Isobactrus

Dorsal view of Isobactrus uniscutatus, from Bartsch & Smit (2006).


Belongs within: Halacaridae.

Isobactrus is a cosmopolitan genus of marine halacarid mites with the ventral armature reduced and not forming a solid shield, and a club-like seta in front of the lateral disti-dorsal seta on tarsi I and II (Newell 1947).

Characters (from Newell 1947): Genital opening of female ventral in position, not guarded by opercula and surrounded by only three pairs of setae. Adults more heavily armoured than nymphs; all dorsal and ventral plates separate. Ventral plates greatly reduced, not forming a solid shield. Femur I greatly swollen, less than 1.5 times as long as high. Tarsi I and II each with a single club-like seta in front of lateral disti-dorsal seta. Two claws on all tarsi.

<==Isobactrus Newell 1947 AB07
    |--I. microdens AB07
    `--+--I. macquariensis AB07
       `--+--I. magnus (Lohmann 1907) AB07, N47 [=Rhombognathus magnus N47]
          `--+--I. calderensis AB07
             `--+--+--I. obesus AB07
                |  `--+--I. luxtoni AB07
                |     `--+--I. asper AB07
                |        `--+--I. ponapensis AB07
                |           `--+--I. australiensis AB07
                |              `--I. pacificus AB07
                `--+--I. peregrinus AB07
                   `--+--+--I. gryposetus AB07
                      |  |--+--I. hamatus AB07
                      |  |  `--I. levis (Viets 1927) AB07, N47 [=Rhombognathus levis N47]
                      |  `--+--I. hartmanni AB07
                      |     `--+--I. dentatus AB07
                      |        |--I. latistriatus AB07
                      |        `--I. tuberculatus AB07
                      `--+--I. ponticus AB07
                         `--+--I. rugosus AB07
                            `--+--I. uniscutatus AB07
                               `--+--I. cryptorhynchus (Trouessart 1901) AB07, N47 [=Rhombognathus cryptorhynchus N47]
                                  `--+--I. ungulatus AB07
                                     `--+--*I. setosus (Lohmann 1889) N47, AB07, N47 (see below for synonymy)
                                        |--I. hutchinsoni Newell 1947 AB07, N47
                                        `--I. pulchellus AB07

*Isobactrus setosus (Lohmann 1889) N47, AB07, N47 [=Aletes setosus N47, Rhombognathus setosus N47]

*Type species of generic name indicated

REFERENCES

[AB07] Abé, H., & I. Bartsch. 2007. Historical biogeography of the halacarid mite genus Isobactrus (Prostigmata, Halacaridae). In: Morales-Malacara, J. B., V. M. Behan-Pelletier, E. Ueckermann, T. M. Pérez, E. G. Estrada-Venegas & M. Badii (eds) Acarology XI: Proceedings of the International Congress pp. 119–126. Instituto de Biología and Faculdad de Ciencias, Universidad Nacional Autónoma de México, Sociedad Latinoamericana de Acarología: México.

[N47] Newell, I. M. 1947. A systematic and ecological study of the Halacaridae of eastern North America. Bulletin of the Bingham Oceanographic Collection 10 (3): 1–232.

Stigmaeus

Dorsal (left) and ventral views of Stigmaeus alvandis, from Khanjani & Ueckermann (2002).


Belongs within: Raphignathoidea.

Stigmaeus is a genus of predatory mites with ten to sixteen dorsal shields on the idiosoma, and four pairs of setae on the prodorsal shield (Khanjani & Ueckermann 2002).

Characters (from Khanjani & Ueckermann 2002): Idiosoma with 10–16 dorsal shields, dimpled or reticulated in most species; prodorsum with four pairs of setae on single large or on a large median and a pair of lateral shields; opisthosomal shielding consisting of an unpaired median shield bearing two or three pairs of setae and unpaired or paired suranal shield and three or six pairs of smaller shields (humerals, marginal, median zonals, lateral zonals and intercalaries) surrounding median shield. Median shield distinct, indistinct or absent. Dorsum with 13–14 pairs of setae.

