `--+--Blapticoxenus Mann 1923 [incl. Termitocolonus Seevers 1941] S57
          |    `--*B. brunneus Mann 1923 [incl. *Termitocolonus ericiogaster Seevers 1941] S57
          |--Termitosodalis Seevers 1941 [incl. Termitomus Silvestri 1947] S57
          |    |--*T. barticae Seevers 1941 S57
          |    `--T. fasciatus (Silvestri 1947) [=*Termitomus fasciatus, Termitohospes fasciatus] S57
          `--Hetairotermitina S57
               |--Termitobra Seevers 1957 S57
               |    `--*T. perinthoides Seevers 1957 S57
               `--Hetairotermes Cameron 1920 [=Termophila Lea 1910 non Grassi 1887] S57
                    |--*H. latebricola (Lea 1910) [=*Termophila latebricola] S57
                    |--H. agilis Cameron 1920 S57
                    |--H. bryanti Cameron 1950 S57
                    |--H. formicicola Oke 1933 S57
                    |--H. insulanus Seevers 1957 S57
                    |--H. piceus Cameron 1920 S57
                    `--H. punctiventris (Lea 1910) [=Termophila punctiventris] S57

*Type species of generic name indicated

REFERENCES

[S57] Seevers, C. H. 1957. A monograph on the termitophilous Staphylinidae (Coleoptera). Fieldiana Zoology 40: 1–334.

Termitusina

Termitusa lativentris, from Seevers (1957).

Belongs within: Staphylinidae.

The Termitusina are a group of limuloid staphylinid beetles found living in the Ethiopian region in association with termites of the genera Cubitermes, Thoracotermes and Pericapritermes.

See also: Life with termites.

Characters (from Seevers 1957): Body-form limuloid. Head strongly deflexed, usually with long axis vertical; small part of vertex visible from above. Pronotum large, shield-like; sides strongly arcuate, converging anteriorly but anterior margin transverse, anterior angles evident. Anterior margin of submentum emarginate. Anterior angles of mentum not produced, apex straight. Abdomen acuminate. Ninth tergite deeply bifid, the two parts elongated as stout processes.

<==Termitusina JK75
    |--Thoracotusa JK75
    |    |--T. alzadae JK75
    |    |--T. carrissoensis JK75
    |    `--T. kymrae JK75
    |--Termitusodes JK75
    |    |--T. congolensis JK75
    |    |--T. lativentris JK75
    |    |--T. leleupi JK75
    |    |--T. media JK75
    |    `--T. summa JK75
    `--Termitusa Wasmann 1905 [incl. Glaphyrinus Bernhauer in Bernhauer & Paulian 1942] S57
         |--*T. sjoestedti Wasmann 1905 S57
         |--T. atlanticus JK75
         |--T. bandai JK75
         |--T. boultoni JK75
         |--T. cameroni Reichensperger 1929 S57
         |--T. cloveri JK75
         |--T. coatoni JK75
         |--T. delignei JK75
         |--T. dybasi JK75
         |--T. escalerae Fauvel 1906 S57
         |--T. hexaseta JK75
         |--T. hystrix Wasmann 1911 S57
         |--T. katangensis JK75
         |--T. lativentris Seevers 1957 S57
         |--T. lightningae JK75
         |--T. lujae Wasmann 1911 S57
         |--T. malawiensis JK75
         |--T. minor JK75
         |--T. mixaseta JK75
         |--T. mundus (Bernhauer in Bernhauer & Paulian 1942) [=*Glaphyrinus mundus, Glaphyrius mirandus ms] S57
         |--T. muneris JK75
         |--T. pasteelsi JK75
         |--T. quadricollis Cameron 1952 S57
         |--T. regina JK75
         |--T. ruellei JK75
         |--T. sheasbyi JK75
         |--T. superhirsutus JK75
         |--T. ugandae JK75
         |--T. viegasi JK75
         |--T. wasmanni JK75
         `--T. williamsi JK75

*Type species of generic name indicated

REFERENCES

[JK75] Jacobson, H. R., & D. H. Kistner. 1975. Numeric analyses of relationships of genera and species of the subtribe Termitusina (Coleoptera: Staphylinidae). Systematic Zoology 24 (2): 191–198.

[S57] Seevers, C. H. 1957. A monograph on the termitophilous Staphylinidae (Coleoptera). Fieldiana Zoology 40: 1–334.

Reichenbachia

Female Reichenbachia, photographed by Mike Quinn.

