Sphaeromatidea

Male Serolis bromleyana, from Beddard (1884).

Belongs within: Scuticoxifera.
Contains: Sphaeromatidae.

The Sphaeromatidea are a clade of marine isopods with the underside of the pleotelson vaulted, defining a branchial chamber (Brandt & Poore 2003).

Characters (from Brandt & Poore 2003): Pleonites variously fused (rarely 1–5 free), pleonite 1 narrower than others (similar in Plakarthrium). Pleotelson underside vaulted, branchial chamber defined by ridges along mesial margin of lateral edge. Antenna 1 without scale. Mandibular lacinia mobilis present on left side, reduced and fused with spine row on right or absent. Mandibular molar a cylindrical process with triturative flat end. Maxillipedal endite reaching at least distal margin of palp article 2, usually distally truncate and setose. Pleopod 3 endopod roughly triangular or at least with inner angle sharper than outer. Dorsal coxal plate 7 similar to others (rarely shorter). Uropodal rami lateral to margin of pleotelson, articulating along longitudinal axis and folding down alongside branchial space.

<==Sphaeromatidea BP03
    |--Sphaeromatoidea BP03
    |    |--Paravireia Chilton 1925 BP03
    |    |    `--P. holdichi Brökeland, Wägele & Bruce 2001 BP03
    |    `--+--Sphaeromatidae BP03
    |       `--+--Tecticeps Richardson 1897 [Tecticipitidae] BP03
    |          |    |--T. alascensis H47
    |          |    |--T. convexus H47
    |          |    |--T. marginalis Gurjanova 1935 BP03
    |          |    `--T. pugettensis Hatch 1947 H47
    |          `--Ancinidae BP03
    |               |--Bathycopea Tattersall 1905 BP03
    |               |    `--B. typhlops Tattersall 1905 BP03
    |               `--Ancinus Milne Edwards 1840 BP03
    |                    |--A. belizensis Kensley & Schotte 1987 BP03
    |                    `--A. depressus (Say 1818) MK83
    `--Seroloidea BP03
         |--Plakarthrium Chilton 1883 [Plakarthriidae] BP03
         |    `--P. australiense Poore & Brandt 2001 BP03
         `--+--Schweglerella [Schweglerellidae] BP03
            |    `--S. strobli Polz 1998 BP03
            `--+--Bathynataliidae BP02
               |    |--Biremia Bruce 1985 BP03
               |    |    `--B. ambocerca Bruce 1985 BP03
               |    `--+--Naudea Kensley 1979 BP03
               |       |    `--N. louwae Kensley 1979 BP03
               |       `--Bathynatalia Kensley 1978 BP03
               |            `--B. gilchristi Barnard 1957 BP03
               `--+--Basserolis Poore 1985 [Basserolidae] BP03
                  |    `--B. kimblae Poore 1985 BP03
                  `--Serolidae BP03
                       |--Serolina holia Poore 1987 BP03
                       `--Serolis Leach 1818 BP03
                            |--*S. paradoxus (Fabricius 1775) [=Oniscus paradoxus; incl. S. fabricii Leach 1818] MK83
                            |--S. bouvieri Richardson 1909 BP03
                            |--S. bromleyana BH88
                            |--S. carinata Lockington 1877 MK83
                            `--S. mgrayi Menzies & Frankenberg 1966 MK83

*Type species of generic name indicated

REFERENCES

[BH88] Boxshall, G. A., & K. Harrison. 1988. New nicothoid copepods (Copepoda: Siphonostomatoida) from an amphipod and from deep-sea isopods. Bulletin of the British Museum (Natural History), Zoology series 54 (6): 285–299.

[BP03] Brandt, A. & G. C. B. Poore. 2003. Higher classification of the flabelliferan and related Isopoda based on a reappraisal of relationships. Invertebrate Systematics 17: 893–923.

[H47] Hatch, M. H. 1947. The Chelifera and Isopoda of Washington and adjacent regions. University of Washington Publications in Biology 10 (5): 155–274.

[MK83] Menzies, R. J., & W. L. Kruczynski. 1983. Isopod Crustacea (exclusive of Epicaridea). Memoirs of the Hourglass Cruises 6 (1): 1–126.

Leptocheliidae

Leptochelia sp., copyright Magdalena Błażewicz-Paszkowycz.

Belongs within: Tanaidacea.

The Leptocheliidae are a family of tanaidaceans characterised by a relatively high number of articles in the uropod endopod (Larsen & Wilson 2002).

Characters (from Larsen & Wilson 2002): Female with medium body calcification. No plates in carapace. Eyes present (except in Mesotanais). Pereonite 1 not reduced. Pleon with five free pleonites, as wide as pereon, no articulated setae on pleonites. Antennule with three to five articles. Antenna with six articles, article 3 with or without (Heterotanaoides) dorsal spiniform setae. Molar process broad. Labium with four lobes, medial spiniform setae absent. Maxilliped basis and endites not fused, endites narrower than basis and with two, or (frequently) three flat setae. Cheliped attachment via sclerite. Marsupium (where known) with four pairs of oostegites. Pereopod coxa present on P1–3 but absent on P4–6. P4–6 dactylus and terminal setae fused to a hook. Pleopods well developed. Uropod endopod with three to six articles, exopod with one or two articles. Male (where known) with body shorter than female. Eyes larger than females. Antennule with more than four articles and aesthetascs. Mouthparts reduced. Cheliped always significantly enlarged and elongated. Pleopods well developed.

<==Leptocheliidae
    |--Heterotanaidinae LW02
    |    |--Heterotanaoides Sieg 1976 LW02
    |    |    `--H. meridionalis Sieg 1986 LW02
    |    |--Mesotanais Dollfus 1897 LW02
    |    |    `--M. dubius Dollfus 1897 LW02
    |    `--Heterotanais Sars 1882 LW02
    |         |--H. melacephala H47
    |         `--H. oerstedii (Kroyer 1842) LW02
    `--Leptocheliinae LW02
         |--‘Hargeria’ Lang 1973 non Lucas 1903 LW02
         |    `--H. rapax (Harger 1879) LW02
         |--Intermedichelia Gutu 1996 LW02
         |    `--I. gracilis Gutu 1996 LW02
         |--Pseudoleptochelia Lang 1973 LW02
         |    `--P. antarctica (Lang 1953) LW02
         |--Pseudonototanais Lang 1973 LW02
         |    `--P. werthi (VanHoeffen 1914) LW02
         `--Leptochelia Dana 1852 LW02
              |--L. dubia (Kroyer 1842) LW02
              `--L. filum H47

*Type species of generic name indicated

REFERENCES

[H47] Hatch, M. H. 1947. The Chelifera and Isopoda of Washington and adjacent regions. University of Washington Publications in Biology 10 (5): 155–274.

[LW02] Larsen, K., & G. D. F. Wilson. 2002. Tanaidacean phylogeny, the first step: the superfamily Paratanaidoidea. Journal of Zoological Systematics and Evolutionary Research 40: 205–222.

Cymothoida

Parasitic isopod Anilocra physodes, copyright Txomin Rivera.

Belongs within: Scuticoxifera.
Contains: Cirolaninae, Eurydicinae, Gnathiidae, Anthuroidea, Corallanidae, Aegidae, Epicaridea, Lironeca, Nerocila, Cymothoa.

The Cymothoida is a diverse group of mostly marine isopods, including both free-living and parasitic forms.

