Carabodoidea

Dorsum of Leptotocepheus trimucronatus, from Balogh & Balogh (1992 vol. 2).

Belongs within: Brachypylina.
Contains: Nippobodidae, Dampfiella, Beckiella, Odontocepheus (Odontocepheus), Diplobodes, Yoshiobodes, Gibbicepheus (Gibbicepheus), Carabodinae, Fissicepheus (Fissicepheus), Pseudotocepheus (Pseudotocepheus), Plenotocepheus, Otocepheinae, Dolicheremaeus.

The Carabodoidea are a primarily tropical group of oribatid mites, many of which spend as least part of their life as burrowers in decaying wood. Members of this group include the Carabodidae, characterised by more or less parallel lamellae on the prodorsum (Balogh & Balogh 1992). Though diverse in adult biology, all species are probably endophagous as nymphs. Adults of many species are arboreal, feeding on fungi and lichens (Norton & Behan-Pelletier 2009). In a number of genera the notogaster bears longitudinal rows of large tubercles, such as in Gibbicepheus or Diplobodes. The genera Spathulocepheus and Trichocarabodes are distinctive in the high number of setae on their genital plates: eight pairs in Trichocarabodes, ten in Spathulocepheus. Other genera have at most seven pairs of genital setae (Balogh & Balogh 1992).

Members of the family Dampfiellidae have the tibiae and tarsi fused and moving as a single segment. The Otocepheidae are characterised by well-developed pedotecta (outgrowths of the body wall underlying the base of the legs). Members of this family are particularly common in damp habitats such as moist litter (Norton & Behan-Pelletier 2009). The family is sometimes divided between two families, Otocepheidae and Tetracondylidae, on the basis of the development of the pedotecta. In Otocepheidae sensu stricto, pedotecta 2+3 are particularly large, so that they are axe-shaped in ventral view or tailfin-shpaed in dorsal view. In 'Tetracondylidae', the pedotecta are distinct but not so extremely developed (Balogh & Balogh 1992).