Belongs within: Goniaceritae.

Reichenbachia is a large cosmopolitan genus of pselaphine beetles, including over 300 species.

Characters (from Park 1942): Ventral surface of head with median longitudinal carina or carinoid swelling; antennae eleven-segmented; maxillary palpi four-segmented; epistomal region generally simple, sometimes slightly expanded laterally or vertically elevated; pronotum with disc simple, evenly convex, three subbasal foveae present with lateral foveae much larger than median fovea; each elytron with distinct dorsal or discal stria and two or three basal foveae; middle coxae contiguous; posterior coxae well separated; abdomen with distinct lateral margin.

<==Reichenbachia
    |--R. affinissima Reitter 1883 NG05
    |--R. amica Reitter 1892 NG05
    |--R. grabowskyi Reitter 1882 NG05
    |--R. integrostriata Reitter 1883 NG05
    |--R. invalida Reitter 1882 NG05
    |--R. lamellicornis Reitter 1882 NG05
    |--R. muscorum Raffray 1904 NG05
    |--R. negligens Reitter 1883 NG05
    |--R. punctithorax Reitter 1882 NG05
    |--R. rufa Schmidt-Göbel 1838 NG05
    |--R. schaufussi Reitter 1882 NG05
    |--R. singapuriensis Raffray 1895 NG05
    |--R. telangensis Reitter 1883 NG05
    `--R. tubericollis Raffray 1891 NG05

*Type species of generic name indicated

REFERENCES

[NG05] Nomura, S., & I. Abd Ghani. 2005. Faunistic notes on the pselaphine species of the supertribes Goniaceritae, Pselaphitae and Clavigeritae from Malaysia and Singapore (Coleoptera: Staphylinidae: Pselaphinae). Serangga 10 (1-2): 1-36.

Park, O. 1942. A study in Neotropical Pselaphidae. Northwestern University: Evanston and Chicago.

Goniaceritae

Specimen of Plagiophorus, from here.

Belongs within: Staphylinidae.
Contains: Natypleurina, Reichenbachia.

The Goniaceritae are a group of pselaphine beetles with clearly separated hind coxae, short trochanters, three-segmented tarsi with the first segment smaller than the other two, asymmetrical or single tarsal claws, and generally complete parameres in the male genitalia (Nomura & Sakchoowong).