Characters (from Brandt & Poore 2003): Pleonites 1–5 free, pleonite 1 similar to others (some pleonites fused in some species). Pleotelson underside flat, without ventrolateral ridges; pleopods not enclosed laterally. Antenna 1 without scale (except in Bathynomus). Mandibular lacinia mobilis absent (present in some Anthuroidea as part of lacinia mobilis). Mandibular molar flat blade-like or conical process (sometimes absent). Maxillipedal endite, if present, rarely longer than palp article 1, distally tapering and with few setae. Dorsal coxal plate 7 similar to others. Pleopods 3–5 endopods more or less oval or rectangular, rounded dorsally. Uropodal rami ventral to pleotelson, articulating from side to side along vertical axis inside branchial spine.

<==Cymothoida BP03
    |--Cirolanidae [Cirolanoidea] BP03
    |    |  i. s.: Ceratolana BBN86
    |    |         Kensleylana Bruce & Herrando-Pérez 2005 F05
    |    |           `--*K. briani Bruce & Herrando-Pérez 2005 F05
    |    |--Cirolaninae BBN86
    |    |--Eurydicinae BBN86
    |    `--+--Conilorpheus BBN86
    |       |--Orphelana BBN86
    |       |--Politolana concharum BBN86, M02
    |       |--Conilera Leach 1818 BBN86, T08
    |       |    |--C. cylindracea Montagu 1804 T08
    |       |    `--C. stygia Packard 1894 BBN86
    |       `--Natatolana Bruce 1986 BBN86, BP03
    |            |--N. albinota BP03
    |            |--N. caeca BBN86
    |            |--N. californiensis BBN86
    |            `--N. woodjonesi (Hale 1924) BP03
    `--Cymothooidea BP03
         |--Anuropus Beddard 1886 [Anuropidae] BP03
         |    |--A. kussakini Vasina 1998 BP03
         |    `--A. pacificus Lincoln & Jones 1973 BP03
         `--+--Protognathia Wägele & Brandt 1988 [Protognathiidae] BP03
            |    |--P. bathypelagica Wägele & Brandt 1988 BP03
            |    `--P. waegeli Kussakin & Rybakov 1995 BP03
            `--+--+--Gnathiidae BP03
               |  `--Anthuroidea BP03
               `--+--Corallanidae BP03
                  `--+--Tridentella Richardson 1905 [Tridentellidae] BP03
                     |    |--T. benguela Brandt & Poore 2001 BP03
                     |    `--T. namibia Brandt & Poore 2001 BP03
                     `--+--Aegidae BP03
                        `--+--Epicaridea BP03
                           `--Cymothoidae BP03
                                |--Lironeca B78
                                |--Aegathoa MK83
                                |--Olencira praegustator (Latrobe 1802) MK83
                                |--Agarna cumulus (Haller 1880) [incl. A. carinata Schiödte & Meinert 1883] MK83
                                |--Telotha henselii (von Martens 1869) MK83
                                |--Nerocila T08
                                |--Renocila ovata Bruce 1987 BP03
                                |--Ourozeuktes Milne Edwards 1840 T08
                                |    `--O. bopyroides (Lesueur 1814) [incl. O. owenii Milne Edwards 1840] T08
                                |--Cteatessa Schioedte & Meinert 1883 T08
                                |    `--C. retusa (Schioedte & Meinert 1878) T08
                                |--Cterissa Schioedte & Meinert 1884 T08
                                |    `--C. pterygota (Koelbel 1878) T08
                                |--Emetha Schioedte & Meinert 1883 T08
                                |    `--E. audouini (Milne Edwards 1840) T08
                                |--Glossobius Schioedte & Meinert 1883 T08
                                |    `--G. impressa (Say 1818) T08
                                |--Lobothorax Bleeker 1856 T08
                                |    `--L. typus Bleeker 1857 (see below for synonymy) T08
                                |--Norileca Bruce 1990 T08
                                |    `--N. indica (Milne Edwards 1840) [=Livoneca indica] T08
                                |--Meinertia MK83
                                |    |--M. deplanata (Bovallius 1885) MK83
                                |    `--M. transversa Richardson 1900 MK83
                                |--Elthusa Schioedte & Meinert 1884 T08
                                |    |--E. raynaudii (Milne Edwards 1840) T08
                                |    `--E. vulgaris (Stimpson 1857) [=Livoneca vulgaris] T08
                                |--Ichthyoxenus Herklots 1870 T08
                                |    |--I. fushanensis Tsai & Dai 1999 TD03
                                |    `--I. jellinghausi Herklots 1870 T08
                                |--Livoneca Leach 1818 T08
                                |    |--L. californica H47
                                |    |--L. redmanii Leach 1818 T08
                                |    `--L. sinuata Koelbel 1878 T08
                                |--Mothocya Costa 1851 T08
                                |    |--M. epimerica Costa 1851 T08
                                |    |--M. longicopa C02
                                |    |--M. nana (Schiödte & Meinert 1884) [=Irona nana] MK83
                                |    |--M. plagulophora (Haller 1880) [incl. Irona vatia Schioedte & Meinert 1884] T08
                                |    |--M. renardi (Bleeker 1857) T08
                                |    `--M. sajori Bruce 1986 T08
                                |--Ceratothoa Dana 1852 T08
                                |    |--C. carinata Bianconi 1869 T08
                                |    |--C. imbricata (Fabricius 1775) [=Codonophilus imbricatus] T08
                                |    |--C. impressa (Say 1818) MK83
                                |    |--C. italica Schioedte & Meinert 1883 T08
                                |    |--C. oestroides (Risso 1826) T08
                                |    |--C. oxyrrhynchaena Koelbel 1878 T08
                                |    `--C. parallela (Otto 1828) T08
                                |--Anilocra Leach 1818 T08
                                |    |--A. acuta Richardson 1910 MK83
                                |    |--A. capensis Leach 1818 T08
                                |    |--A. frontalis Milne Edwards 1840 T08
                                |    |--A. holocentri Williams & Williams 1981 T08
                                |    |--A. laticauda Milne Edwards 1840 MK83
                                |    |--A. leptosoma Bleeker 1857 T08
                                |    |--A. physodes (Linnaeus 1758) T08
                                |    |--A. plebia Schiödte & Meinert 1883 MK83
                                |    `--A. vittata Lucas 1846 E12
                                `--Cymothoa T08

Lobothorax typus Bleeker 1857 [incl. Saophra aurita Schioedte & Meinert 1883, L. aurita] T08

*Type species of generic name indicated

REFERENCES

[BBN86] Botosaneanu, L., N. Bruce & J. Notenboom. 1986. Isopoda: Cirolanidae. In: Botosaneanu, L. (ed.) Stygofauna Mundi: A Faunistic, Distributional, and Ecological Synthesis of the World Fauna inhabiting Subterranean Waters (including the Marine Interstitial) pp. 412–422. E. J. Brill / Dr. W. Backhuys: Leiden.

[BP03] Brandt, A. & G. C. B. Poore. 2003. Higher classification of the flabelliferan and related Isopoda based on a reappraisal of relationships. Invertebrate Systematics 17: 893–923.

[B78] Brusca, R. C. 1978. Studies on the cymothoid fish symbionts of the eastern Pacific (Crustacea: Isopoda: Cymothoidae) II. Systematics and biology of Lironeca vulgaris Stimpson 1857. Occasional Papers of the Allan Hancock Foundation New Series 2: 1–19.

[C02] Collette, B. B. 2002. Needlefishes. Family Belonidae. In: Collette, B. B. & G. Klein-MacPhee (eds) Bigelow and Schroeder’s Fishes of the Gulf of Maine 3^rd ed. pp. 287–289. Smithsonian Institute Press: Washington.

[E12] Evenhuis, N. L. 2012. Publication and dating of the Exploration Scientifique de l'Algérie: Histoire Naturelle des Animaux Articulés (1846–1849) by Pierre Hippolyte Lucas. Zootaxa 3448: 1–61.

