Tenthredo

Tenthredo mesomela, copyright Hectonichus.

Belongs within: Tenthredinidae.

Tenthredo is a diverse genus of often colourful sawflies widespread in Eurasia and North America.

Tenthredo Linnaeus 1758 [incl. Tenthredella Rohwer 1910; Tenthredininae] H04
    |--T. abietis Linnaeus 1758 L58
    |--T. allantosikkimensis Haris 2004 H04
    |--T. alni Linnaeus 1758 L58
    |--T. americana Linnaeus 1758 L58
    |--T. amerinae Linnaeus 1758 L58
    |--T. atra Linnaeus 1758 L58
    |--T. betulae Linnaeus 1758 L58
    |--T. campestris Linnaeus 1758 L58
    |--T. capreae Linnaeus 1758 L58
    |--T. cerasi Linnaeus 1758 L58
    |--T. changbuensis Haris 2000 H04
    |--T. cincta Linnaeus 1758 L58
    |--T. coloripleuris Malaise 1945 H04
    |--T. cynosbati Linnaeus 1758 L58
    |--T. duodecimpunctata Linnaeus 1758 L58
    |--T. erythrocephala Linnaeus 1758 L58
    |--T. faltuum Linnaeus 1758 L58
    |--T. fasciata Linnaeus 1758 L58
    |--T. femorata Linnaeus 1758 L58
    |--T. flava MS01
    |--T. fulva MS01
    |--T. germanica L02
    |--T. gonagra L02
    |--T. intercus Linnaeus 1758 L58
    |--T. juniperi Linnaeus 1758 L58
    |--T. koehleri PK17
    |--T. livida Linnaeus 1758 L58
    |--T. lonicerae Linnaeus 1758 L58
    |--T. lucorum Linnaeus 1758 L58
    |--T. maculipennis Malaise 1945 H04
    |--T. maruana Malaise 1945 H04
    |--T. memoriaescalerai Haris 2004 H04
    |--T. mesomela Linnaeus 1758 L58
    |--T. minutosimplicis Haris 2004 H04
    |--T. moniliata MS01
    |--T. nemoralis Linnaeus 1758 L58
    |--T. nigroypsilon Haris 2004 H04
    |--T. nitens Linnaeus 1758 L58
    |--T. ovata L02
    |--T. pavida L02
    |--T. pratensis Linnaeus 1758 L58
    |--T. pruni Linnaeus 1758 L58
    |--T. reticulata Linnaeus 1758 L58
    |--T. rufipennis MS01
    |--T. rufipes Linnaeus 1758 L58
    |--T. rumicis Linnaeus 1758 L58
    |--T. rustica Linnaeus 1758 L58
    |--T. salicis Linnaeus 1758 L58
    |--T. scrophulariae Linnaeus 1758 L58
    |--T. septentrionalis Linnaeus 1758 L58
    |--T. ulmi Linnaeus 1758 L58
    |--T. ustulata Linnaeus 1758 L58
    |--T. variicolor Malaise 1945 H04
    `--T. viridis Linnaeus 1758 L58

*Type species of generic name indicated

REFERENCES

[H04] Haris, A. 2004. Four new Tenthredo Linnaeus, 1758 species from Sikkim (Hymenoptera, Tenthredinidae). Graellsia 60 (2): 155–161.

[L02] Latreille, P. A. 1802. Histoire Naturelle, générale et particulière des crustacés et des insectes vol. 3. Familles naturelles des genres. F. Dufart: Paris.

[L58] Linnaeus, C. 1758. Systema Naturae per Regna Tria Naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata. Laurentii Salvii: Holmiae.

[MS01] Mocsáry, A., & V. Szépligeti. 1901. Hymenopterák [Hymenopteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 121–169. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[PK17] Peters, R. S., L. Krogmann, C. Mayer, A. Donath, S. Gunkel, K. Meusemann, A. Kozlov, L. Podsiadlowski, M. Petersen, R. Lanfear, P. A. Diez, J. Heraty, K. M. Kjer, S. Klopfstein, R. Meier, C. Polidori, T. Schmitt, S. Liu, X. Zhou, T. Wappler, J. Rust, B. Misof & O. Niehuis. 2017. Evolutionary history of the Hymenoptera. Current Biology 27 (7): 1013–1018.