<==Stigmaeus Koch 1836 KU02
    |--*S. cruentus Koch 1836 KU02
    |--S. africanus E-KO07
    |--S. alvandis Khanjani & Ueckermann 2002 KU02
    |--S. antrodes Berlese 1910 SB63 [incl. S. antrodes var. reticulatus Halb. 1923 G44]
    |--S. bdelloides SS55
    |--S. bicolor Canestrini & Fanzago 1876 CF77
    |--S. elongatus Berlese 1886 [incl. S. luteus Summers 1962] KU02
    |--S. eutrichus Berlese 1910 SB63
    |--S. fissuricola Halbert 1920 [=S. rhodomelas var. fissuricola] S76
    |--S. raynei WHE88
    |--S. rhodomelas Berl. 1910 G44
    |--S. sinai WL09
    |--S. spagneti WHE88
    |--S. unicus Kuznetzov 1977 KU02
    `--S. zaheri E-KO07

*Type species of generic name indicated

REFERENCES

[CF77] Canestrini, G., & F. Fanzago. 1877. Intorno agli Acari Italiani. Atti del Reale Istituto Veneto di Scienze, Lettere ed Arti, Serie 5, 4: 69–208, pls 2–7.

[E-KO07] El-Kammah, K. M., & L. M. I. Oyoun. 2007. Taxonomic records of mite species associated with animal and poultry farms in the Nile delta, vally and N. Sinai Egypt. In: Morales-Malacara, J. B., V. M. Behan-Pelletier, E. Ueckermann, T. M. Pérez, E. G. Estrada-Venegas & M. Badii (eds) Acarology XI: Proceedings of the International Congress pp. 223–232. Instituto de Biología and Faculdad de Ciencias, Universidad Nacional Autónoma de México, Sociedad Latinoamericana de Acarología: México.

[G44] Grandjean, F. 1944. Observations sur les acariens de la famille des Stigmaeidae. Archives des Sciences Physiques et Naturelles, 5me Période 26: 103–131.

[KU02] Khanjani, M., & E. A. Ueckermann. 2002. The stigmaeid mites of Iran (Acari: Stigmaeidae). International Journal of Acarology 28 (4): 317–339.

[SB63] Schweizer, J., & C. Bader. 1963. Die Landmilben der Schweiz (Mittelland, Jura und Alpen): Trombidiformes Reuter, mit 217 Arten und Unterarten und 193 Originalzeichnungen. Denkschriften der Schweizerischen Naturforschenden Gesellschaft [Mémoires de la Société Helvétique des Sciences Naturelles] 84 (2): i–vi, 209–378.

[SS55] Summers, F. M., & E. I. Schlinger. 1955. Mites of the family Caligonellidae (Acarina). Hilgardia 23 (12): 539–561.

[WHE88] Walter, D. E., H. W. Hunt & E. T. Elliott. 1988. Guilds or functional groups? An analysis of predatory arthropods from a shortgrass steppe soil. Pedobiologia 31: 247–260.

[WL09] Walter, D. E., E. E. Lindquist, I. M. Smith, D. R. Cook & G. W. Krantz. 2009. Order Trombidiformes. In: Krantz, G. W., & D. E. Walter (eds) A Manual of Acarology 3rd ed. pp. 233–420. Texas Tech University Press.

Tetranychus

Two-spotted spider mite Tetranychus urticae, copyright J. Holopainen.


Belongs within: Tetranychinae.

Tetranychus is a genus of spider mites with the duplex setae on tarsus I widely separated and the empodium of the female and empodia III and IV of the male consisting of three pairs of hairs with or without a proximodorsal claw (Baker & Pritchard 1960).

<==Tetranychus Dufour 1832 CF77
    |--T. gloveri NG96
    |--T. neocaledonicus (André 1933) NG96, H98 (see below for synonymy)
    `--+--T. (Tetranychus) BT72
       |    |--*T. (T.) lintearius Dufour 1832 BT72, G48 [incl. T. russeolus CF77, T. ulmi CF77, T. vestitus CF77]
       |    |--T. (T.) cinnabarinus (Boisduval 1867) AP80, BP60 (see below for synonymy)
       |    |--T. (T.) ludeni Zacher 1913 AP80, H98
       |    |--T. (T.) mexicanus AP80
       |    |--T. (T.) nakahari AP80
       |    |--T. (T.) urticae Koch 1836 NG96, BT72 (see below for synonymy)
       |    `--T. (T.) yusti AP80
       `--+--T. kanzawai Kishida 1927 NG96, H98 [incl. T. hydrangeae Pritchard & Baker 1955 WL09, H98]
          `--+--T. pacificus NG96
             `--T. (Armenychus Wainstein 1960) BT72
                  |--T. (*A.) armeniaca BT72
                  |--T. (A.) cobrensus Baker & Tuttle 1972 BT72
                  `--T. (A.) mcdanieli McGregor 1931 NG96, BT72