Carabodoidea [Otocepheoidea] S04
    |  i. s.: Nippobodidae S04
    |         Tokunocepheus Aoki 1966 [Tokunocepheidae] S04
    |           `--*T. mizusawai Aoki 1966 S04
    |         Carabocepheus Berlese 1910 [Carabocepheidae] S04
    |           `--*C. lounsburyi (Berlese 1910) [=Carabodes (*Carabocepheus) lounsburyi] S04
    |                |--C. l. lounsburyi S04
    |                `--C. l. latior Balogh & Mahunka 1966 S04
    |--Dampfiellidae S04
    |    |--Dampfiella S04
    |    `--Beckiella PL17
    `--+--Carabodidae SK10
       |    |  i. s.: Apotomocepheus Aoki 1965 S04
       |    |           `--*A. gressitti Aoki 1965 S04
       |    |         Meriocepheus Aoki 1973 S04
       |    |           `--*M. peregrinus Aoki 1973 S04
       |    |         Sagittabodes Balogh & Balogh 1992 S04
       |    |           `--*S. sagitta (Balogh & Mahunka 1966) [=Machadocepheus sagitta] S04
       |    |         Tansocepheus Mahunka 1983 S04
       |    |           `--*T. serratus Mahunka 1983 S04
       |    |         Tectocarabodes Mahunka 1988 S04
       |    |           `--*T. monstruosus Mahunka 1988 S04
       |    |         Tuberocepheus Balogh & Mahunka 1969 S04
       |    |           `--*T. longus (Balogh 1964) [=Machadocepheus longus] S04
       |    |         Singabodes Mahunka 1998 S04
       |    |           `--*S. rarus Mahunka 1998 S04
       |    |         Archegocepheus Aoki 1965 S04
       |    |           |--*A. imadatei Aoki 1965 S04
       |    |           `--A. macrofoliatus Wen 1997 S04
       |    |         Bathocepheus Aoki 1978 [incl. Philippobodes Balogh & Balogh 1992] S04
       |    |           |--*B. concavus Aoki 1978 S04
       |    |           `--B. manguiati (Corpuz-Raros 1979) S04 (see below for synonymy)
       |    |         Cavernocarabodes Mahunka 1974 S04
       |    |           |--*C. perreti Mahunka 1974 S04
       |    |           `--C. trigonosternum (Pérez-Íñigo 1976) [=Carabodes trigonosternum] S04
       |    |         Spathulocepheus Balogh & Mahunka 1969 S04
       |    |           |--*S. amazonicus Balogh & Mahunka 1969 S04
       |    |           `--S. plumiger Balogh & Mahunka 1977 S04
       |    |         Malgasodes Mahunka 2000 S04
       |    |           |--*M. curvisetus Mahunka 2000 S04
       |    |           `--M. hungarorum Mahunka 2000 S04
       |    |         Gymnobodes Balogh 1965 S04
       |    |           |--*G. fraterculus (Balogh 1963) [=Carabodes fraterculus] S04
       |    |           |--G. semengok Mahunka 1996 S04
       |    |           `--G. subnudus (Balogh 1963) [=Carabodes subnudus] S04
       |    |         Pseudocarabodes Mahunka 1991 [incl. Bakobodes Mahunka 1996, Yemenobodes Mahunka 2000] S04
       |    |           |--*P. xenus Mahunka 1991 S04
       |    |           |--P. improvisus (Mahunka 2000) [=Yemenobodes improvisus] S04
       |    |           `--P. orangutan (Mahunka 1996) [=Bakobodes orangutan] S04
       |    |         Trichocarabodes Balogh 1961 [incl. Ceylobodes Balogh & Balogh 1992] S04
       |    |           |--*T. celisi (Balogh 1958) [=Carabodes celisi] S04
       |    |           |--T. capillatus Balogh 1970 S04 [=*Ceylobodes capillatus BB92]
       |    |           |--T. costulatus Mahunka 1984 S04
       |    |           `--T. hettigei Balogh 1970 S04
       |    |         Hardybodes Balogh 1970 [incl. Carabodella Mahunka 1986] S04
       |    |           |--*H. mirabilis Balogh 1970 S04
       |    |           |--H. calcaratus (Mahunka 1986) S04 [=*Carabodella calcarata BB92]
       |    |           |--H. flabellatus Mahunka 1995 S04
       |    |           `--H. penicillatus Mahunka 1995 S04
       |    |         Cubabodes Balogh & Mahunka 1974 S04
       |    |           |--*C. hexagonalis Balogh & Mahunka 1974 S04
       |    |           |--C. confertus Balogh & Mahunka 1980 S04
       |    |           |--C. radiatus Balogh & Mahunka 1974 S04
       |    |           |--C. spathulatus Balogh & Mahunka 1980 S04
       |    |           `--C. verrucatus Balogh & Mahunka 1980 S04
       |    |         Machadocepheus Balogh 1958 S04
       |    |           |--*M. excavatus Balogh 1958 S04
       |    |           |--M. filiferus Mahunka 1986 S04
       |    |           |--M. pauliani Balogh 1962 S04
       |    |           |--M. phyllophorus Balogh 1970 S04
       |    |           |--M. taprobanicus Balogh 1970 S04
       |    |           `--M. tuberculosus Mahunka 1987 S04
       |    |         Odontocepheus Berlese 1913 S04
       |    |           |--O. (Odontocepheus) S04
       |    |           |--O. (Aokiella Balogh & Mahunka 1967) S04
       |    |           |    |--O. (*A.) florens (Balogh & Mahunka 1967) [=*Aokiella florens] S04
       |    |           |    `--O. (A.) rotundus (Hammer 1979) [=Aokiella rotunda] S04
       |    |           `--O. (Indotocepheus Mondal & Kundu 1999) S04
       |    |                `--O. (*I.) himalayensis (Mondal & Kundu 1999) [=*Indotocepheus himalayensis] S04
       |    |         Pasocepheus Aoki 1976 S04
       |    |           |--P. (Pasocepheus) [incl. Opisthocepheus Aoki 1976] S04
       |    |           |    |--*P. (P.) triarcuatus Aoki 1976 S04
       |    |           |    |--P. (P.) bako Mahunka 1996 S04
       |    |           |    |--P. (P.) eremaeozetoides Mahunka 1995 S04
       |    |           |    `--P. (P.) kirai (Aoki 1976) S04 [=*Opisthocepheus kirai BB92]
       |    |           `--P. (Guineobodes Mahunka 1987) S04
       |    |                `--P. (*G.) papuanus (Balogh 1970) [=Machadocepheus papuanus] S04
       |    |         Congocepheus Balogh 1958 S04
       |    |           |--*C. heterotrichus Balogh 1958 S04
       |    |           |--C. hauseri Mahunka 1989 S04
       |    |           |--C. involutus Mahunka 1997 S04
       |    |           |--C. latilamellatus Mahunka 1984 S04
       |    |           |--C. orientalis Mahunka 1989 S04
       |    |           |--C. ornatus Mahunka 1983 S04
       |    |           |--C. taurus Balogh 1961 S04
       |    |           `--C. velatus Mahunka 1986 S04
       |    |         Diplobodes S04
       |    |         Yoshiobodes S04
       |    |--Gibbicepheus Aoki 1958 S04 [Gibbicepheinae CH98]
       |    |    |--G. (Gibbicepheus) S04
       |    |    `--G. (Gibbibodes Mahunka 1986) S04
       |    |         `--G. (*G.) similis (Mahunka 1986) [=*Gibbibodes similis] S04
       |    `--Carabodinae PL17
       `--Otocepheidae [Tetracondylidae] S04
            |  i. s.: Ocellotocepheus Mahunka 1989 S04
            |           `--*O. tuberculatus Mahunka 1989 S04
            |         Samarocepheus Corpuz-Raros 1990 S04
            |           `--*S. cavernicolus Corpuz-Raros 1990 S04
            |         Cerostocepheus Mahunka 1973 S04
            |           `--*C. trisetosus (Balogh 1970) [=Eurostocepheus trisetosus] S04
            |         Ikarotocepheus Mahunka 1988 S04
            |           `--*I. alatus Mahunka 1988 S04
            |         Philippotocepheus Balogh & Balogh 1992 S04
            |           `--*P. siniloanensis (Corpuz-Raros 1979) [=Papuacepheus siniloanensis] S04
            |         Borneremaeus Mahunka 1991 S04
            |           `--*B. hauseri Mahunka 1991 S04
            |         Bulbocepheus Mahunka 1988 S04
            |           `--*B. hauseorum Mahunka 1988 S04
            |         Fernandocepheus Mahunka 1982 S04
            |           `--*F. franzi Mahunka 1982 S04
            |         Flagellocepheus Balogh 1984 S04
            |           `--*F. sagittatus Balogh 1984 S04
            |         Hydroecocepheus Corpuz-Raros 1979 S04
            |           `--*H. romeoi Corpuz-Raros 1979 C-R79
            |         Lophotocepheus Balogh & Balogh 1983 S04
            |           `--*L. simplex Balogh & Balogh 1983 S04
            |         Neotrichocepheus Hammer 1973 S04
            |           `--*N. tongaensis (Hammer 1973) [=Dolicheremaeus (*Neotrichocepheus) tongaensis] S04
            |         Paradolicheremaeus Tseng 1982 S04
            |           `--*P. tabulatus Tseng 1982 S04
            |         Seboetocepheus Mahunka 1985 S04
            |           `--*S. coronatus Mahunka 1985 S04
            |         Trichocondyla Balogh & Balogh 1986 S04
            |           `--*T. multisetosa Balogh & Balogh 1986 S04
            |         Papuacepheus Balogh 1968 S04
            |           |--*P. geminatus (Balogh 1968) [=Megalotocepheus (*Papuacepheus) geminatus] S04
            |           `--P. vagorum Corpuz-Raros 1991 S04
            |         Trichocepheus Balogh & Mahunka 1966 S04
            |           |--*T. lamellatus Balogh & Mahunka 1966 S04
            |           `--T. auriculus Grobler 1995 S04
            |         Wallworkodes Balogh & Balogh 2002 S04
            |           |--*W. carinatus (Wallwork 1977) [=Carabodes carinatus] S04
            |           `--W. hyalinus (Wallwork 1977) [=Carabodes hyalinus] S04
            |         Kalayaan Corpuz-Raros 1998 S04
            |           |--*K. bonifacioi Corpuz-Raros 1998 S04
            |           |--K. centenarius Corpuz-Raros 1998 S04
            |           `--K. rizali Corpuz-Raros 1998 S04
            |         Rimandocepheus Corpuz-Raros 1998 S04
            |           |--*R. leoi Corpuz-Raros 1998 S04
            |           |--R. brevisetosus Corpuz-Raros 1998 S04
            |           `--R. longisetosus Corpuz-Raros 1998 S04
            |         Afrotocepheus Mahunka 1985 S04
            |           |--*A. (Afrotocepheus) sinarmatus Mahunka 1985 S04
            |           `--A. (Didierotocepheus Mahunka 1994) S04
            |                `--A. (*D.) berndi (Mahunka 1994) [=*Didierotocepheus berndi] S04
            |         Trichotocepheus Aoki 1965 S04
            |           |--*T. erabuensis Aoki 1965 S04
            |           |    |--T. e. erabuensis S04
            |           |    `--T. e. modestus Aoki 1973 S04
            |           `--T. amamiensis Aoki 1965 S04
            |         Fissicepheus Balogh & Mahunka 1967 S04
            |           |--F. (Fissicepheus) S04
            |           `--F. (Psammocepheus Aoki 1970) S04
            |                |--F. (*P.) amabilis Aoki 1970 S04
            |                `--F. (P.) haradai Choi 1986 S04
            |         Longocepheus Balogh & Mahunka 1966 S04
            |           |--*L. australis Balogh & Mahunka 1966 S04
            |           |--L. globosus Grobler 1995 S04
            |           |--L. longus (Balogh 1961) [=Pseudotocepheus longus] S04
            |           `--L. youngai (Mahunka 1984) [=Nesotocepheus youngai] S04
            |         Spinotocepheus Hammer 1981 S04
            |           |--*S. javensis Hammer 1981 S04
            |           |--S. foveolatus Hammer 1981 S04
            |           |--S. nigromaculatus Hammer 1981 S04
            |           `--S. tjibodensis Hammer 1981 S04
            |         Cavernocepheus Balogh & Mahunka 1969 S04
            |           |--*C. monstruosus Balogh & Mahunka 1969 S04
            |           |--C. acutus Balogh & Palacios-Vargas 1997 S04
            |           |--C. furcatus Balogh & Palacios-Vargas 1997 S04
            |           |--C. fusifer Balogh & Palacios-Vargas 1997 S04
            |           |--C. monticola Balogh 1989 S04
            |           |--C. obliquus Balogh 2002 S04
            |           `--C. undulatus Balogh 2002 S04
            |         Papillocepheus Balogh & Mahunka 1966 [incl. Clavazetes Hammer 1966] S04
            |           |--*P. heterotrichus Balogh & Mahunka 1966 S04
            |           |--P. areolatus Mahunka 1987 S04
            |           |--P. decoratus Mahunka 1994 S04
            |           |--P. decorus (Hammer 1966) S04 [=*Clavazetes decorus BB92]
            |           |--P. deficiens Balogh & Balogh 1983 S04
            |           |--P. neotropicus (Balogh 1988) [=Clavazetes neotropicus] S04
            |           |--P. reductus Mahunka 1994 S04
            |           `--P. tuberculatus (Mahunka 1978) [=Clavazetes tuberculatus] S04
            |         Pseudotocepheus Balogh 1961 S04
            |           |--P. (Pseudotocepheus) S04
            |           `--P. (Constrictocepheus Grobler 1998) S04
            |                |--P. (*C.) geminatus Balogh & Mahunka 1969 S04
            |                |--P. (C.) coarctatus Balogh 1985 S04
            |                |--P. (C.) curtiseta Hammer 1966 S04
            |                |--P. (C.) gobletus Chakrabarti & Mondal 1978 S04
            |                |--P. (C.) monteithi Balogh & Balogh 1983 S04
            |                |--P. (C.) orientalis Mondal & Kundu 1983 S04
            |                |--P. (C.) punctatus Hammer 1966 S04
            |                `--P. (C.) tenuiseta Hammer 1966 S04
            |         Plenotocepheus PL17
            |--Otocepheinae C-R79
            `--Tetracondylinae C-R79
                 |--Dolicheremaeus NB-P09
                 `--Leptotocepheus Balogh 1961 C-R79, S04
                      |--*L. trimucronatus Balogh 1961 S04
                      |--L. flagellifer Mahunka 1988 S04
                      |--L. macromucronatus Mahunka 1988 S04
                      `--L. orientalis Mahunka 1988 S04