Dolerus

Dolerus varispinus, copyright James K. Lindsey.

Belongs within: Tenthredinidae.

Dolerus is a genus of sawflies found in northern temperate regions with larvae mostly feeding on grasses and horsetails.

Dolerus [Dolerinae] H04
    |--D. ciliatus MS01
    |--D. dubius [incl. D. dubius var. abietis] MS01
    |--D. fissus MS01
    |--D. haematodes MS01
    |--D. palmatus MS01
    |--D. palustris [incl. D. palustris var. saxatilis] MS01
    |--D. picipes MS01
    |--D. pratensis MS01
    |--D. pratorum MS01
    |--D. thoracicus MS01
    `--D. varispinus Hartig 1837 H04

*Type species of generic name indicated

REFERENCES

[H04] Haris, A. 2004. New sawflies from Spain (Hymenoptera, Tenthredinidae). Graellsia 60 (2): 163–165.

[MS01] Mocsáry, A., & V. Szépligeti. 1901. Hymenopterák [Hymenopteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 121–169. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

Ellampus

Ellampus auratus, copyright Kurt Kulac.

Belongs within: Chrysididae.

Ellampus is a genus of small, stoutly built cuckoo wasps with the apical margin of the abdomen truncate or subtruncate and medially incised (Bingham 1903).

Characters (from Bingham 1903): Head transverse, as broad as pronotum or broader, facial cavity subtriangular, arch above narrowing distinctly, moderately deep; antennae slender, cylindrical. Thorax broad, very convex above; pronotum long or short, often shorter than mesonotum, transverse; metanotum and scutellum convex, the former sometimes gibbous anteriorly; mesopleurae convex or flat, apex beneath more or less rounded; postscutellum convex, gibbous or with a mucro at base; posterior lateral angles of median segment produced, apex acute or obtuse. Fore wing with medial cell only complete, first submedial and sometimes radial cell almost complete, generally two-thirds of apical portion of radial nervure obliterated, occasionally second submedial cell faintly outlined; nervures and cells of hind wing obsolete; legs slender, posterior tibiae not incrassate; claws of tarsi multidentate. Abdomen very convex above, base transverse, apical margin rounded at sides, truncate and incised in middle.

<==Ellampus Spin. 1806 [=Elampus; incl. Notozus Först. 1853, Omalus Dahlb. 1854] B03
    |--*E. auratus B03 [=Omalus auratus BM76; incl. E. auratus var. maculatus MS01, E. auratus var. virescens MS01]
    |--‘Omalus’ aeneus BM76
    |--E. chlorosoma Lucas 1849 E12
    |--‘Omalus’ coeruleus BM76
    |--E. constrictus [=Omalus constrictus] BM76
    |--‘Omalus’ cressoni BM76
    |--‘Omalus’ glomeratus BM76
    |--‘Omalus’ interandinus BM76
    |--‘Omalus’ intermedius BM76
    |--‘Omalus’ iridescens BM76
    |--‘Omalus’ janus BM76
    |--E. kashmirensis (Nurse 1902) [=Notozus kashmirensis] B03
    |--E. panzeri BM76
    |--‘Omalus’ punctatus BM76
    |--‘Omalus’ purpuratus BM76
    |--‘Omalus’ pusillus BM76
    |--‘Omalus’ sinuosus BM76
    |--E. spina BM76
    |--E. timidus Nurse 1902 B03
    |--‘Omalus’ trilobatus BM76
    |--‘Omalus’ variatus BM76
    |--‘Omalus’ violaceus BM76
    `--E. viridicyaneus BM76

*Type species of generic name indicated

REFERENCES

[B03] Bingham, C. T. 1903. The Fauna of British India, including Ceylon and Burma. Hymenoptera vol. 2. Ants and Cuckoo-Wasps. Taylor and Francis: London.

[BM76] Bohart, R. M., & A. S. Menke. 1976. Sphecid Wasps of the World. University of California Press: Berkeley.

[E12] Evenhuis, N. L. 2012. Publication and dating of the Exploration Scientifique de l'Algérie: Histoire Naturelle des Animaux Articulés (1846–1849) by Pierre Hippolyte Lucas. Zootaxa 3448: 1–61.