[F05] Fernández, J. 2005. Noticia de nuevos táxones para la ciencia en el ámbito Íbero-Balear y Macaronésico. Nuevos táxones animales descritos en la península Ibérica y Macaronesia desde 1994 (IX). Graellsia 61 (2): 261–282.

[H47] Hatch, M. H. 1947. The Chelifera and Isopoda of Washington and adjacent regions. University of Washington Publications in Biology 10 (5): 155–274.

[MK83] Menzies, R. J., & W. L. Kruczynski. 1983. Isopod Crustacea (exclusive of Epicaridea). Memoirs of the Hourglass Cruises 6 (1): 1–126.

[M02] Musick, J. A. 2002. Sturgeons. Family Acipenseridae. In: Collette, B. B. & G. Klein-MacPhee (eds) Bigelow and Schroeder’s Fishes of the Gulf of Maine 3^rd ed. pp. 83–88. Smithsonian Institute Press: Washington.

[T08] Trilles, J.-P. 2008. Some marine isopods from the Senckenberg Research Institute (Frankfurt am Main, Germany) (Crustacea, Isopoda: Cymothoidae, Aegidae, Corallanidae, Cirolanidae). Senckenbergiana Biologica 88 (1): 21–28.

[TD03] Tsai, M.-L., & C.-F. Dai. 2003. Cannibalism within mating pairs of the parasitic isopod, Ichthyoxenus fushanensis. Journal of Crustacean Biology 23 (3): 662–668.

Limnoriidea

Gribbles Limnoria lignorum, from NOAA.

Belongs within: Scuticoxifera.

The Limnoriidea are a group of marine isopods united by the absence of a mandibular molar (Brandt & Poore 2003). The gribble Limnoria lignorum is a significant pest of marine timbers, gnawing into exposed wood (Hatch 1947).

Characters (from Brandt & Poore 2003): Pleonites 1–5 or 1–4 free, pleonite 1 similar to others or narrower. Pleotelson underside flat, without ventrolateral ridges (pleopods not enclosed laterally). Antenna 1 with or without scale. Mandibular lacinia mobilis present or absent. Mandibular molar absent. Maxillipedal enditre reaching at least distal margin of palp article 4, non-tapering, slender (except in Keuphylia). Dorsal coxal plate 7 similar to others. Pleopods 3–5 endopods more or less oval or rectangular, rounded distally. Uropodal rami ventral to pleotelson, articulating from side to side along vertical axis or distal.

<==Limnoriidea [Limnorioidea] BP03
    |--Keuphylia [Keuphyliidae] BP03
    |    `--K. nodosa Bruce 1980 BP03
    `--+--Hadromastax Bruce 1988 [Hadromastacidae] BP03
       |    `--H. polynesica Bruce & Müller 1991 BP03
       `--Limnoriidae BP03
            |--Paralimnoria andrewsi (Calman 1910) BP03
            |--Lynseia [Lynseiidae] BP03
            `--Limnoria BP03
                 |--L. lignorum H47
                 |--L. pfefferi Stebbing 1904 MK83
                 |--L. platycauda Menzies 1957 MK83
                 |--L. quadripunctata Holthuis 1949 BP03
                 |--L. saseboensis Menzies 1957 MK83
                 |--L. simulata Menzies 1957 MK83
                 |--L. stephenseni BP03
                 `--L. tripunctata Menzies 1951 MK83

*Type species of generic name indicated

REFERENCES

[BP03] Brandt, A. & G. C. B. Poore. 2003. Higher classification of the flabelliferan and related Isopoda based on a reappraisal of relationships. Invertebrate Systematics 17: 893–923.

[H47] Hatch, M. H. 1947. The Chelifera and Isopoda of Washington and adjacent regions. University of Washington Publications in Biology 10 (5): 155–274.

[MK83] Menzies, R. J., & W. L. Kruczynski. 1983. Isopod Crustacea (exclusive of Epicaridea). Memoirs of the Hourglass Cruises 6 (1): 1–126.

Cirolaninae

Cirolana harfordi, copyright Peter J. Bryant.

Belongs within: Cymothoida.

The Cirolaninae are a group of mostly marine isopods with a secondary unguis present on the pereopodal dactyli (Kensley & Schotte 1989).

Characters (from Kensley & Schotte 1989): Frontal lamina short, flat. Clypeus flattened, not projecting. Antennal peduncular articles 4 and 5 subequal, longer than articles 1–3. Pereopods with secondary unguis on dactyli. Penes reduced or absent. Pleonite 5 always overlapped by pleonite 4. Pleopod 2 in male with copulatory stylet articulating basally.

<==Cirolaninae
    |--Parabathynomus BBN86
    |--Hansenolana sphaeromiformis (Hansen 1890) BBN86, MK83
    |--Neocirolana BBN86
    |--Cartetolana BBN86
    |--Gnatholana BBN86
    |--Oncilorpheus stebbingi Paul & Menzies 1971 BBN86, MK83
    |--Creaseriella Rioja 1953 BBN86
    |    `--C. anops (Creaser 1936) BBN86
    |--Saharolana Monod 1930 BBN86
    |    `--S. seurati Monod 1930 BBN86
    |--Antrolana Bowman 1964 BBN86
    |    `--A. lira Bowman 1964 BBN86
    |--Cirolanides Benedict 1896 BBN86
    |    `--C. texensis Benedict 1896
    |--Mexilana Bowman 1975 BBN86
    |    `--M. saluposi Bowman 1975 BBN86
    |--Faucheria Dollfus & Viré 1905 BBN86
    |    `--F. faucheri (Dollfus & Viré 1900) BBN86
    |--Bathynomus Milne Edwards 1879 BBN86, BP03
    |    |--B. giganteus Milne Edwards 1879 MK83
    |    `--B. immanis Bruce 1986 BP03
    |--Haptolana Bowman 1966 BBN86
    |    |--H. pholeta Bruce & Humphreys 1993 PH13
    |    |--H. somala Messana & Chelazzi 1984 BBN86
    |    `--H. trichostoma Bowman 1966 BBN86
    |--Bahalana Carpenter 1981 BBN86
    |    |--B. cardiopus Notenboom 1981 BBN86
    |    `--B. geracei Carpenter 1981 BBN86
    |--Turcolana Argano & Pesce 1980 BBN86
    |    |--T. cariae Argano & Pesce 1980 BBN86
    |    `--T. rhodica Botosaneanu, Boutin & Henry 1985 BBN86
    |--Skotobaena Ferrara & Monod 1972 BBN86
    |    |--S. monodi Ferrara & Lanza 1978 BBN86
    |    `--S. mortoni Monod 1972 BBN86
    |--Sphaerolana Cole & Minckley 1972 BBN86
    |    |--S. affinis Cole & Minckley 1970 BBN86
    |    `--S. interstitialis Cole & Minckley 1970 BBN86
    |--Anopsilana Paulian & Delamare Deboutteville 1956 (see below for synonymy) BBN86
    |    |--A. acanthura (Notenboom 1981) BBN86
    |    |--A. crenata Bowman & Franz 1982 BBN86
    |    |--A. cubensis (Hay 1903) BBN86
    |    |--A. poissoni Paulian & Delamare Deboutteville 1956 BBN86
    |    `--A. radicicola (Notenboom 1981) BBN86
    |--Speocirolana Bolivar y Pieltain 1950 BBN86
    |    |--S. bolivari (Rioja 1953) BBN86
    |    |--S. endeca Bowman 1981 BBN86
    |    |--S. guerrai Contreras-Balderas & Purata-Velarde 1982 BBN86
    |    |--S. pelaezi (Bolivar y Pieltain 1950) BBN86
    |    |--S. pubens Bowman 1981 BBN86
    |    `--S. thermydronis Cole & Minckley 1966 BBN86
    |--Sphaeromides Dollfus 1897 BBN86
    |    |--S. bureschi Strouhal 1963 BBN86
    |    |    |--S. b. bureschi BBN86
    |    |    `--S. b. serbica Pljakič 1968 BBN86
    |    |--S. polateni Angelov 1968 BBN86
    |    |--S. raymondi Dollfus 1897 BBN86
    |    `--S. virei (Brian 1923) [=Troglaega virei] BBN86
    |         |--S. v. virei BBN86
    |         |--S. v. mediodalmatina Sket 1964 BBN86
    |         `--S. v. montenigrina Sket 1957 BBN86
    |--Typhlocirolana Racovitza 1905 BBN86
    |    |--T. buxtoni Racovitza 1912 BBN86
    |    |--T. fontis (Gurney 1908) BBN86
    |    |--T. gurneyi Racovitza 1912 BBN86
    |    |--T. leptura Botosaneanu, Boutin & Henry 1985 BBN86
    |    |--T. lulli Pujiula 1911 BBN86
    |    |--T. moraguesi Racovitza 1905 BBN86
    |    |--T. reichi Por 1962 BBN86
    |    |--T. rifana Margalef 1958 BBN86
    |    `--T. steinitzi Strouhal 1960 BBN86
    `--Cirolana Leach 1818 MK83
         |--*C. cranchii Leach 1818 MK83
         |--C. borealis Lilljeborg 1851 [incl. C. spinipes Bate & Westwood 1868] MK83
         |--C. chiltoni H47
         |--C. gracilis Hansen 1890 MK83
         |--C. harfordi (Lockington 1877) HJ08
         |--C. kincaidi Hatch 1947 H47
         |--C. koepckei Bott 1954 B54
         |--C. lata BBN86
         |--C. leptanga Bruce 1994 BP03
         |--C. linguifrons B54
         |--C. minuta Hansen 1890 MK83
         |--C. obtruncata Richardson 1901 MK83
         |--C. parva Hansen 1890 [incl. C. albida Richardson 1901, C. diminuta Menzies 1962] MK83
         |--C. polita (Stimpson 1854) [=Aega polita, Conilera polita] MK83
         |--C. salvadorensis Schuster 1954 S54
         |--C. troglexuma FBI01
         `--C. vancouverensis [=C. chiltoni vancouverensis] H47