Bathocepheus manguiati (Corpuz-Raros 1979) S04 [=Machadocepheus manguiati S04, *Philippobodes manguiati BB92]

*Type species of generic name indicated

REFERENCES

[BB92] Balogh, J., & P. Balogh. 1992. The Oribatid Mites Genera of the World vol. 1. Hungarian Natural History Museum: Budapest.

[CH98] Colloff, M. J., & R. B. Halliday. 1998. Oribatid Mites: A catalogue of Australian genera and species. CSIRO Publishing: Collingwood.

[C-R79] Corpuz-Raros, L. A. 1979. Philippine Oribatei (Acarina). I. Preliminary list of species and descriptions of forty new species. Philippine Agriculturist 62 (1): 1–82.

[NB-P09] Norton, R. A., & V. M. Behan-Pelletier. 2009. Suborder Oribatida. In: Krantz, G. W., & D. E. Walter (eds) A Manual of Acarology 3^rd ed. pp. 430–564. Texas Tech University Press.

[PL17] Pachl, P., A. C. Lindl, A. Krause, S. Scheu, I. Schaefer & M. Maraun. 2017. The tropics as an ancient cradle of oribatid mite diversity. Acarologia 57 (2): 309–322.

[SK10] Schäffer, S., S. Koblmüller, T. Pfingstl, C. Sturmbauer & G. Krisper. 2010. Ancestral state reconstruction reveals multiple independent evolution of diagnostic morphological characters in the "Higher Oribatida" (Acari), conflicting with current classification. BMC Evolutionary Biology 10: 246.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

Otocepheus (Acrotocepheus)

Dorsal (left) and ventral views of Otocepheus excelsus, from Norton & Behan-Pelletier (2009).

Belongs within: Otocepheidae.

The subgenus Acrotocepheus of the genus Otocepheus is a primarily southeast and southern Asian group of otocepheid oribatid mites characterised by the presence of median condyles on the prodorsum.

Characters (from Balogh & Balogh 1992): Median prodorsal condyles present. Notogaster with ten pairs of setae; lateral notogastral condyles narrowly spaced. Epimeral seta formula 3-1-3-3; apodemata I absent; pedotecta 2+3 conspicuously developed, axe-shaped in ventral view and tailfin-shaped in dorsal view, anterior and posterior expansions nearly equal in size. Four pairs of genital setae; one pair of aggenital setae. two pairs of anal setae; three pairs of adanal setae; fissura iad adanal.

<==Otocepheus (Acrotocepheus Aoki 1965) NB-P09, S04
    |--O. (*A.) quateorum Aoki 1965 S04
    |--‘Acrotocepheus’ (A.) besucheti Mahunka 1973 S04
    |--‘Acrotocepheus’ (A.) bucephalus Balogh 1970 S04
    |--‘Acrotocepheus’ (A.) burckhardti Mahunka 1987 S04
    |--O. (A.) caudatus Hammer 1981 S04
    |--‘Acrotocepheus’ (A.) consimilis Balogh 1970 S04
    |--‘Acrotocepheus’ (A.) curvisetiger Aoki 1984 S04
    |--‘Acrotocepheus’ (A.) diehli Mahunka 1989 S04
    |--O. (A.) duplicornutus Aoki 1965 S04
    |    |--O. d. duplicornutus S04
    |    `--‘Acrotocepheus’ d. discrepans Balogh & Mahunka 1967 S04
    |--O. (A.) excelsus Aoki 1965 NB-P09, S04
    |--‘Acrotocepheus’ (A.) gracilis Aoki 1973 S04
    |--‘Acrotocepheus’ (A.) grandis Mahunka 1973 S04
    |--O. (A.) heterotrichus Balogh & Mahunka 1968 S04
    |--O. (A.) holtmanni Aoki 1965 S04
    |-‘Acrotocepheus’. (A.) horakae Mahunka 1987 S04
    |--‘Acrotocepheus’ (A.) lienhardi Mahunka 1989 S04
    |--‘Acrotocepheus’ (A.) loebli Mahunka 1973 S04
    |--O. (A.) macrodentatus Hammer 1981 S04
    |--‘Acrotocepheus’ (A.) pangasuganensis Corpuz-Raros 1990 S04
    |--O. (A.) philippinensis Aoki 1965 S04
    |--‘Acrotocepheus’ (A.) surigaoensis Corpuz-Raros 1979 S04
    |--‘Acrotocepheus’ (A.) triplicicornutus Balogh & Mahunka 1967 S04
    |--‘Acrotocepheus’ (A.) tupasae Corpuz-Raros 1990 S04
    `--‘Acrotocepheus’ (A.) wallacei Mahunka 1989 S04

*Type species of generic name indicated

REFERENCES

Balogh, J., & P. Balogh. 1992. The Oribatid Mites Genera of the World vol. 1. Hungarian Natural History Museum: Budapest.