[MS01] Mocsáry, A., & V. Szépligeti. 1901. Hymenopterák [Hymenopteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 121–169. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

Chrysis

Chrysis ignita, copyright Louise Hislop.

Belongs within: Chrysididae.

Chrysis is a genus of cuckoo-wasps parasitic on fossorial and other wasps, many species of which have prominent teeth on the apical margin of the abdomen (Bingham 1903).

Characters (from Bingham 1903): Female with head varying in shape, triangular or subtriangular, rounded and convex above, occasionally globose, sometimes broader than long, rarely quadrate; mandibles pointed at apex or bidentate; clypeus generally broad, often subporrect medially or emarginate anteriorly; face up to front below anterior ocellus and between eyes generally hollow, forming facial cavity in which antennae are folded at rest; antennae of male and female simple, generally cylindrical; scape moderately long; front more or less convex, sometimes slightly hollow around anterior ocellus; ocelli always present; eyes proportionately rather large, lateral; cheeks below and sides of head behind eyes generally short and restricted; vertex more or less convex; occiput transverse or emarginate, occasionally rounded. Thorax with prothorax short or moderately long, posterior lateral angles prolonged slightly towards base of wings; pronotum above more or less transversely rectangular, convex, sides parallel or slightly convergent anteriorly; propleurae generally more or less excavate; mesonotum and scutellum distinct; mesopleurae generally rugose, very often obliquely grooved, groove formed by foveate distinct depressions; apex acute or rounded, sometimes bidentate, often excavate and margined; postscutellum well developed, most often convex, rounded posteriorly, sometimes with mucro projecting backwards from base; median segment always more or less suppressed beneath postscutellum, posterior lateral angles always produced, flattened and usually acute at apex. Wings with few nervures and cells; forewing with costal, medial and first discoidal cells complete, first submedial and radial cells sometimes complete, sometimes incomplete, second submedial and first cubital incomplete, costal cell always very narrow, almost obliterated, first cubital generally entirely open at apex, rarely the transverse cubital nervure closing it is faintly outlined; hind wing with neuration still more incomplete, mere vestiges of cells and nervures being faintly outlined. Legs feeble, unarmed; femora and tibiae normal; claws simple, not dentate. Abdomen pseudosessile, more or less convex above, very variable in proportion of length to breadth; sides may be parallel or convergent posteriorly; base concave, transverse, or with one or more impressions above; second segment generally the longest, very often medially longitudinally carinate; third segment often transversely incrassate in middle, with anteapical row of foveae present as a rule, exceptionally absent or obliterated; apical margin with or without teeth.