Tetranychus incertae sedis:
  T. abacae MF07
  T. andrei Baker & Pritchard 1960 BP60
  T. armipennis W99
  T. bastosi S91
  T. browningi Baker & Pritchard 1960 BP60
  T. canadensis NMF07
  T. crataegi Hirst 1920 H23
  T. desertorum Banks 1900 [incl. T. opuntiae Banks 1908] H98
  T. dianellae Davis 1967 H98
  T. dugesi Cano & Alsacio 1886 Hof07
  T. evansi Baker & Pritchard 1960 BP60
  T. frater NMF07
  T. gibbus Koch & Berendt 1854 S93
  T. lambi Pritchard & Baker 1955 H98
  T. latus CF77
  T. lombardinii Baker & Pritchard 1960 BP60
  T. macfarlanei Baker & Pritchard 1960 BP60
  T. marianae McGregor 1950 H98
  T. ogmophallos MF07
  T. papayae G01
  T. piercei Ho07
  T. pilosus CF77
  T. pueraricola WL09
  T. quercivorus NG96
  T. rhagodiae Miller 1966 H98
  T. savenkoae NG96
  T. sayedi Baker & Pritchard 1960 BP60
  T. schoenei NMF07
  T. spinigerus Lucas 1847 (n. d.) E12 [=Metatetranychus spinigerus BP60]
  T. truncatus NMF07
  T. tumidosus Baker & Pritchard 1960 BP60
  T. turkestani Ugarov & Nikolski 1937 KB91 [incl. T. atlanticus WL09]

Tetranychus neocaledonicus (André 1933) NG96, H98 [=Eotetranychus neocaledonicus H98; incl. T. cucurbitae Rahman & Sapra 1958 BP60]

Tetranychus (Tetranychus) cinnabarinus (Boisduval 1867) AP80, BP60 [=Acarus cinnabarinus BP60; incl. T. cucurbitacearum Sayed 1946 BP60]

Tetranychus (Tetranychus) urticae Koch 1836 NG96, BT72 [incl. T. arabicus K07, Epitetranychus alceae Linnaeus 1758 S61, T. altheae S61, Acarus cucumeris Boisduval 1867 H98, T. cucumeris H98, A. rosarum Boisduval 1867 H98, T. rosarum H98, A. sambuci Schranck 1781 S61, A. telarius Linnaeus 1758 (nom. rej.) H98, Eotetranychus telarius SB63, Tetranychus telarius BP60]

*Type species of generic name indicated

REFERENCES

[AP80] Andrews, K. L., & S. L. Poe. 1980. Spider mites of El Salvador, Central America (Acari: Tetranychidae). Florida Entomologist 63 (4): 502–505.

[BP60] Baker, E. W., & A. E. Pritchard. 1960. The tetranychoid mites of Africa. Hilgardia 29 (11): 455–574.

[BT72] Baker, E. W., & D. M. Tuttle. 1972. New species and further notes on the Tetranychoidea mostly from the southwestern United States (Acarina: Tetranychidae and Tenuipalpidae). Smithsonian Contributions to Zoology 116: 1–37.

[CF77] Canestrini, G., & F. Fanzago. 1877. Intorno agli Acari Italiani. Atti del Reale Istituto Veneto di Scienze, Lettere ed Arti, Serie 5, 4: 69–208, pls 2–7.

[E12] Evenhuis, N. L. 2012. Publication and dating of the Exploration Scientifique de l'Algérie: Histoire Naturelle des Animaux Articulés (1846–1849) by Pierre Hippolyte Lucas. Zootaxa 3448: 1–61.

[G48] Grandjean, F. 1948. Quelques caractères des tétranyques. Bulletin du Muséum, 2e série 20 (6): 517–524.