[NB-P09] Norton, R. A., & V. M. Behan-Pelletier. 2009. Suborder Oribatida. In: Krantz, G. W., & D. E. Walter (eds) A Manual of Acarology 3^rd ed. pp. 430–564. Texas Tech University Press.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

Nippobodidae

Dorsal view of Leobodes anulatus, from Norton & Behan-Pelletier (2009).

Belongs within: Circumdehiscentiae.

The Nippobodidae are a group of oribatid mites found in eastern Asia characterised by large projections on both the prodorsum and the anterior of the notogaster.

Characters (from Norton & Behan-Pelletier 2009): Length 600–750 µm. Prodorsum with large processes in middle portion; with true lamellae. Subcapitulum diarthric; chelicerae chelate-dentate; subcapitular mentum without tectum. Notogaster with large subtriangular or subrectangular humeral processes projecting dorsally over bothridium, interlocked or not with prodorsal processes; octotaxic system absent.

<==Nippobodidae
    |--Leobodes Aoki 1965 S04
    |    |--*L. mirabilis Aoki 1965 S04
    |    |--L. anulatus Aoki 1965 S04
    |    |--L. ligiangensis Aoki 2000 S04
    |    |--L. monstruosus Jeleva & Vu 1987 S04
    |    `--L. yinae Aoki 2000 S04
    `--Nippobodes Aoki 1959 S04
         |--*N. insolitus Aoki 1959 S04
         |--N. brevisetiger Aoki 1981 S04
         |--N. chejuensis Choi 1996 S04
         |--N. latus Aoki 1970 S04
         |--N. tamlaensis Choi 1996 S04
         |--N. tokaraensis Aoki 1989 S04
         `--N. yuwanensis Aoki 1984 [=N. brevisetiger yuwanensis] S04

*Type species of generic name indicated

REFERENCES

Norton, R. A., & V. M. Behan-Pelletier. 2009. Suborder Oribatida. In: Krantz, G. W., & D. E. Walter (eds) A Manual of Acarology 3rd ed. pp. 430–564. Texas Tech University Press.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

Carabodes (Klapperiches)

Carabodes chandleri, from Canadian Biodiversity Information Facility.

Belongs within: Carabodidae.

The subgenus Klapperiches of the genus Carabodes is a cosmopolitan group of carabodid oribatid mites with an unusually shortened ventral region.

Characters (from Balogh & Balogh 1992): Prodorsum and notogaster without high elevation(s) or deep excavation(s); notogaster with ten pairs of well-developed setae. Ventral region shortened, genital and anal plates near each other; genital plates with four pairs of setae; aggenital setae absent.

<==Carabodes (Klapperiches Mahunka 1979) S04
    |--C. (*K.) nigrosetosus (Mahunka 1979) [=*Klapperiches nigrosetosus] S04
    |--C. (K.) agenjoi Pérez-Íñigo 1969 S04
    |--C. (K.) andasibe Mahunka 1993 S04
    |--C. (K.) atrichosus Mahunka 1984 S04
    |--C. (K.) borhidii Balogh & Mahunka 1979 S04
    |--C. (K.) california Reeves & Behan-Pelletier 1998 S04
    |--C. (K.) chandleri Reeves 1992 S04
    |--C. (K.) coronatus Mahunka 1986 S04
    |--C. (K.) depilatus Balogh & Mahunka 1969 S04
    |--C. (K.) dickinsoni Reeves & Behan-Pelletier 1998 S04
    |--C. (K.) dissimilis Bernini 1976 S04
    |--C. (K.) ecuadoriensis Balogh 1988 S04
    |--C. (K.) excellens Balogh & Mahunka 1969 S04
    |--C. (K.) globiger Balogh 1970 S04
    |--C. (K.) granosus Sellnick 1959 S04
    |--C. (K.) higginsi Reeves 1988 S04
    |--C. (K.) jamaicaensis Woolley 1967 S04
    |--C. (K.) kusseri Balogh & Balogh 1983 S04
    |--C. (K.) littoristicus (Reeves 1997) [=Klapperiches littoristicus] S04
    |--C. (K.) luteoauratus Hammer 1972 S04
    |--C. (K.) manganoi Bernini 1976 S04
    |--C. (K.) manifera Hammer 1977 S04
    |--C. (K.) microtrichus Mahunka 1984 S04
    |--C. (K.) minusculus Berlese 1923 [incl. C. castrii Mahunka 1966] S04
    |--C. (K.) ornatissimus Hammer 1966 S04
    |--C. (K.) penicillatus Berlese 1916 S04
    |--C. (K.) penicillus (Mahunka 1998) [=Klapperiches penicillus] S04
    |--C. (K.) pocsi Mahunka 1983 S04
    |--C. (K.) poggii Bernini 1976 S04
    |--C. (K.) problematicus Mahunka 1985 S04
    |--C. (K.) pulcher Bernini 1976 S04
    |    |--C. p. pulcher S04
    |    `--C. p. occidentalis Pérez-Íñigo & Peña 1996 S04
    |--C. (K.) purpurarius Pérez-Íñigo & Peña 1996 S04
    |--C. (K.) radiatus Berlese 1916 [incl. C. dendroetus Reeves 1987] S04
    |--C. (K.) samoensis Balogh & Balogh 1986 S04
    |--C. (K.) schatzi Bernini 1976 S04
    |--C. (K.) similis Ruiz, Subías & Kahwash 1989 [incl. C. translamellatus Pérez-Íñigo 1990] S04
    |--C. (K.) strinovichi Balogh & Mahunka 1978 S04
    |--C. (K.) tenerifensis Pérez-Íñigo 1976 S04
    |--C. (K.) variabilis Hammer 1966 S04
    `--C. (K.) willmanni Bernini 1975 S04

*Type species of generic name indicated

REFERENCES

Balogh, J., & P. Balogh. 1992. The Oribatid Mites Genera of the World vol. 1. Hungarian Natural History Museum: Budapest.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

Carabodes (Carabodes)

Carabodes labyrinthicus, copyright Biodiversity Institute of Ontario.

Belongs within: Carabodinae.