Anopsilana Paulian & Delamare Deboutteville 1956 [incl. Haitilana Notenboom 1981, Troglocirolana Rioja 1956] BBN86

*Type species of generic name indicated

REFERENCES

[BBN86] Botosaneanu, L., N. Bruce & J. Notenboom. 1986. Isopoda: Cirolanidae. In: Botosaneanu, L. (ed.) Stygofauna Mundi: A Faunistic, Distributional, and Ecological Synthesis of the World Fauna inhabiting Subterranean Waters (including the Marine Interstitial) pp. 412–422. E. J. Brill/Dr. W. Backhuys: Leiden.

[B54] Bott, R. 1954. Ein neuer litoraler Isopod aus Peru. Senckenbergiana Biologica 35: 107–108.

[BP03] Brandt, A. & G. C. B. Poore. 2003. Higher classification of the flabelliferan and related Isopoda based on a reappraisal of relationships. Invertebrate Systematics 17: 893–923.

[FBI01] Fosshagen, A., G. A. Boxshall & T. M. Iliffe. 2001. The Epacteriscidae, a cave-living family of calanoid copepods. Sarsia 86: 245–318.

[H47] Hatch, M. H. 1947. The Chelifera and Isopoda of Washington and adjacent regions. University of Washington Publications in Biology 10 (5): 155–274.

[HJ08] Huisman, J. M., D. S. Jones, F. E. Wells & T. Burton. 2008. Introduced marine biota in Western Australian waters. Records of the Western Australian Museum 25 (1): 1–44.

Kensley, B., & M. Schotte. 1989. Guide to the Marine Isopod Crustaceans of the Caribbean. Smithsonian Institution Press: Washington.

[MK83] Menzies, R. J., & W. L. Kruczynski. 1983. Isopod Crustacea (exclusive of Epicaridea). Memoirs of the Hourglass Cruises 6 (1): 1–126.

[PH13] Poore, G. C. B. & W. F. Humphreys. 2013. Bunderanthura bundera gen. et sp. nov. from Western Australia, first anchialine Leptanthuridae (Isopoda) from the Southern Hemisphere. Records of the Western Australian Museum 28 (1): 21–29.

[S54] Schuster, O. 1954. Zwei neue Crustaceen von der pazifischen Küste Mittel-Amerikas (Amphipoda und Isopoda). Senckenbergiana Biologica 35: 103–105.

Bopyridae

Dorsal (left) and ventral (centre) views of female and dorsal view of male (right) of Parabopyrella richardsonae, from Marilyn Schotte.

Belongs within: Epicaridea.

The Bopyridae are a group of isopods parasitic as adults on decapod crustaceans, while larvae (where known) are parasites of calanoid copepods (Markham 1985).

Characters (from Markham 1985): Female distinctly segmented, slightly to greatly asymmetrical, depressed. Head occasionally fused with first pereomere, sometimes enclosed by frontal lamina; antenna rudimentary; maxilliped of two segments, often with anteromedial palp; posteroventral border of head usually with one or two lateral projections on each side. All seven pereomeres usually distinct; pereopods reduced, usually of seven pairs, though up to six occasionally absent from one side; pereopods prehensile, dorsal to lateral, isomorphic; oostegites of five (rarely seven) pairs often produced into lateral plates resembling pleopodal rami; pleopods generally present on all but final pleomere; pleopods rudimentary or modified for respiration, uniramous or biramous, isomorphic; uropods, when present, terminal, uniramous or biramous, often of same form as pleopods or lateral plates. Male much smaller than female, at least twice as long as wide, symmetrical, distinctly segmented. Head rounded anteriorly, occasionally fused with first pereomere; antennae often prominent. Pereon of seven distinct pereomeres; pair of ventral prehensile pereopods on each pereomere, usually isomorphic except first and sometimes second rarely conspicuously larger than others. Pleon of one to six pleomeres; if unsegmented, usually lacking appendages; if multisegmented, often with sessile to flaplike uniramous or biramous pleopods on each pleomere but last; uropods, if present, uniramous or biramous and terminal.