[G01] Gupta, S. K. 2001. A conspectus of natural enemies of phytophagous mites and mites as potential biocontrol agents of agricultural pests in India. In: Halliday, R. B., D. E. Walter, H. C. Proctor, R. A. Norton & M. J. Colloff (eds) Acarology: Proceedings of the 10th International Congress pp. 484–497. CSIRO Publishing: Melbourne.

[H98] Halliday, R. B. 1998. Mites of Australia: A checklist and bibliography. CSIRO Publishing: Collingwood.

[H23] Hirst, S. 1923. On some new or little-known species of Acari. Proceedings of the Zoological Society of London 1923: 971–1000.

[Ho07] Ho, C.-C. 2007. Crop mite pests of southeast Asia and tropical far east. In: Morales-Malacara, J. B., V. M. Behan-Pelletier, E. Ueckermann, T. M. Pérez, E. G. Estrada-Venegas & M. Badii (eds) Acarology XI: Proceedings of the International Congress pp. 259–266. Instituto de Biología and Faculdad de Ciencias, Universidad Nacional Autónoma de México, Sociedad Latinoamericana de Acarología: México.

[Hof07] Hoffmann, A. 2007. Acarology in Mexico: prehispanic to modern times. In: Morales-Malacara, J. B., V. M. Behan-Pelletier, E. Ueckermann, T. M. Pérez, E. G. Estrada-Venegas & M. Badii (eds) Acarology XI: Proceedings of the International Congress pp. 21–26. Instituto de Biología and Faculdad de Ciencias, Universidad Nacional Autónoma de México, Sociedad Latinoamericana de Acarología: México.

[K07] Knapp, M. 2007. Important mite crop pests in Africa. In: Morales-Malacara, J. B., V. M. Behan-Pelletier, E. Ueckermann, T. M. Pérez, E. G. Estrada-Venegas & M. Badii (eds) Acarology XI: Proceedings of the International Congress pp. 267–277. Instituto de Biología and Faculdad de Ciencias, Universidad Nacional Autónoma de México, Sociedad Latinoamericana de Acarología: México.

[KB91] Kreiter, S., F. Brian, C. Magnien, G. Sentenac & G. Valentin. 1991. Spider mites and chemical control of weeds: interactions. In Dusbábek, F. & V. Bukva (eds) Modern Acarology: Proceedings of the VIII International Congress of Acarology, held in České Budĕjovice, Czechoslovakia, 6–11 August 1990 vol. 2 pp. 725–736. SPB Academic Publishing: The Hague.

[MF07] Moraes, G. J. de, & C. H. W. Flechtmann. 2007. Phytophagous mites of tropical crops in eastern South America. In: Morales-Malacara, J. B., V. M. Behan-Pelletier, E. Ueckermann, T. M. Pérez, E. G. Estrada-Venegas & M. Badii (eds) Acarology XI: Proceedings of the International Congress pp. 279–288. Instituto de Biología and Faculdad de Ciencias, Universidad Nacional Autónoma de México, Sociedad Latinoamericana de Acarología: México.

[NG96] Navajas, M., & J. Gutierrez. 1996. Molecular phylogeny in spider mites (Tetranychidae). In: Mitchell, R., D. J. Horn, G. R. Needham & W. C. Welbourn (eds) Acarology IX vol. 1. Proceedings pp. 305–308. Ohio Biological Survey: Columbus (Ohio).

[NMF07] Navia, D., G. J. de Moraes & C. H. W. Flechtmann. 2007. Phytophagous mites as invasive alien species: quarantine procedures. In: Morales-Malacara, J. B., V. M. Behan-Pelletier, E. Ueckermann, T. M. Pérez, E. G. Estrada-Venegas & M. Badii (eds) Acarology XI: Proceedings of the International Congress pp. 307–316. Instituto de Biología and Faculdad de Ciencias, Universidad Nacional Autónoma de México, Sociedad Latinoamericana de Acarología: México.

[S91] Sabelis, M. W. 1991. Life-history evolution of spider mites. In: Schuster, R., & P. W. Murphy (eds) The Acari: Reproduction, development and life-history strategies pp. 23–49. Chapman & Hall: London.

[SB63] Schweizer, J., & C. Bader. 1963. Die Landmilben der Schweiz (Mittelland, Jura und Alpen): Trombidiformes Reuter, mit 217 Arten und Unterarten und 193 Originalzeichnungen. Denkschriften der Schweizerischen Naturforschenden Gesellschaft [Mémoires de la Société Helvétique des Sciences Naturelles] 84 (2): i–vi, 209–378.