The type subgenus of the genus Carabodes is a subcosmopolitan group of carabodid oribatid mites with an evenly arched notogaster. The type species, C. coriaceus, is widespread in forest soils in Europe with less certain records from other parts of the Holarctic region.

Characters (from Balogh & Balogh 1992): Prodorsum and notogaster without high elevation(s) or deep excavation(s); notogaster with ten pairs of well-developed setae. Ventral region not shortened, genital and anal plates normally separated. Genital plates with four setae; aggenital setae usually present.

Carabodes Koch 1835 (Carabodes) [incl. Neocepheus Willmann 1936] S04
    |--C. (C.) marginatus (Michael 1884) SK10, S04 [=Tegeocranus marginatus S04; incl. C. pontiger Berlese 1913 S04]
    `--+--C. femoralis group CA86
       |    |--C. (C.) femoralis (Nicolet 1855) SK10, S04 [=Tegeocranus femoralis S04]
       |    |--C. (C.) rimosus Aoki 1959 CA86
       |    |--C. (C.) rugosior Berlese 1916 CA86, S04 [=C. femoralis var. rugosior S04]
       |    |--C. (C.) transversarius Choi & Aoki 1986 CA86
       |    `--C. (C.) tsushimaensis Aoki 1970 CA86, S04 [=C. rimosus tsushimaensis S04]
       |         |--C. t. tsushimaensis S04
       |         `--C. t. ‘dorsalis’ Choi & Aoki 1986 non C. dorsalis Banks 1896 S04
       `--C. (C.) labyrinthicus (Michael 1879) SK10, S04 (see below for synonymy)

Carabodes (Carabodes) incertae sedis:
  *C. (C.) coriaceus Koch 1835 [incl. C. nepos Hull 1914, C. papillosus Lucas 1849] S04
  C. (C.) affinis Berlese 1913 S04
  C. (C.) apuanicus Bernini 1979 S04
  C. (C.) arduinii Valle 1955 S04
  C. (C.) areolatus Berlese 1916 S04
  C. (C.) auriculatus Mahunka 1987 S04
  C. (C.) azoricus Pérez-Íñigo & Pérez-Íñigo 1996 S04
  C. (C.) basilewskyi Balogh 1958 (n. d.) S04
  C. (C.) bellus Aoki 1959 S04
  C. (C.) berninii Mahunka 1983 S04
  C. (C.) bicolor Balogh 1958 (n. d.) S04
  C. (C.) brevis Banks 1896 S04
  C. (C.) cephalotes Koch 1836 (n. d.) [incl. C. cynocephalus Koch 1839] S04
  C. (C.) cerebrum Fujikawa 1993 S04
  C. (C.) cherokee Reeves 1993 S04
  C. (C.) chirstlus Mahunka 1987 [incl. C. foliatus Morell 1990] S04
  C. (C.) clavatus Warburton 1912 (n. d.) [=C. lalbyrinthicus clavatus] S04
  C. (C.) cochleaformis Reeves 1990 S04
  C. (C.) colorado Reeves & Behan-Pelletier 1998 S04
  C. (C.) comas Kulijev 1979 S04
  C. (C.) coweetaensis Reeves 1995 S04
  C. (C.) dudichi Mahunka & Mahunka-Papp 1999 S04
  C. (C.) egregius Djaparidze 1990 S04
  C. (C.) erectus Reeves 1992 S04
  C. (C.) falcatus Jacot 1937 S04
  C. (C.) floridus Berlese 1913 S04
  C. (C.) gibbiceps Berlese 1916 S04
  C. (C.) grandjeani Bernini 1977 S04
  C. (C.) granulatus Banks 1895 [incl. C. omo Jacot 1937] S04
  C. (C.) gregorioi Gil-Martín & Subías 1997 S04
  C. (C.) guadarramicus Subías & Arillo 2001 S04
  C. (C.) hispanicus Pérez-Íñigo 1966 S04
  C. (C.) hoh Reeves & Behan-Pelletier 1998 S04
  C. (C.) hummelincki (Willmann 1936) S04 [=*Neocepheus hummelincki CH98]
  C. (C.) hungaricus Balogh 1943 S04
  C. (C.) ikeharai Aoki 1988 S04
  C. (C.) interruptus Reeve 1992 S04
  C. (C.) magnus Kunst 1961 S04
  C. (C.) montanus Bernini 1979 S04
  C. (C.) nantahalaensis Reeves 1993 S04
  C. (C.) neonominatus Subías 2004 (see below for synonymy) S04
  C. (C.) niger Banks 1895 S04
  C. (C.) nitens Johnston 1847 (n. d.) S04
  C. (C.) ornatus Storkan 1925 [incl. C. forsslundi Sellnick 1953] S04
  C. (C.) palmifer Berlese 1904 [incl. C. peniculatus Aoki 1970] S04
  C. (C.) paraspinosus Kulijev 1968 S04
  C. (C.) pentasetosus Reeves 1992 S04
  C. (C.) perezinigoi Salinas 1971 S04
  C. (C.) phylliformis Reeves 1993 S04
  C. (C.) pirenaicus Subías & Arillo 2001 S04
  C. (C.) pirenensis Kunst 1961 S04
  C. (C.) polyporetes Reeves 1991 S04
  C. (C.) prunum Fujikawa 1993 S04
  C. (C.) quadrangulus Bernini 1979 [incl. C. flagellifer Mihelčič 1966] S04
  C. (C.) reticulatus Berlese 1913 [=C. coriaceus reticulatus] S04
  C. (C.) spiniformis Reeves 1995 S04
  C. (C.) spinosus Storkan 1925 S04
  C. (C.) subalpinus Thor 1937 S04
  C. (C.) subarcticus Trägardh 1902 S04 [=C. elongatus var. subarctica S22]
  C. (C.) subcarinatus Mahunka & Mahunka-Papp 2000 (see below for synonymy) S04
  C. (C.) tenuis Forsslund 1953 S04
    |--C. t. tenuis S04
    `--C. t. longisetosus Kulijev 1968 S04
  C. (C.) wettsteini Mahunka 1996 S04
  C. (C.) wonalancetanus Reeves 1990 S04

Carabodes (Carabodes) labyrinthicus (Michael 1879) SK10, S04 [=Tegeocranus labyrinthicus S04; incl. Cepheus heimi Oudemans 1903 S04, Carabodes marginepunctatus Trägardh 1902 S04, Ca. scymnus Hull 1914 S04, Ca. vermiculatus Berlese 1916 S04]

Carabodes (Carabodes) neonominatus Subías 2004 [=C. gibbiceps clavata Jacot 1938 non C. labyrinthicus clavatus Warburton 1912] S04

Carabodes (Carabodes) subcarinatus Mahunka & Mahunka-Papp 2000 [incl. C. carinatus Mahunka & Mahunka-Papp 1999 non Wallwork 1977] S04

*Type species of generic name indicated

REFERENCES

Balogh, J., & P. Balogh. 1992. The Oribatid Mites Genera of the World vol. 1. Hungarian Natural History Museum: Budapest.