<==Goniaceritae
    |--Iniocyphini NG05
    |    |--Natypleurina NG05
    |    `--Sunorfa [Iniocyphina] NG05
    |         `--S. clavata (Raffray 1895) NG05
    |--Proterini NG05
    |    |--Mechanicus cribratus Raffray 1893 NG05
    |    |--Proterus punctatus NG05
    |    `--Bythinoderes grabowskyi Reitter 1883 NG05
    |--Atychodea [Tychini] NG05
    |    |--A. lenticornis Reitter 1883 NG05
    |    |--A. raffrayi Reitter 1883 NG05
    |    |--A. simoniana Reitter 1883 NG05
    |    `--A. singularis Reitter 1883 NG05
    |--Arnylliini NG05
    |    |--Awas Löbl 1994 NG05
    |    |    |--*A. giraffa Löbl 1994 NG05
    |    |    `--A. rajah Nomura 2004 NG05
    |    `--Harmophorus Motschulsky 1851 [incl. Arnyllium Reitter 1884] NG05
    |         |--H. ciliatum (Raffray 1894) NG05
    |         |--H. cultratum (Raffray 1894) NG05
    |         |--H. ensipes (Reitter 1883) NG05
    |         |--H. parviceps (Reitter 1883) NG05
    |         `--H. pectinatum (Reitter 1883) NG05
    |--Brachyglutina [Brachyglutini, Pselaptina] NG05
    |    |--Ephymata mucronata (Raffray 1895) NG05
    |    |--Trissemus NG05
    |    |--*Noduliceps latifrons NG05
    |    |--Reichenbachia NG05
    |    |--Atenisodus Raffray 1904 NG06
    |    |    `--A. longicornis (Raffray 1895) NG05
    |    |--Eupines King 1866 NG05
    |    |    |--E. atomus (Schaufuss 1887) NG05
    |    |    `--E. sphaerica (Motchulsky 1851) NG05
    |    |--Comatopselaphus Schaufuss 1882 NG05
    |    |    |--C. opacicollis Schaufuss 1882 NG05
    |    |    `--C. puncticollis Raffray 1895 NG05
    |    |--Rybaxis NG05
    |    |    |--R. amica Reitter 1883 NG05
    |    |    |--R. imperatrix Schaufuss 1882 NG05
    |    |    |--R. nubile Reitter 1882 NG05
    |    |    `--R. sarawakensis Schaufuss 1882 NG05
    |    `--Batraxis Reitter 1881 NG05
    |         |--B. brevis Raffray 1895 NG05
    |         |--B. curvispina Raffray 1892 NG05
    |         |--B. elegans Raffray 1895 NG05
    |         |--B. foveata Raffray 1895 NG05
    |         |--B. gracilicornis Raffray 1895 NG05
    |         |--B. instabilis Raffray 1895 NG05
    |         |--B. obliqua Raffray 1895 NG05
    |         |--B. pumilio Raffray 1895 NG05
    |         |--B. quadrata Raffray 1895 NG05
    |         |--B. truncaticornis Raffray 1895 NG05
    |         `--B. variabilis Raffray 1890 NG05
    `--Plagiophorus Motschulsky 1851 [incl. Cyathiger King 1865; Cyathigerini] NG05
         |--P. baumeisteri (Reitter 1883) NG05
         |--P. besucheti (Burckhardt 1985) NG05
         |--P. borneensis (Reitter 1883) NG05
         |--P. bulbifer (Bryant 1915) NG05
         |--P. cribratus (Raffray 1895) NG05
         |--P. decamerus (Burckhardt 1985) NG05
         |--P. dispar (Bryant 1915) NG05
         |--P. elongatus (Bryant 1915) NG05
         |--P. endecamerus (Burckhardt 1985) NG05
         |--P. heterocerus (Raffray 1895) NG05
         |--P. impar (Raffray 1895) NG05
         |--P. inexpectus (Raffray 1895) NG05
         |--P. magnicornis ((Burckhardt 1985) NG05
         |--P. malayanus (Bryant 1915) NG05
         |--P. matangensis (Bryant 1915) NG05
         |--P. mirandus (Bryant 1915) NG05
         |--P. murphyi (Burckhardt 1985) NG05
         |--P. myrmecophilus (Bryant 1915) NG05
         |--P. octomerus (Burckhardt 1985) NG05
         |--P. penangensis (Burckhardt 1985) NG05
         |--P. pseudoheptamerus (Burckhardt 1985) NG05
         |--P. raffrayi (Bryant 1915) NG05
         |--P. rufus (Raffray 1895) NG05
         |--P. sarawakensis (Bryant 1915) NG05
         |--P. schaufussi Reitter 1883 NG05
         |--P. sericeus (Raffray 1895) NG05
         |--P. simoni (Reitter 1883) NG05
         |--P. variolosus (Raffray 1895) NG05
         |--P. vicinus (Burckhardt 1985) NG05
         `--P. viduus (Raffray 1895) NG05

*Type species of generic name indicated

REFERENCES

[NG05] Nomura, S., & I. Abd Ghani. 2005. Faunistic notes on the pselaphine species of the supertribes Goniaceritae, Pselaphitae and Clavigeritae from Malaysia and Singapore (Coleoptera: Staphylinidae: Pselaphinae). Serangga 10 (1-2): 1-36.

Pselaphitae

Centrophthalmus punctipennis, from here.

Belongs within: Staphylinidae.

The Pselaphitae are a large group of beetles that usually have eleven-segmented antennae and greatly developed maxillary palpi (Imms et al. 1977).