[S93] Selden, P. A. 1993. Arthropoda (Aglaspidida, Pycnogonida and Chelicerata). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 297–320. Chapman & Hall: London.

[S61] Southcott, R. V. 1961. Studies on the systematics and biology of the Erythraeoidea (Acarina), with a critical revision of the genera and subfamilies. Australian Journal of Zoology 9: 367–610.

[W99] Walter, D. E. 1999. Cryptic inhabitants of a noxious weed: mites (Arachnida: Acari) on Lantana camara L. invading forests in Queensland. Australian Journal of Entomology 38: 197–200.

[WL09] Walter, D. E., E. E. Lindquist, I. M. Smith, D. R. Cook & G. W. Krantz. 2009. Order Trombidiformes. In: Krantz, G. W., & D. E. Walter (eds) A Manual of Acarology 3rd ed. pp. 233–420. Texas Tech University Press.

Ecpleopini

Leposoma nanodactylus, from the Reptile Database.


Belongs within: Gymnophthalmidae.

The Ecpleopini are a clade of microteiid lizards supported by molecular data (Pellegrino et al. 2001).

<==Ecpleopini
    |--Arthrosaura Boulenger 1885 PR01
    |    |--A. kochii PR01
    |    `--A. reticulata PR01
    `--+--Ecpleopus Duméril & Bibron 1839 PR01
       |    `--*E. gaudichaudii PR01
       `--+--+--Colobosauroides Cunha & Lima Verde 1991 PR01
          |  |    `--C. cearensis PR01
          |  `--Anotosaura Amaral 1933 PR01
          |       |--A. collaris PR01
          |       `--A. vanzolinia PR01
          `--+--Amapasaurus Cunha 1970 PR01
             `--Leposoma Spix 1825 PR01
                  |--L. nanodactylus PR01
                  |--L. oswaldoi PR01
                  |--L. parietale DS86
                  `--L. percarinatum PR01

*Type species of generic name indicated

REFERENCES

[DS86] Dixon, J. R,. & P. Soini. 1986. The Reptiles of the Upper Amazon Basin, Iquitos Region, Peru. Milwaukee Public Museum: Milwaukee.

[PR01] Pellegrino, K. C. M., M. T. Rodrigues, Y. Yonenaga-Yassuda & J. W. Sites Jr. 2001. A molecular perspective on the evolution of microteiid lizards (Squamata, Gymnophthalmidae), and a new classification for the family. Biological Journal of the Linnean Society 74: 315–338.

Cheyletus

Ventral view of Cheyletus eruditus, photographed by J. Hubert.


Belongs within: Cheyletini.

Cheyletus is a genus of free-living predatory mites. One species, C. eruditus, has been used as a biological control agent for grain mites.

Characters (from Baker 1949): Eyes absent; palpal tarsus with two sickle-like and two comb-like setae; two dorsal shields present; dorsal marginal setae feather-like or pilose; dorsal setae, when present, staghorn-like; leg I with two claws.