[CA86] Choi, S. S., & J. Aoki. 1986. The oribatid mites (Acari: Cryptostigmata) of Korea (4). On the femoralis-group of the genus Carabodes C.L. Koch, 1836. Korean Journal of Entomology 16 (1): 27–32.

[CH98] Colloff, M. J., & R. B. Halliday. 1998. Oribatid Mites: A catalogue of Australian genera and species. CSIRO Publishing: Collingwood.

[S22] Schweizer, J. 1922. Beitrag zur Kenntnis der terrestrischen Milbenfauna der Schweiz. Verhandl. Naturf. Ges. Basel 33: 23–112, 4 pls.

[SK10] Schäffer, S., S. Koblmüller, T. Pfingstl, C. Sturmbauer & G. Krisper. 2010. Ancestral state reconstruction reveals multiple independent evolution of diagnostic morphological characters in the "Higher Oribatida" (Acari), conflicting with current classification. BMC Evolutionary Biology 10: 246.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

Last updated: 15 December 2018.

Austrocarabodes (Austrocarabodes)

Austrocarabodes elegans, photographed by S. E. Thorpe.

Belongs within: Carabodidae.

The type subgenus of the genus Austrocarabodes is a cosmopolitan group of generalised carabodid oribatid mites.

Characters (from Balogh & Balogh 1992): Body not noticeably elongate. Prodorsum with wide lamellae. Notogaster evenly arched; notogastral setae long; humeral region without setae. Genital plates with four pairs of setae.

<==Austrocarabodes Hammer 1966 (Austrocarabodes) S04
    |--*A. (A.) ensifer (Sellnick 1931) [=Carabodes ensifer] S04
    |--A. (A.) agalawatta Balogh 1988 S04
    |--A. (A.) agressor Balogh & Mahunka 1978 S04
    |--A. (A.) albidus (Balogh 1961) [=Carabodes albidus] S04
    |--A. (A.) alveolatus Hammer 1973 S04
    |--A. (A.) angulatus (Balogh 1958) (n. d.) [=Carabodes angulatus] S04
    |--A. (A.) arrogans Pérez-Íñigo 1967 S04
    |--A. (A.) australis (Balogh & Csiszár 1963) [=Carabodes australis] S04
    |--A. (A.) bacilliger Mahunka 1978 S04
    |--A. (A.) bellicosus Balogh 1988 S04
    |--A. (A.) boninensis (Aoki 1978) [=Carabodes australis boninensis] S04
    |--A. (A.) butiae Pérez-Íñigo & Sarasola 1998 S04
    |--A. (A.) cadeti Mahunka 1978 S04
    |--A. (A.) cellularis (Balogh 1962) [=Carabodes cellularis] S04
    |--A. (A.) corpulentus (Trägardh 1931) (n. d.) [=Cepheus corpulentus] S04
    |--A. (A.) costulatus (Balogh 1958) [=Carabodes costulatus] S04
    |--A. (A.) crenellatus Mahunka 1983 S04
    |--A. (A.) crinitus (Trägardh 1931) (n. d.) [=Cepheus crinitus] S04
    |--A. (A.) curvisetiger Aoki 1982 S04
    |--A. (A.) davisi (Balogh & Mahunka 1969) [=Carabodes davisi] S04
    |--A. (A.) elegans Hammer 1966 S04
    |--A. (A.) erectus Mahunka 1984 S04
    |--A. (A.) falcatus Hammer 1973 S04
    |--A. (A.) fenestratus (Wallwork 1977) [=Carabodes fenestratus] S04
    |--A. (A.) flabellifer Mahunka 1986 S04
    |--A. (A.) foliaceisetus Krivolutsky 1971 S04
    |--A. (A.) glabrus Mahunka 1982 S04
    |--A. (A.) gressitti Balogh & Mahunka 1978 S04
    |--A. (A.) haradai (Aoki 1978) [=Carabodes haradai] S04
    |--A. (A.) horridus (Wallwork 1977) [=Carabodes horridus] S04
    |--A. (A.) imperfectus (Sellnick 1959) S04 [=Carabodes imperfecta S59]
    |    |--A. i. imperfectus S04
    |    `--A. i. squamosus Hammer 1970 S04
    |--A. (A.) intermedius Ruiz, Subías & Kahwash 1989 S04
    |--A. (A.) latissimus Mahunka 2000 S04
    |--A. (A.) lepidus Aoki 1978 S04
    |--A. (A.) longulus (Balogh 1958) (n. d.) [=Carabodes longulus] S04
    |--A. (A.) lunaris (Balogh 1962) [=Carabodes lunaris] S04
    |--A. (A.) maculatus Hammer 1966 S04
    |--A. (A.) microlaminatus Mahunka 1983 S04
    |--A. (A.) mixtus Mahunka 1996 S04
    |--A. (A.) nodosus Hammer 1966 S04
    |--A. (A.) obsoletus (Berlese 1916) [=Carabodes obsoletus] S04
    |--A. (A.) ocellatus Mahunka 1987 S04
    |--A. (A.) picturatus Mahunka 1987 S04
    |--A. (A.) pinnatus Mahunka 1986 S04
    |--A. (A.) polytrichus Balogh & Mahunka 1978 S04
    |--A. (A.) pseudoreticulatus Covarrubias 1967 S04
    |--A. (A.) rimosus Mahunka 1987 S04
    |--A. (A.) rugosus Mahunka 1969 S04
    |--A. (A.) schauenbergi Mahunka 1978 S04
    |--A. (A.) schwartzi (Balogh & Mahunka 1969) [=Carabodes schwartzi] S04
    |--A. (A.) similis Mahunka 1978 S04
    |--A. (A.) sinuosociliatus Mahunka 1983 S04
    |--A. (A.) sordidus (Balogh 1958) (n. d.) [=Carabodes sordidus] S04
    |--A. (A.) spathulatus Mahunka 1978 S04
    |--A. (A.) sphaeroideus Mahunka 1978 S04
    |--A. (A.) sphaerula Balogh 1970 S04
    |--A. (A.) szentivanyi (Balogh & Mahunka 1967) [=Carabodes szentivanyi] S04
    |--A. (A.) tarandus Mahunka 1986 S04
    |--A. (A.) travei (Balogh & Csiszár 1963) [=Carabodes travei] S04
    |--A. (A.) vaucheri Mahunka 1984 S04
    `--A. (A.) verrucatus (Trägardh 1931) [=Carabodes verrucatus] S04

*Type species of generic name indicated

REFERENCES

[S59] Sellnick, M. 1959. Acarina from southeastern Polynesia—II (Oribatidae). Occasional Papers of the Bernice P. Bishop Museum 22 (9): 109–152.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

Odontocepheus (Odontocepheus)

(Left to right) right lateral, dorsal and ventral views of Odontocepheus elongatus, from Balogh & Balogh (1992 vol. 2).

Belongs within: Carabodidae.

The type subgenus of the genus Odontocepheus is a group of relatively elongate carabodid oribatid mites found in the Holarctic and Australian regions.