<==Chrysis Linn. 1767 [incl. Pyria Lepel. & Serv. 1825] B03
    |--*C. cyanea B03
    |--C. abuensis Nurse 1902 B03
    |--C. affinis Lucas 1849 E12
    |--C. agilis B85
    |--C. amoenula (Mocsáry 1899) [=Pentachrysis amoenula] B03
    |--C. angustata (Mocsáry 1892) [=Tetrachrysis angustata] B03
    |--C. annulata du Buyss. 1887 (see below for synonymy) B03
    |--C. apiata du Buyss. 1900 [incl. C. lucinda Nurse 1903] B03
    |--C. arrestans Nurse 1903 B03
    |--C. ausae Bohart 1985 B85
    |--C. australis Bohart 1985 B85
    |--C. autocrata Nurse 1903 B03
    |--C. bahadur Nurse 1903 B03
    |--C. barbara Lucas 1849 E12
    |--C. bayadera du Buysson 1896 B03
    |--C. bengalensis (Mocsáry 1889) [=Pentachrysis bengalensis] B03
    |--C. bhavanae Bingham 1903 B03
    |--C. bipartita B85
    |--C. blanchardi Lucas 1849 E12
    |--C. burmanica Mocsáry 1892 B03
    |--C. cembricola HR11
    |--C. chavanae Nurse 1902 B03
    |--C. cirtana Lucas 1849 E12
    |--C. comottii Grib. 1884 B03
    |--C. conserta du Buyss. 1891 B03
    |--C. cotesi du Buysson 1893 B03
    |--C. crictovallensis B85
    |--C. cupreiventris Bingh. 1898 B03
    |--C. curtisensis Linsenmaier 1982 B85
    |--C. danae Bingham 1903 B03
    |--C. dentifrontis Linsenmaier 1982 B85
    |--C. disparilis Cameron 1897 B03
    |--C. dissimilanda du Buysson 1896 B03
    |--C. dives Lucas 1849 E12
    |--C. durga Bingham 1903 B03
    |--C. elizabethae Bingham 1903 B03
    |--C. feana (Mosc. 1892) [=Hexachrysis feana] B03
    |--C. festina B85
    |--C. fossulata B85
    |--C. fulgida MS01
    |--C. furcifera Bingham 1903 B03
    |--C. furiosa Cameron 1897 B03
    |--C. fuscipennis Brullé 1846 [=Tetrachrysis fuscipennis] B03
    |--C. gracilenta (Mocsáry 1889) [=Chrysis gracilenta] B03
    |--C. greeni Bingham 1903 B03
    |--C. gujaratica Nurse 1903 B03
    |--C. hedychroides Bingham 1903 B03
    |--C. ignifascia Mocsáry 1892 B03
    |--C. ignita BM76 [incl. C. ignita var. longula MS01]
    |--C. imperiosa Smith 1874 [=Pentachrysis imperiosa] B03
    |--C. impostor B85
    |--C. indica (Mocsáry 1889) [=Hexachrysis indica] B03
    |--C. indogotea Duf. & Perr. 1840 [=Tetrachrysis indogotea] B03
    |--C. insignis Lucas 1849 E12
    |--C. interceptor B85
    |--C. intrudens B85
    |--C. ionophris (Mocsáry 1892) [=Tetrachrysis ionophris] B03
    |--C. jalala Nurse 1902 B03
    |--C. kashmirensis Nurse 1902 B03
    |--C. lanka Bingham 1903 B03
    |--C. leachi BM76
    |--C. libita du Buysson 1896 B03
    |--C. lincea B85
    |--C. lusca Fabr. 1804 [=Pentachrysis lusca; incl. C. lusca var. concinna] B03
    |--C. mendicalis Cameron 1897 B03
    |--C. microtrema (n. d.) B85
    |--C. nila Bingham 1903 B03
    |--C. norsemanae Bohart 1985 B85
    |--C. notidana Bohart 1985 B85
    |--C. nugatrix Bingham 1903 B03
    |--C. nursei Bingham 1903 B03
    |--C. obliterata (Mocsáry 1889) [=Tetrachrysis obliterata] B03
    |--C. obscura Smith 1861 [=Hexachrysis obscura] B03
    |--C. oculata Fabr. 1775 [=Hexachrysis oculata, Pyria oculata; incl. C. oculata var. sumptuosa] B03
    |--C. oraniensis Le Peletier 1849 E12
    |--C. orientalis (Guér. 1842) [=Pyria orientalis, Hexachrysis orientalis] B03
    |--C. paria Bingham 1903 B03
    |--C. perfecta Cameron 1897 [incl. C. hoggei Nurse 1903] B03
    |--C. perplexa B85
    |--C. perthensis B85
    |--C. principalis Smith 1874 [=Hexachrysis principalis] B03
    |--C. quaerita Nurse 1902 B03
    |--C. rufa [incl. C. neglecta] MS01
    |--C. saginata B85
    |--C. sandaracata Bingham 1903 B03
    |--C. saraksensis Radoszk. 1891 [=C. seraxensis] B03
    |--C. schioedtei Dahlb. 1854 (see below for synonymy) B03
    |--C. scioensis Grib. 1879 [=Trichrysis scioensis] B03
    |--C. shanghaiensis Smith 1874 [incl. Pentachrysis crassiscuta Mocsáry 1889, P. himalayensis Mocsáry 1889] B03
    |--C. simulans B85
    |--C. singalensis (Mocsáry 1889) [=Trichrysis singalensis] B03
    |--C. sollicita B85
    |--C. speculata du Buysson 1896 B03
    |--C. succincta BM76
    |--C. tafnensis Le Peletier 1849 E12
    |--C. tasmaniaca B85
    |--C. thalia Nurse 1903 B03
    |--C. triacantha (Mocsáry 1889) [=Trichrysis triacantha] B03
    |--C. tricolor Lucas 1849 E12
    |--C. unicolor Lucas 1849 E12
    |--C. versicolor Lucas 1849 E12
    |--C. viridula PK17
    |--C. wroughtoni du Buysson 1896 B03
    |--C. xysa Bohart 1985 B85
    |--C. yallingupia B85
    `--C. zylla Bohart 1985 B85