<==Cheyletus Latreille 1796 BF01 [=Cheletes B49; incl. Eutarsus Hessling 1852 B49]
    |--*C. eruditus (Schrank 1781) (see below for synonymy) B49
    |--C. acer Oudemans 1904 [=Cheletes acer] B49
    |--C. alacer Oudemans 1904 [=Cheletes alacer] B49
    |--C. audax Oudemans 1904 [=Cheletes audax] B49
    |--C. aversor Rodendorf 1940 B49
    |--C. baloghi BF01
    |--C. beauchampi Baker 1949 B49
    |--C. burmiticus Cockerell 1917 (n. d.) B49
    |--C. butleri Hughes 1948 B49
    |--C. cacahuamilpensis Baker 1949 B49
    |--C. carnifex Zakhvatkin 1940 B49
    |--C. clavispinus Banks 1902 (n. d.) B49
    |--C. cocos H07
    |--C. davisi Baker 1949 B49
    |--C. doddi Baker 1949 B49
    |--C. ferox Trouessart 1889 B49
    |--C. fortis Oudemans 1904 [=Cheletes fortis] B49
    |--C. furibundus Rodendorf 1940 B49
    |--C. hendersoni Baker 1949 B49
    |--C. intrepidus Oudemans 1903 [=Cheletes intrepidus] B49
    |--C. linsdalei Baker 1949 B49
    |--C. malaccensis Oudemans 1903 [=Cheletes malaccensis] H98
    |--C. munroi Hughes 1948 B49
    |--C. parumsetosus Karpelles 1884 B49
    |--C. patagiatus Nordenskiold 1900 (n. d.) B49
    |--C. praedabundus Kuzin in Rodendorf 1940 B49
    |--C. promptus Oudemans 1904 B49
    |--C. rabiosus Rodendorf 1940 B49
    |--C. rapax Oudemans 1903 [=Cheletes rapax] B49
    |--C. saevus Oudemans 1906 (n. d.) [=Cheletes saevus] B49
    |--C. schneideri Oudemans 1904 [=Cheletes schneideri] B49
    |--C. strenuus Oudemans 1904 [=Cheletes strenuus] B49
    |--C. tenuipilis Fain et al. 1980 MR01
    |--C. trouessarti Oudemans 1902 H98 [=Cheletes trouessarti H98; incl. Cheyletus macrocherus Hardy in Andre 1933 B49]
    |--C. trux Rodendorf 1940 B49
    |--C. venator Vitzthum 1920 B49
    `--C. vorax Oudemans 1903 [=Cheltes vorax] B49

*Cheyletus eruditus (Schrank 1781) [=Acarus eruditus; incl. *Eutarsus cancriformis Hessling 1852, Cheyletus eburneus Hardy in Andre 1933, C. ferox Banks 1906 non Trouessart 1889] B49

*Type species of generic name indicated

REFERENCES

[B49] Baker, E. W. 1949. A review of the mites of the family Cheyletidae in the United States National Museum. Proceedings of the United States National Museum 99 (3238): 267–320.

[BF01] Bochkov, A. V., & A. Fain. 2001. Phylogeny and system of the Cheyletidae (Acari: Prostigmata) with special reference to their host-parasite associations. Bulletin de l’Institut Royal des Sciences Naturelles de Belgique Entomologie 71: 5–36.

[H98] Halliday, R. B. 1998. Mites of Australia: A checklist and bibliography. CSIRO Publishing: Collingwood.

[H07] Haq, M. A. 2007. Coconut mite tragedy in peninsular India. In: Morales-Malacara, J. B., V. M. Behan-Pelletier, E. Ueckermann, T. M. Pérez, E. G. Estrada-Venegas & M. Badii (eds) Acarology XI: Proceedings of the International Congress pp. 427–432. Instituto de Biología and Faculdad de Ciencias, Universidad Nacional Autónoma de México, Sociedad Latinoamericana de Acarología: México.

[MR01] Morris, M. J. A., & J. Rimmer. 2001. Allergenicity of the predator dust mite Cheyletus tenuipilis (Acari: Cheyletidae): a preliminary study. In: Halliday, R. B., D. E. Walter, H. C. Proctor, R. A. Norton & M. J. Colloff (eds) Acarology: Proceedings of the 10th International Congress pp. 558–564. CSIRO Publishing: Melbourne.

Diaspididae

Coconut scales Aspidiotus destructor, photographed by Scot Nelson.


Belongs within: Coccoidea.
Contains: Parlatoriini.

The Diaspididae, armoured scales, are a diverse group of scale insects covered by a wax-like, fibrous scale formed from pygidial secretions, anal excreta and the exuviae of previous instars (Carver et al. 1991).

Characters (from Carver et al. 1991): Adult female legless, without antennae; fourth and following segments of abdomen usually fused to form pygidium, marginally bearing complex of tubular ducts, marginal spines, plates and gland spines; operculum absent. Scale incorporating exuviae of first and second instars, adult female sometimes remaining enclosed within exuviae of second instar.