Characters (from Balogh & Balogh 1992): Body elongate, more than twice as long as wide. Prodorsum with wide lamellae; sensillus short with dilated head. Notogaster evenly arched; notogastral setae more or less evenly spaced; anterior margin of notogaster with four pairs of anteriorly directed setae; posteromarginal area with four pairs of setae, relatively short but otherwise similar to remaining notogastral setae. Genital plates with four pairs of setae.

<==Odontocepheus Berlese 1913 (Odontocepheus) S04
    |--*O. (O.) elongatus (Michael 1879) [=Tegeocranus elongatus; incl. Hermannia oblonga Karpelles 1893] S04
    |--O. (O.) bandae Subías & Arillo 2001 S04
    |--O. (O.) beijingensis Wang 1997 S04
    |--O. (O.) curtiseta Ruiz, Subías & Kahwash 1989 S04
    |--O. (O.) espatulatus Saloña & Iturrondobeitia 1989 S04
    |--O. (O.) hungaricus Mahunka 1991 S04
    |--O. (O.) immarginatus Jacot 1934 S04
    |--O. (O.) laeviusculus (Canestrini & Fanzago 1877) (n. d.) [=Claviceps laeviusculus] S04
    |--O. (O.) oblongus (Banks 1895) [=Carabodes oblongus] S04
    |--O. (O.) oglasae Baratti & Bernini 1994 S04
    |--O. (O.) piramidalis Mahunka 1987 S04
    |--O. (O.) rumbleseatus Reeves 1995 S04
    |--O. (O.) villosus Mahunka 1987 S04
    `--O. (O.) zaballosi Subías & Arillo 2001 S04

*Type species of generic name indicated

REFERENCES

Balogh, J., & P. Balogh. 1992. The Oribatid Mites Genera of the World vol. 1. Hungarian Natural History Museum: Budapest.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

Gibbicepheus (Gibbicepheus)

Gibbicepheus frondosus, from here.

Belongs within: Carabodidae.

The type subgenus of the genus Gibbicepheus is a primarily pantropical group of carabodid oribatid mites with four rows of large tubercles on the notogaster.

Characters (from Balogh & Balogh 1992): Prodorsum with wide lamellae. Notogaster with large tubercles aligned in four longitudinal rows; humeral setae absent; medial setae widely distributed, about equidistant; posteromarginal region with four pairs of setae. Genital plates with four pairs of setae.

<==Gibbicepheus Aoki 1958 (Gibbicepheus) S04
    |--*G. (G.) elevatus Balogh 1958 S04
    |--G. (G.) austroamericanus Mahunka 1984 S04
    |--G. (G.) baccanensis Jeleva & Hu 1987 S04
    |--G. (G.) chinensis Wen 1992 S04
    |--G. (G.) ensifer Balogh 1958 (n. d.) S04
    |--G. (G.) fenestralis Hammer 1979 S04
    |--G. (G.) frondosus (Aoki 1959) [=Diplobodes frondosus] S04
    |--G. (G.) latohumeralis Hammer 1982 S04
    |--G. (G.) micheli Mahunka 1978 S04
    |--G. (G.) novus Hammer 1973 S04
    |--G. (G.) rugosus Mahunka 1978 S04
    |--G. (G.) sisri Sanyal 1990 S04
    |--G. (G.) tuberculatus Balogh 1970 S04
    |--G. (G.) ventrostriatus Hammer 1979 S04
    `--G. (G.) waterhousei Balogh & Mahunka 1978 S04

*Type species of generic name indicated

REFERENCES

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

Pseudotocepheus (Pseudotocepheus)

Dorsal (left) and ventral views of Pseudotocepheus sturmi, from Balogh & Balogh (1992 vol. 2).

Belongs within: Otocepheidae.

The type subgenus of the genus Pseudotocepheus is a pantropical and southern subtropical group of fairly generalised otocepheid oribatid mites.

Characters (from Balogh & Balogh 1992): Notogaster with ten pairs of setae. Epimeral setal formula 3-1-3-3. Three pairs of genital setae; one pair of aggenital setae; two pairs of anal setae; three pairs of adanal setae. Fissure iad apoanal.

<==Pseudotocepheus Balogh 1961 (Pseudotocepheus) [incl. Nesotocepheus Hammer 1972] S04
    |--*P. (P.) pauliani Balogh 1961 S04
    |--P. (P.) amonstruosus Mahunka 1973 (see below for synonymy) S04
    |--P. (P.) andinus (Fernández, Martínez & Eguaras 1990) [=Nesotocepheus andinus] S04
    |--P. (P.) aokii Grobler 1997 S04
    |--P. (P.) asymmetricus Balogh & Balogh 1983 S04
    |--P. (P.) australis (Mahunka 1980) [=Nesotocepheus australis] S04
    |--P. (P.) bacilliger Balogh & Balogh 1983 S04
    |--P. (P.) clavigerus (Mahunka 1978) [=Nesotocepheus clavigerus] S04
    |--P. (P.) contractus Grobler 1997 S04
    |--P. (P.) curtipilus (Trägardh 1931) [=Odontocepheus curtipilus] S04
    |--P. (P.) dentatus Grobler 1998 S04
    |--P. (P.) flagellatus (Mahunka 1987) [=Plenotocepheus flagellatus] S04
    |--P. (P.) foveolatus Hammer 1966 S04
    |--P. (P.) granulatus Mahunka 1985 S04
    |--P. (P.) hammerae Chakrabarti & Kundu 1978 S04
    |--P. (P.) hauseri (Mahunka 1980) [=Nesotocepheus hauseri] S04
    |--P. (P.) lanceolatus Grobler 1998 S04
    |--P. (P.) lienhardi Mahunka 1993 S04
    |--P. (P.) medius Balogh & Mahunka 1966 S04
    |--P. (P.) neonominatus Subías 2004 [=P. longus Mahunka 1973 non Balogh 1961] S04
    |--P. (P.) parallelus Balogh & Balogh 1983 S04
    |--P. (P.) pauliensis Pérez-Íñigo & Baggio 1993 S04
    |--P. (P.) promissus Grobler 1998 S04
    |--P. (P.) pygmaeus Balogh 1962 S04
    |--P. (P.) radiatus Hammer 1973 S04
    |--P. (P.) setiger (Hammer 1972) [=Nesotocepheus setiger] S04
    |--P. (P.) sexdentatus (Trägardh 1931) [=Odontocepheus sexdentatus] S04
    |--P. (P.) sexdimorphus (Vasiliu & Calugar 1977) [=Nesotocepheus sexdimorphus] S04
    |--P. (P.) shauni Grobler 1997 S04
    |--P. (P.) sturmi Balogh 1984 S04
    |--P. (P.) szentivanyorum Balogh & Mahunka 1978 S04
    |--P. (P.) tolanaro Mahunka 1997 S04
    `--P. (P.) vicarius Balogh & Mahunka 1978 S04

Pseudotocepheus (Pseudotocepheus) amonstruosus Mahunka 1973 [incl. P. septemtuberculatus Balogh & Mahunka 1978, P. simplex Pérez-Íñigo & Baggio 1980, P. transversalis Mahunka 1978] S04

*Type species of generic name indicated

REFERENCES

Balogh, J., & P. Balogh. 1992. The Oribatid Mites Genera of the World vol. 1. Hungarian Natural History Museum: Budapest.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

Fissicepheus (Fissicepheus)

Fissicepheus clavatus, from here.