Chrysis annulata du Buyss. 1887 [=Tetrachrysis annulata; incl. C. balucha Nurse 1903, T. fulvicornis Mocs. 1889, C. fulvicornis, C. quettaensis Nurse 1903] B03

Chrysis schioedtei Dahlb. 1854 [=Hexachrysis schioedtei; incl. C. gemmatus Smith 1874, Hexachrysis siva Mocsáry 1889, C. siva] B03

*Type species of generic name indicated

REFERENCES

[B03] Bingham, C. T. 1903. The Fauna of British India, including Ceylon and Burma. Hymenoptera vol. 2. Ants and Cuckoo-Wasps. Taylor and Francis: London.

[B85] Bohart, R. M. 1985. A key to the Australian Chrysis with description of new species (Hymenoptera: Chrysididae). Insecta Mundi 1 (1): 47–52.

[BM76] Bohart, R. M., & A. S. Menke. 1976. Sphecid Wasps of the World. University of California Press: Berkeley.

[E12] Evenhuis, N. L. 2012. Publication and dating of the Exploration Scientifique de l'Algérie: Histoire Naturelle des Animaux Articulés (1846–1849) by Pierre Hippolyte Lucas. Zootaxa 3448: 1–61.

[HR11] Heraty, J., F. Ronquist, J. M. Carpenter, D. Hawks, S. Schulmeister, A. P. Dowling, D. Murray, J. Munro, W. C. Wheeler, N. Schiff & M. Sharkey. 2011. Evolution of the hymenopteran megaradiation. Molecular Phylogenetics and Evolution 60: 73–88.

[MS01] Mocsáry, A., & V. Szépligeti. 1901. Hymenopterák [Hymenopteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 121–169. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[PK17] Peters, R. S., L. Krogmann, C. Mayer, A. Donath, S. Gunkel, K. Meusemann, A. Kozlov, L. Podsiadlowski, M. Petersen, R. Lanfear, P. A. Diez, J. Heraty, K. M. Kjer, S. Klopfstein, R. Meier, C. Polidori, T. Schmitt, S. Liu, X. Zhou, T. Wappler, J. Rust, B. Misof & O. Niehuis. 2017. Evolutionary history of the Hymenoptera. Current Biology 27 (7): 1013–1018.

Lasius

Lasius neoniger, copyright Tom Murray.

Belongs within: Lasiini.

Lasius, moisture ants, is a genus of largely nocturnally foraging ants that is widespread in temperate regions of the Northern Hemisphere.

Characters (from Bingham 1903; Wheeler & Wheeler 1990): Worker: Mandibles short, elongate, triangular, masticatory margin dentate and somewhat oblique, the apical tooth long, curved and acute; clypeus trapeziform, convex from side to side, its anterior margin rounded, posterior margin distinct; frontal area not distinctly limited, about twice as broad as high; antennal carinae short, wide apart, parallel and vertical; antennae 12-segmented, originating close to rear margin of clypeus, scape surpassing occipital margin by less than one-third its length, often much shorter; eyes of moderate size; ocelli present, but not distinct, very small. Thorax with pronotum rather short, strongly rounded, mesonotum broad, convex; metanotum short, basal level portion about half length of obliquely sloping flat apical portion; pro-mesonotal suture very distinct; thorax at mesometanotal suture deeply emarginate; propodeal spiracle round; legs somewhat short and stout, claws simple. Pedicel with small quadrangular, more or less laminate node; abdomen massive, broadly oval, not produced in front or overhanging the pedicel. Erect body hairs short and golden, not coarse. Queen: Head and pedicel as in worker; ocelli larger and more distinct. Thorax very massive, pronotum short, vertical, not forming part of dorsum; mesonotum very large, convex; scutellum broad, somewhat flat; metanotum short, rounded posteriorly; abdomen large, oval. Male: Mandibles broad, masticatory margin acute, with only one apical tooth, or whole masticatory margin dentate. Antennae 13-segment, flagellum filiform, segments nearly equal in length, first segment thicker than others. Antennal carinae short, diverging posteriorly. Genitalia very small on under side of abdomen, covered by tectiform dorsum of apical abdominal segments. Exterior genital plates narrowing towards apex, twice as long as at base, broad, and rounded apical plate crescentic. Ventral plate not emarginate.