<==Diaspididae T02
    |--Ulucoccus Takagi, Tho & Khoo 1990 [Ulucoccinae] T02
    |    |--U. danumensis T02
    |    `--U. gombakensis T02
    |--Diaspidinae T02
    |    |--Lepidosaphidini [Lepidosaphedini] T02
    |    |    |  i. s.: Mitulaspis malayana T02
    |    |    |--Coccomytilina T02
    |    |    |--Howardiina T02
    |    |    |--Mempelaspis [Mempelaspidina] T02
    |    |    |    `--M. serpentina Takagi 2000 T02
    |    |    `--Lepidosaphes M27 [Lepidosaphidina T02]
    |    |         |--L. beckii ACW01
    |    |         |--L. cordylinidis M27
    |    |         `--L. ulmi WP99
    |    `--Diaspidini T02
    |         |--Kuwanaspidina T02
    |         |--Protodiaspidina T02
    |         |--Fiorinia [Fioriniina] T02
    |         |    `--F. (Adiscofiorinia) pygosema Green & Laing 1923 [=Mongrovaspis pygosema] T02
    |         |--Diaspis G26 [Diaspidina T02]
    |         |    `--D. echinocacti G26
    |         `--Chionaspis G33 [Chionaspidina T02]
    |              |--C. citri G33
    |              `--C. eucalypti G29
    `--Aspidiotinae T02
         |--Parlatoriini T02
         |--Odonaspis [Odonaspidini] T02
         |    `--O. arcusnotata Ben-Dov 1988 T02
         |--Thysanaspis Ferris 1955 [Thysanaspidini] T02
         |    |--*T. acalyptus Ferris 1955 T02
         |    |--T. litseae Takagi 1961 T02
         |    `--T. perkinsi Takagi 1969 T02
         |--Smilacicola Takagi 1969 [Smilacicolini] T02
         |    |--*S. apicalis Takagi 1969 T02
         |    |--S. crenatus Takagi 1983 T02
         |    `--S. heimi (Balachowsky 1947) [=Rugaspidiotus heimi] T02
         |--Leucaspidini T02
         |    |--Leucaspis cordylinidis M27
         |    |--Lopholeucaspis japonica (Cockerell 1897) T02
         |    `--Mongrovaspis Bodenheimer 1951 T02
         |         `--*M. quadrispinosa (Green 1934) [=Leucaspis quadrispinosa] T02
         `--Aspidiotus Bouché 1833 K00 [Aspidiotini T02]
              |--A. destructor S07
              |--A. hederae [incl. A. buddleiae, A. carpodeti, A. epidendri] M27
              |--A. juglansregiae Comstock 1880 S69
              |--A. nerii S07
              |--A. proteus Pampaloni 1902 K00
              |--A. secretus G17
              |--A. uvae B49
              `--A. vitiensis K08

Diaspididae incertae sedis:
  Lapazia Ferris 1937 OK06
  Pinnaspis strachani MHG04
  Aonidiella GE05
    |--A. aurantii C81 [=Aspidiotus aurantii D37]
    `--A. citrina WD04
  Xanthophthalma T02
  Salicicola T02
    |--S. kermanensis (Lindinger 1906) T02
    `--S. vayssierei Balachowsky 1928 T02
  Melanaspis KU07
    |--M. glomerata G01
    `--M. inopinata KU07
  Pseudaulacaspis pentagona KU07
  Chrysomphalus CGW91
    |--C. aonidum CGW91
    |--C. aurantii K08
    |--C. dictyospermi [incl. C. dictyospermi var. pinnulifera] K08
    `--C. ficus WEE70
  Unaspis citri S07
  Quadraspidiotus CGW91
    |--Q. lenticularis CGW91
    |--Q. ostreaeformis CGW91
    |--Q. perniciosus CGW91
    `--Q. pyri WEE70
  Berlesaspis spinifera CGW91
  Maskellia globosa CGW91

*Type species of generic name indicated

REFERENCES

[ACW01] Aguilar, H., C. C. Childers & W. C. Welbourn. 2001. Relative abundance and seasonal occurrence of mites in the family Tydeidae on citrus in Florida. In: Halliday, R. B., D. E. Walter, H. C. Proctor, R. A. Norton & M. J. Colloff (eds) Acarology: Proceedings of the 10th International Congress pp. 376–380. CSIRO Publishing: Melbourne.

[B49] Baker, E. W. 1949. A review of the mites of the family Cheyletidae in the United States National Museum. Proceedings of the United States National Museum 99 (3238): 267–320.

[C81] Caltagirone, L. E. 1981. Landmark examples in classical biological control. Annual Review of Entomology 26: 213–232.