<==Bopyridae [Bopyrina]
    |  i. s.: Leidya distortus (Leidy 1855) [=Cepon distortus] M85
    |         Munidion parvum H47
    |         Phyllodurus abdominalis H47
    |--Athelginae M85
    |--Ione H47 [Ioninae M85]
    |    `--I. cornuta [incl. I. brevicauda Bonnier 1900] H47
    |--Argeia Dana 1852 [Argeiinae] M85
    |    |--*A. pugettensis Dana 1852 M85
    |    `--A. atlantica Markham 1977 M85
    |--Pseudioninae M85
    |    |--Pontobopyrus Markham 1979 M85
    |    |    `--*P. abyssorum Markham 1979 M85
    |    `--Pseudione Kossmann 1881 M85
    |         |--*P. callianassae Kossmann 1881 M85
    |         |--P. affinis (Sars 1882) [=Pleurocrypta affinis; incl. Palaegyge plesionikae Barnard 1920] M85
    |         |--P. cognata Markham 1985 M85
    |         |--P. elongata M85
    |         |--P. galacanthae H47
    |         |--P. giardi H47
    |         |--P. indica M85
    |         |--P. magna Shiino 1951 M85
    |         `--P. tattersalli M85
    |--Hemiarthrinae [Phrixinae, Phryxinae] M85
    |    |--Allodiplophryxus Markham 1985 M85
    |    |    `--*A. floridanus Markham 1985 M85
    |    |--Loki Markham 1972 M85
    |    |    `--*L. circumsaltanus Markham 1972 M85
    |    |--Azygopleon Markham 1985 M85
    |    |    `--*A. schmitti (Pearse 1932) [=Hemiarthrus schmitti] M85
    |    |--Eophrixus Caroli 1930 [incl. Hypophryxus Shiino 1934] M85
    |    |    |--*E. lysmatae (Caroli 1930) [=Phrixus (Eophrixus) lysmatae] M85
    |    |    `--E. subcaudalis (Hay 1917) (see below for synonymy) M85
    |    |--Dicropleon Markham 1972 M85
    |    |    |--*D. periclimenis Markham 1972 M85
    |    |    `--D. morator Markham 1982 M85
    |    |--Hyperphrixus Nierstrasz & Brender à Brandis 1931 M85
    |    |    |--*H. tattersalli Nierstrasz & Brender à Brandis 1931 M85
    |    |    `--H. castrensis Markham 1985 M85
    |    |--Metaphrixus Nierstrasz & Brender à Brandis 1931 M85
    |    |    |--*M. carolii Nierstrasz & Brender à Brandis [=Metaphryxus carolii] M85
    |    |    `--M. intutus Bruce 1965 M85
    |    |--Hemiarthrus Giard & Bonnier 1887 [=Phryxus (preoc.), Phrixus Caroli 1930] M85
    |    |    |--*H. abdominalis (Krøyer 1840-1841) (see below for synonymy) M85
    |    |    |--H. nematocarcini Stebbing 1914 M85
    |    |    `--H. synalphei (Pearse 1950) [=Diplophryxus synalphei] M85
    |    `--Diplophryxus Richardson 1904 M85
    |         |--*D. jordani Richardson 1904 M85
    |         |--D. alphei Shiino 1934 M85
    |         |--D. kempi Chopra 1930 M85
    |         `--D. richardsoni Chopra 1930 M85
    `--Bopyrinae M85
         |--Schizobopyrina M85
         |--Bopyrus Latreille 1802 M85
         |    `--*B. crangorum (Fabricius 1798) [=Monoculus crangorum, B. squillarum Latreille 1802] M85
         |--Bathygyge Hansen 1897 M85
         |    `--*B. grandis Hansen 1897 M85
         |--Probopyrinella Nierstrasz & Brender à Brandis 1929 M85
         |    `--*P. latreuticola (Gissler 1882) (see below for synonymy) M85
         |--Urobopyrus Richardson 1904 M85
         |    `--*U. processae Richardson 1904 [=U. processi (l. c.); incl. U. provisorius Rouch & Taberly 1961] M85
         |--Parabopyriscus Markham 1985 M85
         |    `--*P. stellatus Markham 1985 M85
         |--Probopyria Markham 1985 M85
         |    `--*P. alphei (Richardson 1900) (see below for synonymy) M85
         |--Ovobopyrus Markham 1985 M85
         |    `--*O. alphezemiotes Markham 1985 M85
         |--Bopyriscus Richardson 1905 M85
         |    `--*B. calmani Richardson 1905 M85
         |--Bopyroides Stimpson 1864 M85
         |    |--B. hippolytes (Krøyer 1838) (see below for synonymy) M85
         |    `--B. cluthae M85
         |--Bopyrinella Nierstrasz & Brender à Brandis 1925 M85
         |    |--B. thorii (Richardson 1904) (see below for synonymy) M85
         |    |--B. albida Shiino 1958 M85
         |    `--B. nipponica Shiino 1936 [=B. antillensis var. nipponica] M85
         |--Probopyrus Giard & Bonnier 1888 [incl. Palaegyge Giard & Bonnier 1888, Palegyge] M85
         |    |--*P. ascendens (Semper 1881) [=Bopyrus ascendens] M85
         |    |--‘*Palaegyge’ borrei Giard & Bonnier 1888 M85
         |    |--P. palaemoni Lemos de Castro & Brasil Lima 1974 M85
         |    `--P. pandalicola (Packard 1879) (see below for synonymy) M85
         |--Bopyrione Bourdon & Markham 1980 M85
         |    |--*B. synalphei Bourdon & Markham 1980 M85
         |    |--B. longicapitata Markham 1982 M85
         |    |--B. toloensis Markham 1982 M85
         |    `--B. woodmasoni (Chopra 1923) M85
         |--Synsynella Hay 1917 [incl. Bopyro Pearse 1932, Prosynsynella Nierstrasz & Brender à Brandis 1929] M85
         |    |--*S. deformans Hay 1917 [=Bopyrella deformans] M85
         |    |--‘*Prosynsynella’ hayi Nierstrasz & Brender à Brandis 1929 (see below for synonymy) M85
         |    |--‘Bopyrella’ inoi Shiino 1949 M85
         |    `--S. integra Bourdon 1981 M85
         |--Bopyrella Bonnier 1900 M85
         |    |--*B. thomsoni Bonnier 1900 M85
         |    |--B. harmopleon Bowman 1956 M85
         |    |--B. macginitiei M85
         |    `--B. malensis M85
         |--Bopyrina Kossmann 1881 M85
         |    |--*B. ocellata (Czerniavsky 1869) [=Bopyrus ocellatus; incl. Bopyrina giardi] M85
         |    |--B. abbreviata Richardson 1904 M85
         |    |--B. choprae M85
         |    |--B. crangona Pearse 1953 S69
         |    |--B. gigas M85
         |    |--B. pleurocephala M85
         |    `--B. sewelli M85
         `--Parabopyrella Markham 1985 M85
              |--*P. mortenseni (Nierstrasz & Brender à Brandis 1929) [=Bopyrella mortenseni] M85
              |--P. delagoae (Bourdon 1982) [=Bopyrella delagoae] M85
              |--P. incisa (Bourdon 1982) [=Bopyrella incisa] M85
              |--P. lata (Nierstrasz & Brender à Brandis 1929) [=Bopyrella lata] M85
              |--‘Bopyrella’ nierstraszi M85
              |--P. richardsonae (Nierstrasz & Brender à Brandis 1929) [=Bopyrella richardsonae] M85
              |--P. saronae (Bourdon & Bruce 1979) [=Bopyrella saronae] M85
              `--P. thomasi (Nierstrasz & Brender à Brandis 1929) [=Bopyrella thomasi] M85

Bopyrinella thorii (Richardson 1904) [=Bopyrina thorii; incl. *Bopyrinella antillensis Nierstrasz & Brender à Brandis 1925]

Bopyroides hippolytes (Krøyer 1838) [=Bopyrus hippolytes, Bobyroides (l. c.) hippolytes, Gyge hippolytes, Pleurocrypta hippolytes; incl. *Bopyroides acutimarginatus Stimpson 1864] M85

Eophrixus subcaudalis (Hay 1917) [=Phryxus subcaudalis, Hemiarthrus subcaudalis, Phrixus (Paraphrixus) subcaudalis] M85

*Hemiarthrus abdominalis (Krøyer 1840-1841) [=Bopyrus abdominalis, *Phrixus abdominalis, *Phryxus abdominalis; incl. Phryxus hippolytes Rathke 1843] M85

*Probopyria alphei (Richardson 1900) [=Bopyrus alphei, Bopyrella alphei, Probopyrus alphei; incl. Capitetragonia asperotibialis Pearse 1953] M85

*Probopyrinella latreuticola (Gissler 1882) [=Bopyroides latreuticola, Bopyrina latreuticola, Probopyrus latreuticola; incl. Bopyrella asymmetrica Nierstrasz & Brender à Brandis 1929, Bopyrus latreutis Bate 1888] M85