<==Lasius Fabr. 1804 B03
    |--*L. niger B03
    |--L. affinis MS01
    |--L. alienobrunneus Forel 1894 [=L. niger brunneus var. alienobrunneus] B03
    |--L. alienoflavus Forel in Bingham 1903 B03
    |--L. alienus (Förster 1850) B03 [=Formica aliena B03, L. niger alienus MS01]
    |--L. atopus Cole 1958 WW90
    |--L. brunneus W01
    |--L. californicus BS06
    |--L. crinitus (Smith 1858) [=Formica crinita] B03
    |--L. distinguendus ES12
    |--L. flavus S61
    |--L. fuliginosus (Latr. 1798) [=Formica fuliginosa] B03
    |--L. grandis ES12
    |--L. hikosanus Yamauchi 1979 I92
    |--L. himalayanus Forel 1894 [=L. niger brunneus var. himalayanus] B03
    |--L. murphyi ES12
    |--L. neglectus ES12
    |--L. neoniger GE05
    |--L. nevadensis Cole 1956 WW90
    |--L. nitigaster WD16
    |--L. pallitarsis ES12
    |--L. platythorax WD16
    `--L. umbratus A71

*Type species of generic name indicated

REFERENCES

[A71] Askew, R. R. 1971. Parasitic Insects. Heinemann Educational Books: London.

[B03] Bingham, C. T. 1903. The Fauna of British India, including Ceylon and Burma. Hymenoptera vol. 2. Ants and Cuckoo-Wasps. Taylor and Francis: London.

[BS06] Brady, S. G., T. R. Schultz, B. L. Fisher & P. S. Ward. 2006. Evaluating alternative hypotheses for the early evolution and diversification of ants. Proceedings of the National Academy of Sciences of the USA 103 (48): 18172–18177.

[ES12] Espadaler, X., & S. Santamaria. 2012. Ecto- and endoparasitic fungi on ants from the Holarctic region. Psyche 2012: 168478.

[GE05] Grimaldi, D., & M. S. Engel. 2005. Evolution of the Insects. Cambridge University Press: New York.

[I92] Iwahashi, J. (ed.) 1992. Reddo Deeta Animaruzu: a pictorial of Japanese fauna facing extinction. JICC: Tokyo.

[MS01] Mocsáry, A., & V. Szépligeti. 1901. Hymenopterák [Hymenopteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 121–169. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[S61] Southcott, R. V. 1961. Studies on the systematics and biology of the Erythraeoidea (Acarina), with a critical revision of the genera and subfamilies. Australian Journal of Zoology 9: 367–610.

[WW90] Wheeler, G. C., & J. Wheeler. 1990. Insecta: Hymenoptera Formicidae. In: Dindal, D. L. (ed.) Soil Biology Guide pp. 1277–1294. John Wiley & Sones: New York.

[WD16] Wrzosek, M., G. Dubiel, M. Gorczak, J. Pawłowska, M. Tischer & S. Bałazy. 2016. New insights on the phylogeny and biology of the fungal ant pathogen Aegeritella. Journal of Invertebrate Pathology 133: 1–7.

[W01] Wurst, E. 2001. The life cycle of Lemanniella minotauri n. sp. and the erection of the new family Lemanniellidae (Acari: Astigmata). Stuttgarter Beiträge zur Naturkunde Serie A (Biologie) 621: 1–34.

Ichneumonini

Ichneumon extensorius, copyright James K. Lindsey.

Belongs within: Ichneumonidae.