[CGW91] Carver, M., G. F. Gross & T. E. Woodward. 1991. Hemiptera (bugs, leafhoppers, cicadas, aphids, scale insects etc.) In: CSIRO. The Insects of Australia: A textbook for students and research workers vol. 1 pp. 429–509. Melbourne University Press: Carlton (Victoria).

[D37] Dobzhansky, T. 1937. Genetics and the Origin of Species. Columbia University Press: New York.

[G17] Girault, A. A. 1917. Descriptiones stellarum novarum. Privately published (reprinted Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 80–101).

[G26] Girault, A. A. 1926. New pests from Australia. II. Privately published (reprinted Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 200–202).

[G29] Girault, A. A. 1929. Description of a case of lunacy in Homo and of six-legged articulates. Privately published (reprinted Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 262–265).

[G33] Girault, A. A. 1933. Some beauties inhabitant not of commercial boudoirs but of Nature’s bosom, notably new insects. Privately published (reprinted Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 299–303).

[GE05] Grimaldi, D., & M. S. Engel. 2005. Evolution of the Insects. Cambridge University Press: New York.

[G01] Gupta, S. K. 2001. A conspectus of natural enemies of phytophagous mites and mites as potential biocontrol agents of agricultural pests in India. In: Halliday, R. B., D. E. Walter, H. C. Proctor, R. A. Norton & M. J. Colloff (eds) Acarology: Proceedings of the 10th International Congress pp. 484–497. CSIRO Publishing: Melbourne.

[KU07] Khanjani, M., & E. A. Ueckermann. 2007. Natural enemies of the walnut pest Aceria tristriatus (Nal.) (Acari: Eriophyidae) in western Iran. In: Morales-Malacara, J. B., V. M. Behan-Pelletier, E. Ueckermann, T. M. Pérez, E. G. Estrada-Venegas & M. Badii (eds) Acarology XI: Proceedings of the International Congress pp. 451–453. Instituto de Biología and Faculdad de Ciencias, Universidad Nacional Autónoma de México, Sociedad Latinoamericana de Acarología: México.

[K08] Kirkaldy, G. W. 1908. A catalogue of the Hemiptera of Fiji. Proceedings of the Linnean Society of New South Wales 33: 345–391, pl. 4.

[K00] Koteja, J. 2000. Advances in the study of fossil coccids (Hemiptera: Coccinea). Polskie Pismo Entomologiczne 69: 187–218.

[MHG04] Maynard, G. V., J. G. Hamilton & J. F. Grimshaw. 2004. Quarantine—phytosanitary, sanitary and incursion management: an Australian entomological perspective. Australian Journal of Entomology 43 (3): 318–328.

[M27] Myers, J. G. 1927. On the nomenclature of New Zealand Homoptera. Transactions and Proceedings of the New Zealand Institute 57: 685–690.

[OK06] Özdikmen, H., & A. B. Kury. 2006. Three homonymous generic names in Araneae and Opiliones. Journal of Arachnology 34 (1): 279–280.

[S07] Ślipiński, A. 2007. Australian Ladybird Beetles (Coleoptera: Coccinellidae): Their biology and classification. Australian Biological Resources Study: Canberra.

[S69] Steyskal, G. C. 1969. The mistreatment of the Latin genitive case in forming names of parasites. Systematic Zoology 18 (3): 339–342.

[T02] Takagi, S. 2002. One new subfamily and two new tribes of the Diaspididae (Homoptera: Coccoidea). Insecta Matsumurana, new series 59: 55–100.

[WP99] Walter, D. E., & H. C. Proctor. 1999. Mites: Ecology, Evolution and Behaviour. CABI Publishing: Wallingford (UK).

[WD04] Wardhaugh, C. W., & R. K. Didham. 2004. The effect of introduced wasp (Vespula vulgaris, Hymenoptera: Vespidae) predation on the dispersive life history stages of beech scale insects (Ultracoelostoma spp., Homoptera: Margarodidae). New Zealand Entomologist 27: 91–101.

[WEE70] Woodward, T. E., J. W. Evans & V. F. Eastop. 1970. Hemiptera (bugs, leafhoppers, etc.) In: CSIRO. The Insects of Australia: A textbook for students and research workers pp. 387–457. Melbourne University Press.