Probopyrus pandalicola (Packard 1879) [=Bopyrus pandalicola, Pr. pandicola (l. c.); incl. Pr. bithynis Richardson 1904, Pr. bythinis (l. c.), Pr. creaseri Pearse 1936, Pr. floridensis Richardson 1904, Pr. bithynis var. gigas Nierstrasz & Brender à Brandis 1929, Pr. floridensis var. gigas, Bopyrus manhattensis Gissler 1881, Palaegyge meeki Richardson 1912, Probopyrus meeki, Pr. oviformis Nierstrasz & Brender à Brandis 1929, Pa. oviformis, Bopyrus palaemoneticola Gissler 1882, Probopyrus palaemoneticola, Pr. panamensis Richardson 1912, Pr. papaloapanensis Rioja 1949, Pr. papaloanensis (l. c.)] M85

‘*Prosynsynella’ hayi Nierstrasz & Brender à Brandis 1929 [incl. *Bopyro choprae Pearse 1932, Bopyrina choprae, Bopyrus chopre (l. c.)] M85

*Type species of generic name indicated

REFERENCES

[H47] Hatch, M. H. 1947. The Chelifera and Isopoda of Washington and adjacent regions. University of Washington Publications in Biology 10 (5): 155–274.

[M85] Markham, J. C. 1985. A review of the bopyrid isopods infesting caridean shrimps in the northwestern Atlantic Ocean, with special reference to those collected during the Hourglass Cruises in the Gulf of Mexico. Memoirs of the Hourglass Cruises 7 (3): 1–156.

S69] Steyskal, G. C. 1969. The mistreatment of the Latin genitive case in forming names of parasites. Systematic Zoology 18 (3): 339–342.

Asellus (Asellus)

Water hoglouse Asellus aquaticus, copyright Peter Pfeiffer.

Belongs within: Asellidae.

The subgenus Asellus of the genus freshwater isopod genus Asellus is characterised by a lack of spines on the propodite of the male first legs (Hatch 1947).

Characters (from Hatch 1947): Body usually elongate, depressed. Head usually narrower and shorter than first thoracic segment; eyes usually present; mandible with three-segmented palp. Abdomen usually not longer than broad. Male with inner margin of posterior surface of propodite of first legs not produced in teeth or immovable spines.

<==Asellus Geoffroy 1762 (Asellus) HLM86
    |--A. (A.) amamiensis Matsumoto 1961 HLM86
    |--A. (A.) aquaticus (Linnaeus 1758) HLM86
    |    |--A. a. aquaticus HLM86
    |    |--A. a. carsicus Karaman 1952 HLM86
    |    |--A. a. cavernicolus Racovitza 1925 HLM86
    |    |--A. a. cyclobranchialis Sket 1965 HLM86
    |    |--A. a. fribergensis Schneider 1887 HLM86
    |    `--A. a. longicornis Sket 1965 HLM86
    |--A. (A.) birsteini Levanidov 1976 HLM86
    |--A. (A.) ezoensis Matsumoto 1962 HLM86
    |--A. (A.) hilgendorfi Bovallius 1886 HLM86
    |    |--A. h. hilgendorfi Bovallius 1886 HLM86
    |    `--A. h. ryukyuensis Ueno 1938 HLM86
    |--A. (A.) hyugaensis Matusmoto 1960 HLM86
    |--A. (A.) kumaensis Matsumoto 1960 HLM86
    |--A. (A.) monticola [incl. A. monticola var. fontinalis Birstein 1936] HLM86
    |--A. (A.) musashiensis Matsumoto 1961 HLM86
    |--A. (A.) shikokuensis Matsumoto 1960 HLM86
    |--A. (A.) tamaensis Matsumoto 1961 HLM86
    `--A. (A.) tomalensis H47

*Type species of generic name indicated

REFERENCES

[H47] Hatch, M. H. 1947. The Chelifera and Isopoda of Washington and adjacent regions. University of Washington Publications in Biology 10 (5): 155–274.

[HLM86] Henry, J.-P., J. J. Lewis & G. Magniez. 1986. Isopoda: Asellota: Aselloidea, Gnathostenetroidoidea, Stenetrioidea. In: Botosaneanu, L. (ed.) Stygofauna Mundi: A Faunistic, Distributional, and Ecological Synthesis of the World Fauna inhabiting Subterranean Waters (including the Marine Interstitial) pp. 434–464. E. J. Brill/Dr. W. Backhuys: Leiden.

Asellidae

One-spotted waterlouse Proasellus meridianus, copyright Keith Lugg.

Belongs within: Asellota.
Contains: Asellus (Asellus).

The Asellidae are a group of freshwater crustacean of the Holarctic region (Henry et al. 1986).

Characters (from Henry et al. 1986): Antenna bearing a squama; maxillipedal oostegite reduced to a "Wasserstrudelapparat" (water swirling apparatus); pleonites 1 and 2 reduced both in length and width, forming a stalk between pereion and pleotelson; male copulatory organ (endopodite of pleopods 2) secondarily uniarticulate, but complex.