The Ichneumonini is a diverse but poorly defined group of generalised Ichneumoninae mostly parasitic on large Lepidoptera (Gauld 1984).

<==Ichneumonini [Acanthojoppini, Joppini] NI11
    |--Stirexephanes melanarius Cameron 1912 NI11
    |--Crytea G84
    |--Chiaglas longicornis Cameron 1903 [=Stenichneumon longicornis; incl. C. tinctipennis Cameron 1903] NI11
    |--Hytophatnus lineatus Cameron 1907 NI11
    |--Stenaoplus ornatitarsis (Cameron 1904) (see below for synonymy) NI11
    |--Aculicoxa Gauld 1984 G84
    |    `--*A. striata Gauld 1984 G84
    |--Algathia Cameron 1902 G84
    |    `--*A. maculiceps G84
    |--Longichneumon Heinrich 1934 G84
    |    `--*L. annaelisae G84
    |--Phaeneumon Gauld 1984 G84
    |    `--*P. phoenix Gauld 1984 G84
    |--Setanta Cameron 1901 G84
    |    `--*S. rufipes G84
    |--Uloola Gauld 1984 G84
    |    `--*U. brevis Gauld 1984 G84
    |--Barichneumonites Heinrich 1934 G84
    |    |--*B. sphaeriscutellatus G84
    |    `--B. australasiae G84
    |--Lissosculpta Heinrich 1934 G84
    |    |--*L. impexa [=Ichneumon impexus] G84
    |    `--L. basalis G84
    |--Eutanyacra Cameron 1903 G84
    |    |--*E. pallidicoxis G84
    |    `--E. licitatorius [=Ichneumon licitatorius] G84
    |--Tricholabus Thomson 1894 [incl. Epiopelmidea Viereck 1913, Otohimea Uchida 1926] G84
    |    |--*T. femoralis G84
    |    |--‘*Epiopelmidea’ erythrogaster G84
    |    `--‘*Otohimea’ nigra G84
    |--Losgna NI11
    |    |--L. cariniscutis (Cameron 1905) [=Agarenes cariniscutis] NI11
    |    `--L. nigrita NI11
    |         |--L. n. nigrita NI11
    |         `--L. n. maculate Heinrich 1965 NI11
    |--Ctenochares Foerster 1869 (see below for synonymy) G84
    |    |--C. bicolorus (see below for synonymy) G84
    |    |--‘*Stenophorus’ amoenus G84
    |    |--‘Hoplismenus’ fulvidus [incl. *Talimeda pallidiceps] G84
    |    |--‘*Anisojoppa’ lutea G84
    |    `--‘*Henicophatnus’ rufithorax [incl. *Celmis blanditus] G84
    |--Gavrana Cameron 1906 [incl. Rossitera Cheesman 1953] G84
    |    |--*G. maculipes G84
    |    |--G. conica Gauld 1984 G84
    |    |--‘Ichneumon’ ischioleucus [=Levansa ischioleuca] G84
    |    |--‘Cryptus’ tiphiipuppis [=*Rossitera tiphiipuppis] G84
    |    `--G. spinosa Gauld 1984 G84
    `--Ichneumon Linnaeus 1758 (see below for synonymy) G84
         |--*I. extensorius Linnaeus 1758 G84, L58
         |--I. abbreviatus L02
         |--I. albiger PK17
         |--I. annulicornis Szépligeti in Mocsáry & Szépligeti 1901 MS01
         |--I. aphidum Linnaeus 1758 L58
         |--I. appendigaster Linnaeus 1758 L58
         |--I. ariolator Linnaeus 1758 L58
         |--I. assectator Linnaeus 1758 L58
         |--I. camelus Linnaeus 1758 L58
         |--I. caudator Fabricius 1775 G84
         |--I. cerinthius P93
         |--I. circumflexus Linnaeus 1758 L58
         |--I. coccorum Linnaeus 1758 L58
         |--I. compressus [=Banchus compressus] L02
         |--I. compunctor Linnaeus 1758 L58
         |--I. constrictorius Linnaeus 1758 L58
         |--I. coronatus [=Stephanus coronatus] L02
         |--I. coruscator Linnaeus 1758 L58
         |--I. crispatorius Linnaeus 1758 L58
         |--I. culpatorius Linnaeus 1758 L58
         |--I. cultriformis L02
         |--I. cyniphidis Linnaeus 1758 L58