<==Pselaphitae
    |--Ctenistes [Ctenistini] NG05
    |    |--C. mitis Schaufuss 1882 NG05
    |    `--C. parviceps G89
    |--Odontalgus [Odontalgini] NG05
    |    `--O. vestitus Schaufuss 1886 NG05
    |--Sabarhytus Löbl 2000 [Arhytodini] NG05
    |    `--S. kinabalu Löbl 2000 NG05
    |--Pselaphus [Pselaphini] NG05
    |    |--P. biocellatus Reitter 1883 NG05
    |    |--P. calopygaeus Schaufuss 1887 NG05
    |    |--P. lativentris Reitter 1882 NG05
    |    |--P. parvipalpis Reitter 1882 [incl. P. bivestitus Schaufuss 1887] NG05
    |    |--P. sexstriatus Reitter 1883 NG05
    |    `--P. unipunctatus Reitter 1883 NG05
    |--Hybocephalini NG05
    |    |--Stipesa [incl. Filiger] NG05
    |    |--Pseudapharina prima (Reitter 1883) NG05
    |    |--Apharinodes NG05
    |    |    |--A. miranda Raffray 1895 NG05
    |    |    `--A. squamosa Raffray 1890 NG05
    |    |--Apharina NG05
    |    |    |--A. borneensis Raffray 1896 NG05
    |    |    `--A. fuscipennis Reitter 1883 NG05
    |    `--Hybocephalodes NG05
    |         |--H. dentiventris Reitter 1883 NG05
    |         |--H. informis Reitter 1883 NG05
    |         `--H. telangensis Reitter 1883 NG05
    |--Tmesiphorini NG05
    |    |--Ctenotillus NG05
    |    |--Raphitreus dentimanus Raffray 1890 NG05
    |    |--Chandleriella Hlavac 2000 NG05
    |    |    `--*C. termitophila (Bryant 1915) [=Lasinus termitophilus] NG05
    |    |--Tmesiphorus NG05
    |    |    |--T. armatus Raffray 1882 NG05
    |    |    |--T. clavatus Raffray 1895 NG05
    |    |    `--T. matangensis Bryant 1915 NG05
    |    |--Dacnotillus NG05
    |    |    |--D. pilosus Raffray 1895 NG05
    |    |    |--D. transversus Raffray 1895 NG05
    |    |    `--D. tuberculatus Raffray 1895 NG05
    |    |--Ancystrocerus NG05
    |    |    |--A. longicornis Raffray 1895 NG05
    |    |    |--A. pallidus Raffray 1895 NG05
    |    |    |--A. punctatus Raffray 1895 NG05
    |    |    `--A. rugicollis Raffray 1895 NG05
    |    `--Pseudophanias Raffray 1890 NG05
    |         |--*P. malaianus Raffray 1890 NG05
    |         |--P. cribricollis Raffary 1895 NG05
    |         |--P. heterocerus Raffray 1895 NG05
    |         |--P. pilosus Raffray 1895 NG05
    |         |--P. puberulus Raffray 1904 NG05
    |         `--P. punctatus Raffray 1904 NG05
    `--Tyrini NG05
         |--Centrophthalmina NG05
         |    |--Enantius NG05
         |    |    |--E. fortis Raffray 1895 NG05
         |    |    |--E. rostratus Reitter 1883 NG05
         |    |    `--E. singapuriensis Raffray 1895 NG05
         |    `--Centrophthalmus NG05
         |         |--C. bispina Reitter 1883 NG05
         |         |--C. divisus Reitter 1883 NG05
         |         `--C. punctipennis NG05
         |              |--C. p. punctipennis NG05
         |              `--C. p. punctatissimus Schaufuss 1882 NG05
         `--Tyrina NG05
              |--Labomimus heterocerus (Raffray 1882) [=Pselaphodes heterocerus; incl. P. antennatus Bryant 1915] NG05
              |--Paralasinus NG05
              |--Taiwanophodes Hlavác 2002 NG05
              |--Nomuraius Hlavác 2002 NG05
              |--Linan Hlavác 2002 NG05
              |--Indophodes Hlavác 2002 NG05
              |--Tyrodes clavatus Raffray 1895 NG05
              |--Ryxabis anthicoides Westwood 1870 NG05
              |--Tyrus NG05
              |    |--T. corticalis (Raffray 1890) NG05
              |    `--T. myrmecophilus Bryant 1915 NG05
              `--Pselaphodes NG05
                   |--P. clavatus Raffray 1895 NG05
                   |--P. foveolatus Raffray 1892 NG05
                   `--P. villosulus Westwood 1870 NG05

*Type species of generic name indicated

REFERENCES

[G89] Gestro, R. 1889. Viaggio ab Assab nel Mar Rosso dei signori G. Doria ed O. Beccari con il R. Avviso «Esploratore» dal 16 Novembre 1879 al 26 Febbraio 1880. – IV. Coleotteri. Annali del Museo Civico di Storia Naturale di Genova, Serie 2^a 7: 5-72.

Imms, A. D., O. W. Richards & R. G. Davies. 1977. Imms' General Textbook of Entomology. Springer.

[NG05] Nomura, S., & I. Abd Ghani. 2005. Faunistic notes on the pselaphine species of the supertribes Goniaceritae, Pselaphitae and Clavigeritae from Malaysia and Singapore (Coleoptera: Staphylinidae: Pselaphinae). Serangga 10 (1-2): 1-36.