Belongs within: Otocepheidae.

The type subgenus of the genus Fissicepheus is an Eurasian group of oribatid mites possessing condyles in the aggenital region.

Characters (from Balogh & Balogh 1992): Notogaster with ten pairs of setae. Epimeral setal formula 2-0-1-3. Pedotecta 2+3 distinct but not conspicuously developed. Four pairs of genital setae; aggenital condyles present; one pair of aggenital setae; two pairs of anal setae; three pairs of adanal setae. Fissures iad apoanal, far from anal plates, distance ad3–ad3 about same length as iad–iad.

<==Fissicepheus Balogh & Mahunka 1967 (Fissicepheus) S04
    |--*F. (F.) elegans Balogh & Mahunka 1967 S04
    |--F. (F.) chinensis Wen 1993 S04
    |--F. (F.) clavatus (Aoki 1959) [=Tetracondyla clavata] S04
    |--F. (F.) claviopsis Mahunka 1971 S04
    |--F. (F.) coronarius Aoki 1967 S04
    |    |--F. c. coronaris S04
    |    `--F. c. koreensis Mahunka 1971 S04
    |--F. (F.) curvisetosus Kubota 2001 S04
    |--F. (F.) elegans Balogh & Mahunka 1967 S04
    |--F. (F.) mitis Aoki 1970 S04
    |--F. (F.) mitratus Mahunka 1971 S04
    |--F. (F.) nakanei Aoki 1986 S04
    |--F. (F.) ornithorhynchus Wen 1993 S04
    |--F. (F.) sitnikovae Sergienko 1976 S04
    |--F. (F.) steinmanni Mahunka 1971 S04
    |--F. (F.) subclavatus Mahunak 1971 S04
    `--F. (F.) vicinus Aoki 1986 S04

*Type species of generic name indicated

REFERENCES

Balogh, J., & P. Balogh. 1992. The Oribatid Mites Genera of the World vol. 1. Hungarian Natural History Museum: Budapest.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

Dampfiellidae

Dorsum, venter and leg I of Dampfiella cejansis, from Norton & Behan-Pelletier (2009).

Belongs within: Circumdehiscentiae.

The Dampfiellidae are a pantropical group of oribatid mites that are diverse in lowland rain forests (Norton & Behan-Pelletier 2009).

Characters (from Norton & Behan-Pelletier 2009): Length 600–1400 µm. Prodorsum with costulae; porelike alveolar vestige of second exobothridial seta present. Notogaster narrow, about twice as long as wide, with strong depression laterally between setae c and la. Genital plates with three or four pairs of setae. Tarsi and tibiae functionally fused, moving as single segment.

<==Dampfiellidae S04
    |--Dampfiella Sellnick 1931 S04
    |    |--*D. procera Sellnick 1931 S04
    |    |--D. ambigua Pérez-Íñigo 1976 S04
    |    |--D. angusta Hammer 1979 S04
    |    |--D. dubia Hammer 1971 S04
    |    |--D. euaensis Hammer 1973 S04
    |    |--D. foliata Mahunka 1974 S04
    |    |--D. jandai Starý 1993 S04
    |    |--D. kinabalu Mahunka 2000 S04
    |    |--D. luzonica Corpuz-Raros 1997 S04
    |    |--D. mahunkai Starý 1993 S04
    |    |--D. nebulosa Mahunka 2000 S04
    |    |--D. ovalis Corpuz-Raros 1997 S04
    |    |--D. papuana Balogh & Balogh 1986 S04
    |    |--D. paratina (Mahunka 1983) [=Beckiella paratina] S04
    |    |--D. peseki Starý 1993 S04
    |    |--D. philippica Corpuz-Raros 1997 S04
    |    |--D. prostata Aoki 1965 S04
    |    |--D. putinglupa Corpuz-Raros 1997 S04
    |    |--D. sepilok Mahunka 2000 S04
    |    |--D. setosa Mahunka 1984 S04
    |    |--D. similis Hammer 1971 S04
    |    |--D. similoides Corpuz-Raros 1997 S04
    |    |--D. tupi Pérez-Íñigo & Baggio 1989 S04
    |    |--D. vanalinearis Corpuz-Raros 1997 S04
    |    `--D. zellwegeri Mahunka 1997 S04
    `--Beckiella Grandjean 1964 S04
         |--*B. cejansis (Beck 1962) [=Dampfiella cejansis] S04
         |--B. acuta Balogh & Mahunka 1978 S04
         |--B. africana (Balogh 1958) [=Dampfiella africana] S04
         |--B. arcta Pérez-Íñigo & Baggio 1986 S04
         |--B. bloszyki Starý 1989 S04
         |--B. borhidii Balogh & Mahunka 1978 S04
         |--B. bucephala Balogh & Mahunka 1978 S04
         |--B. capitulum Balogh & Mahunka 1978 S04
         |--B. carinata (Beck 1962) [=Dampfiella carinata] S04
         |--B. clavata Mahunka 1988 S04
         |--B. cubana Starý 1989 S04
         |--B. deficiens Balogh & Mahunka 1978 S04
         |--B. discoidalis Balogh & Mahunka 1978 S04
         |--B. duplicata Balogh & Mahunka 1978 S04
         |--B. elongata Balogh & Mahunka 1969 S04
         |--B. foveolata Balogh & Mahunka 1969 S04
         |--B. fratercula Balogh & Mahunka 1978 S04
         |--B. garciai Balogh & Mahunka 1979 S04
         |--B. inca Starý 1992 S04
         |--B. interlamellaris Balogh & Mahunka 1978 S04
         |--B. irmayi Balogh & Mahunka 1969 S04
         |--B. lamellata Balogh & Mahunka 1969 S04
         |--B. latirostris Starý 1992 S04
         |--B. microseta Balogh & Mahunka 1979 S04
         |--B. opposita Mahunka 1982 S04
         |--B. recta Balogh & Mahunka 1978 S04
         |--B. reticulofemorata Balogh & Mahunka 1979 S04
         |--B. sellnicki (Hammer 1961) [=Dampfiella sellnicki] S04
         |--B. silvai Balogh & Mahunka 1979 S04
         |--B. synlamellata Balogh & Mahunka 1974 S04
         `--B. vitiosa Mahunka 1985 S04

*Type species of generic name indicated

REFERENCES

Norton, R. A., & V. M. Behan-Pelletier. 2009. Suborder Oribatida. In: Krantz, G. W., & D. E. Walter (eds) A Manual of Acarology 3rd ed. pp. 430–564. Texas Tech University Press.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.