<==Asellidae
    |--Baicalasellus HLM86
    |--Gallasellus Henry & Magniez 1977 HLM86
    |    `--G. heilyi (Legrand 1956) HLM86
    |--Stygasellus Chappuis 1943 HLM86
    |    `--S. phreaticus (Chappuis 1943) [=Stygonectes phreaticus] HLM86
    |--Uenasellus Matsumoto 1962 HLM86
    |    `--U. iyoensis (Matsumoto 1960) HLM86
    |--Salmasellus Bowman 1975 HLM86
    |    `--S. steganothrix Bowman 1975 HLM86
    |--Psammasellus Braga 1968 HLM86
    |    `--P. capitatus Braga 1968 HLM86
    |--Calasellus Bowman 1981 HLM86
    |    |--C. californicus (Miller 1933) HLM86
    |    `--C. longus Bowman 1981 HLM86
    |--Lirceolus Bowman & Longley 1976 HLM86
    |    |--L. pilus (Steeves 1968) HLM86
    |    `--L. smithii (Ulrich 1902) HLM86
    |--Lirceus Rafinesque 1820 HLM86
    |    |--L. culveri Holsinger & Bowman 1973 HLM86
    |    |--L. fontinalis HLM86
    |    `--L. usdagalum Estes & Holsinger 1976 HLM86
    |--Nipponasellus Matsumoto 1962 HLM86
    |    |--N. aioii (Chappuis 1955) HLM86
    |    |--N. hubrichti (Matsumoto 1956) HLM86
    |    |--N. kagaensis (Msatsumoto 1958) HLM86
    |    |--N. takefuensis (Matsumoto 1961) HLM86
    |    `--N. tonensis (Matsumoto 1961) HLM86
    |--Bragasellus Henry & Magniez 1968 HLM86
    |    |--B. aireyi Henry & Magniez 1980 HLM86
    |    |--B. boui Henry & Magniez 1974 HLM86
    |    |--B. comasi Henry & Magniez 1976 HLM86
    |    |--B. comasioides Magniez & Brehier 2004 F05
    |    |--B. conimbricensis (Braga 1946) HLM86
    |    |--B. escolai Henry & Magniez 1978 HLM86
    |    |--B. frontellum (Braga 1964) HLM86
    |    |--B. incurvatus Afonso 1984 HLM86
    |    |--B. lagari Henry & Magniez 1973 HLM86
    |    |--B. pauloae (Braga 1958) HLM86
    |    `--B. seabrai (Braga 1943) HLM86
    |--Asellus Geoffroy 1762 HLM86
    |    |  i. s.: A. cavaticus T30
    |    |         A. meridianus N29
    |    |         A. nipponensis Nicholls 1929 N29
    |    |--A. (Asellus) HLM86
    |    |--A. (Conasellus) communis H47
    |    |--A. (Mesoasellus Birstein 1939) HLM86
    |    |    |--A. (M.) dentifer Birstein & Levanidov 1952 HLM86
    |    |    `--A. (M.) strinatii Chappuis 1955 HLM86
    |    `--A. (Phreatosellus Matsumoto 1962) HLM86
    |         |--A. (P.) akiyoshiensis (Ueno 1927) HLM86
    |         |--A. (P.) higoensis Matsumoto 1960 HLM86
    |         |--A. (P.) iriei Matsumoto 1978 HLM86
    |         |--A. (P.) kawamurai (Tattersall 1921) HLM86
    |         |--A. (P.) minatoi Matsumoto 1978 HLM86
    |         |--A. (P.) miurai Chappuis 1955 HLM86
    |         |--A. (P.) uenoi Matsumoto 1978 HLM86
    |         `--A. (P.) yoshinoensis Matsumoto 1960 HLM86
    |--Synasellus Braga 1944 HLM86
    |    |--S. albicastrensis Braga 1960 HLM86
    |    |--S. barcelensis Noodt & Galhano 1969 HLM86
    |    |--S. brigantinus Braga 1959 HLM86
    |    |--S. exiguus Braga 1944 HLM86
    |    |--S. fragilis Braga 1946 HLM86
    |    |--S. insignis Afonso 1984 HLM86
    |    |--S. lafonensis Braga 1959 HLM86
    |    |--S. longicauda Braga 1959 HLM86
    |    |--S. mariae (Braga 1942) HLM86
    |    |    |--S. m. mariae HLM86
    |    |    `--S. m. favaiensis Eiras 1974 HLM86
    |    |--S. mateusi Braga 1954 HLM86
    |    |--S. meirelesi Braga 1959 HLM86
    |    |--S. minutus Braga 1967 HLM86
    |    |--S. nobrei Braga 1967 HLM86
    |    |--S. pireslimai Braga 1959 HLM86
    |    |--S. serranus Braga 1967 HLM86
    |    `--S. transmontanus Braga 1959 HLM86
    |--Caecidotea Packard 1871 HLM86
    |    |--C. acuticarpa Mackin & Hubricht 1940 HLM86
    |    |--C. adenta Mackin & Hubricht 1940 HLM86
    |    |--C. alabamensis Stafford 1911 HLM86
    |    |--C. ancyla (Fleming 1972) HLM86
    |    |--C. antricola (Creaser 1931) HLM86
    |    |--C. barri (Steeves 1965) HLM86
    |    |--C. beattyi Lewis & Bowman 1981 HLM86
    |    |--C. bicrenata (Steeves 1963) HLM86
    |    |    |--C. b. bicrenata HLM86
    |    |    `--C. b. whitei Lewis & Bowman 1981 HLM86
    |    |--C. bisetus (Steeves 1968) HLM86
    |    |--C. bowmani Lewis 1980 HLM86
    |    |--C. brevicauda HLM86
    |    |--C. cannula (Steeves 1963) HLM86
    |    |--C. carolinensis Lewis & Bowman 1977 HLM86
    |    |--C. catachaetus (Fleming & Steeves 1972) HLM86
    |    |--C. chiapas Bowman 1975 HLM86
    |    |--C. circulus (Steeves 1968) HLM86
    |    |--C. cyrtorhynchus (Fleming & Steeves 1972) HLM86
    |    |--C. dentadactyla HLM86
    |    |--C. dimorpha Mackin & Hubricht 1940 HLM86
    |    |--C. extensolingualus (Fleming 1972) HLM86
    |    |--C. filicispeluncae Bowman & Hobbs 1983 HLM86
    |    |--C. franzi Lewis 1981 HLM86
    |    |--C. fustis Lewis 1981 HLM86
    |    |--C. henroti (Bresson 1955) HLM86
    |    |--C. hobbsi Maloney 1939 HLM86
    |    |--C. holsingeri (Steeves 1963) HLM86
    |    |--C. incurva (Steeves 1968) HLM86
    |    |--C. jordani (Eberly 1965) HLM86
    |    |--C. kendeighi (Steeves & Seidenbergh 1971) HLM86
    |    |--C. lesliei Lewis & Bowman 1981 HLM86
    |    |--C. macropropoda Chase & Blair 1937 [incl. C. ozarkana Chase & Blair 1937] HLM86
    |    |--C. metcalfi (Fleming 1972) HLM86
    |    |--C. mitchelli Argano 1972 HLM86
    |    |--C. nickajackensis Packard 1881 HLM86
    |    |--C. nortoni (Steeves 1966) HLM86
    |    |--C. oculata Mackin & Hubricht 1940 HLM86
    |    |--C. packardi Mackin & Hubricht 1940 HLM86
    |    |--C. parva (Steeves 1964) HLM86
    |    |--C. pasquinii Argano 1972 HLM86
    |    |--C. paurotrigonus (Fleming 1972) HLM86
    |    |--C. pricei Levi 1949 HLM86
    |    |--C. recurvata (Steeves 1963) HLM86
    |    |--C. reddelli (Steeves 1968) HLM86
    |    |--C. richardsonae Hay 1901 HLM86
    |    |--C. rotunda Bowman & Lewis 1984 HLM86
    |    |--C. salemensis Lewis 1981 HLM86
    |    |--C. scrupulosa HLM86
    |    |--C. scyphus (Steeves 1968) HLM86
    |    |--C. sequoiae Bowman 1975 HLM86
    |    |--C. serrata (Fleming 1972) HLM86
    |    |--C. simonini (Bresson 1955) HLM86
    |    |--C. sinuncus (Steeves 1965) HLM86
    |    |--C. spatulata Mackin & Hubricht 1940 HLM86
    |    |--C. steevesi (Fleming 1972) HLM86
    |    |--C. stiladactyla Mackin & Hubricht 1940 HLM86
    |    |--C. stygia Packard 1871 HLM86
    |    |--C. teresae Lewis 1982 HLM86
    |    |--C. tridentata Hungerford 1922 HLM86
    |    |--C. vandeli (Bresson 1955) HLM86
    |    |--C. vomeroi Argano 1972 HLM86
    |    `--C. zullinii Argano 1972 HLM86
    `--Proasellus Dudich 1925 HLM86
         |--P. acutianus Argano & Henry 1972 HLM86
         |--P. adriaticus Argano & Pesce 1979 HLM86
         |--P. albigensis (Magniez 1965) HLM86
         |--P. ambracicus Pesce & Argano 1980 HLM86
         |--P. amiterninus Argano & Pesce 1979 HLM86
         |--P. anophtalmus (Karaman 1934) HLM86
         |    |--P. a. anophtalmus HLM86
         |    |--P. a. bosnicus (Sket 1965) HLM86
         |    |--P. a. dalmatinus (Karaman 1955) HLM86
         |    |--P. a. longisetus (Remy 1941) HLM86
         |    |--P. a. radikanus (Karaman 1955) HLM86
         |    |--P. a. rhausinus (Remy 1941) HLM86
         |    `--P. a. serbicus (Karaman 1955) HLM86
         |--P. aquaecalidae (Racovitza 1922) HLM86
         |--P. arnautovici HLM86
         |    |--P. a. arnautovici HLM86
         |    `--P. a. elongatus (Karaman 1953) HLM86
         |--P. arthrodilus (Braga 1945) HLM86
         |--P. bagradicus Henry & Magniez 1972 HLM86
         |--P. bardaunii Alouf, Henry & Magniez 1982 HLM86
         |--P. basnosanui (Codreanu & Codreanu 1962) HLM86
         |--P. bellesi Henry & Magniez 1982 HLM86
         |--P. beroni Henry & Magniez 1968 HLM86
         |--P. boui Henry & Magniez 1969 HLM86
         |--P. burgundus Henry & Magniez 1969 HLM86
         |--P. cantabricus Henry & Magniez 1968 HLM86
         |--P. cavaticus (Leydig 1871) HLM86
         |--P. chappuisi Henry & Magniez 1968 HLM86
         |--P. chauvini Henry & Magniez 1978 HLM86
         |--P. coiffaiti Henry & Magniez 1972 HLM86
         |--P. comasi Henry & Magniez 1982 HLM86
         |--P. coxalis (Dollfus 1892) HLM86
         |    |--P. c. coxalis HLM86
         |    |--P. c. africanus (Monod 1924) HLM86
         |    |--P. c. banyulensis (Racovitza 1919) HLM86
         |    |--P. c. cephallenus (Strouhal 1942) HLM86
         |    |--P. c. cyrenaicus (Arcangeli 1942) HLM86
         |    |--P. c. gabriellae (Margalef 1950) HLM86
         |    |--P. c. leucadius (Strouhal 1942) HLM86
         |    |--P. c. lucifugus (Deeleman-Reinhold 1965) HLM86
         |    |--P. c. perarmatus (Remy 1936) HLM86
         |    `--P. c. peyerimhoffi (Racovitza 1919) HLM86
         |--P. cretensis Pesce & Argano 1980 HLM86
         |--P. danubialis (Codreanu & Codreanu 1962) HLM86
         |--P. delhezi Henry & Magniez 1973 HLM86
         |--P. deminutus (Sket 1959) HLM86
         |    |--P. d. deminutus HLM86
         |    |--P. d. intermedius (Sket 1965) HLM86
         |    `--P. d. vulgaris (Sket 1965) HLM86
         |--P. elegans (Codreanu & Codreanu 1962) HLM86
         |--P. escolai Henry & Magniez 1982 HLM86
         |--P. espanoli Henry & Magniez 1982 HLM86
         |--P. exiguus Afonso 1983 HLM86
         |--P. franciscoloi (Chappuis 1955) HLM86
         |--P. gardinii (Arcangeli 1942) HLM86
         |--P. gauthieri (Monod 1924) HLM86
         |    |--P. g. gauthieri HLM86
         |    `--P. g. theodori (Birstein 1951) [incl. P. gauthieri numidicus Monod in Gauthier 1928 (n. n.)] HLM86
         |--P. gjorgjevici (Karaman 1933) HLM86
         |--P. gourbaultae Henry & Magniez 1981 HLM86
         |--P. hercegovinensis (Karaman 1933) HLM86
         |--P. hermallensis (Arcangeli 1938) HLM86
         |--P. hussoni (Remy 1941) HLM86
         |    |--P. h. hussoni HLM86
         |    `--P. h. curvifrons (Remy 1941) HLM86
         |--P. hypogeus (Racovitza 1922) HLM86
         |--P. infirmus (Birstein 1936) HLM86
         |--P. istrianus (Stammer 1932) HLM86
         |--P. jaloniacus Henry & Magniez 1977 HLM86
         |--P. karamani (Remy 1934) HLM86
         |--P. lagari Henry & Magniez 1982 HLM86
         |--P. lescherae Henry & Magniez 1977 HLM86
         |--P. linearis (Birstein 1967) HLM86
         |--P. ljovuschkini (Birstein 1967) HLM86
         |--P. lusitanicus (Frade 1938) HLM86
         |--P. lykaonicus Argano & Pesce 1978 HLM86
         |--P. maleri Henry 1977 HLM86
         |--P. margalefi Henry & Magniez 1982 HLM86
         |--P. mateusorum Afonso 1982 HLM86
         |--P. meridianus (Racovitza 1919) HLM86
         |    |--P. m. meridianus HLM86
         |    `--P. m. belgicus (Arcangeli 1935) HLM86
         |--P. minoicus Pesce & Argano 1980 HLM86
         |--P. monodi (Strouhal 1942) HLM86
         |--P. monsferratus (Braga 1948) HLM86
         |--P. montenigrinus (Karaman 1934) HLM86
         |    |--P. m. montenigrinus HLM86
         |    `--P. m. macedonicus (Karaman 1955) HLM86
         |--P. nolli (Karaman 1952) HLM86
         |--P. notenboomi Henry & Magniez 1982 HLM86
         |--P. parvulus (Sket 1960) HLM86
         |--P. patrizzi (Arcangeli 1952) HLM86
         |--P. pavani (Arcangeli 1942) HLM86
         |    |--P. p. pavani HLM86
         |    `--P. p. orientalis (Sket 1965) HLM86
         |--P. racovitzai Henry & Magniez 1972 HLM86
         |--P. rectangulatus Afonso 1982 HLM86
         |--P. rectus Afonso 1982 HLM86
         |--P. remyi [incl. P. remyi f. nudus (Karaman 1953)] HLM86
         |--P. rouchi Henry 1980 HLM86
         |--P. similis (Birstein 1967) HLM86
         |--P. sketi Henry 1975 HLM86
         |--P. slavus (Remy 1948) HLM86
         |    |--P. s. slavus HLM86
         |    |--P. s. histriae (Sket 1963) HLM86
         |    |--P. s. salisburgensis (Strouhal 1957) HLM86
         |    |--P. s. serbiae (Sket 1963) HLM86
         |    |--P. s. styriacus (Sket 1963) HLM86
         |    |--P. s. vindobonensis (Karaman 1955) HLM86
         |    `--P. s. zeii (Sket 1963) HLM86
         |--P. slovenicus (Sket 1957) HLM86
         |--P. solanasi Henry & Magniez 1972 HLM86
         |--P. spelaeus (Racovitza 1922) HLM86
         |--P. spinipes Afonso 1979 HLM86
         |--P. strouhali (Karaman 1955) HLM86
         |    |--P. s. strouhali HLM86
         |    `--P. s. puteanus (Henry 1966) HLM86
         |--P. synaselloides Henry 1963 HLM86
         |--P. thermonyctophilus (Monod 1924) HLM86
         |--P. valdensis (Chappuis 1948) HLM86
         |--P. vandeli Magniez & Henry 1969 HLM86
         |--P. variegatus Afonso 1982 HLM86
         |--P. vignai Argano & Pesce 1977 HLM86
         `--P. walteri (Chappuis 1948) HLM86

*Type species of generic name indicated

REFERENCES

[F05] Fernández, J. 2005. Noticia de nuevos táxones para la ciencia en el ámbito Íbero-Balear y Macaronésico. Nuevos táxones animales descritos en la península Ibérica y Macaronesia desde 1994 (IX). Graellsia 61 (2): 261–282.

[H47] Hatch, M. H. 1947. The Chelifera and Isopoda of Washington and adjacent regions. University of Washington Publications in Biology 10 (5): 155–274.

[HLM86] Henry, J.-P., J. J. Lewis & G. Magniez. 1986. Isopoda: Asellota: Aselloidea, Gnathostenetroidoidea, Stenetrioidea. In: Botosaneanu, L. (ed.) Stygofauna Mundi: A Faunistic, Distributional, and Ecological Synthesis of the World Fauna inhabiting Subterranean Waters (including the Marine Interstitial) pp. 434–464. E. J. Brill/Dr. W. Backhuys: Leiden.

[N29] Nicholls, G. E. 1929. A new species of Asellus (A. nipponensis) from Japan. Archives de Zoologie Expérimentale et Générale 68 (1): 33–38.

[T30] Tattersall, W. M. 1930. Asellus cavaticus, a blind isopod new to the British fauna, from a well in Hampshire. Proceedings of the Linnean Society of London 141: 7.