         |--I. deliratorius Linnaeus 1758 L58
         |--I. delusor Linnaeus 1758 L58
         |--I. denigrator Linnaeus 1758 L58
         |--I. deprimator L02
         |--I. desertor Linnaeus 1758 L58
         |--I. designatorius Linnaeus 1758 L58
         |--I. dubitator L02
         |--I. edictorius linnaeus 1758 L58
         |--I. elongator L02
         |--I. enervator Linnaeus 1758 L58
         |--I. exarator Linnaeus 1758 L58
         |--I. extensor Linnaeus 1758 L58
         |--I. flavator L02
         |--I. flavomaculatus Lucas 1849 E12
         |--‘*Coreojoppa flavomaculata’ Uchida 1926 non Ichneumon flavomaculatus Lucas 1849 G84
         |--I. formicarius L02
         |--I. fossorius Linnaeus 1758 L58
         |--I. glaucopterus Linnaeus 1758 L58
         |--I. globatus Linnaeus 1758 L58
         |--I. glomeratus Linnaeus 1758 L58
         |--I. hemipteron Riche 1802 R02
         |--I. incubitor Linnaeus 1758 L58
         |--I. inculcator Linnaeus 1758 L58
         |--I. insidiosus P93
         |--I. jaculator Linnaeus 1758 L58
         |--I. juniperi Linnaeus 1758 L58
         |--I. juvencus Linnaeus 1758 L58
         |--I. latrator [incl. *Brachypterus means, *Pterocormus means] G84
         |--I. lotatorius [incl. *Colobacis forticornis] G84
         |--I. luctatorius Linnaeus 1758 L58
         |--I. mandator Linnaeus 1758 L58
         |--I. manifestator Linnaeus 1758 L58
         |--I. migrator L02
         |--I. moderator Linnaeus 1758 L58
         |--I. muscarum Linnaeus 1758 L58
         |--I. nominator L02
         |--I. nunciator L02
         |--I. ovulorum Linnaeus 1758 L58
         |--I. pendulator Latreille 1800 L00
         |--I. peregrinator Linnaeus 1758 L58
         |--I. perturbator L02
         |--I. pisorius Linnaeus 1758 L58
         |--I. praerogator Linnaeus 1758 L58
         |--I. primatorius [incl. *Matsumuraius grandis] G84
         |--I. profligator L02
         |--I. promissorius G84 (see below for synonymy)
         |--I. pugillator Linnaeus 1758 L58
         |--I. pugillator Linnaeus 1758 L58
         |--I. quadrialbatus MS01
         |--I. raptorius Linnaeus 1758 L58
         |--I. reluctator Linnaeus 1758 L58
         |--I. resinellae Linnaeus 1758 L58
         |--I. rubrocinctus Lucas 1849 E12
         |--‘*Tyanites’ rufipes [incl. *Vabsaris forticornis] G84
         |--I. ruspator Linnaeus 1758 L58
         |--I. sarcitorius Linnaeus 1758 L58 [=*Euichneumon sarcitorius G84]
         |--I. saturatorius Linnaeus 1758 L58
         |--I. secalis Linnaeus 1758 L58
         |--I. serrator L02
         |--I. sessilis [=Evania sessilis] L02
         |--I. spectrum Linnaeus 1758 L58
         |--I. strobilellae Linnaeus 1758 L58
         |--I. subcutaneus Linnaeus 1758 L58
         |--I. sugillatorius Linnaeus 1758 L58
         |--I. titillator Linnaeus 1758 L58
         |--I. tuberosus [incl. *Bureschias balcanicus] G84
         |--I. turionellae Linnaeus 1758 L58
         |--I. urinator L02
         |--I. vaginatorius Linnaeus 1758 L58
         |--I. varius [=Banchus varius] L02
         |--I. venator Linnaeus 1758 L58 [=Banchus venator L02]
         |--I. vesparum Rossi 1793 R93
         |--I. vespoides L02
         `--I. volutatorius Linnaeus 1758 L58

Ctenochares Foerster 1869 [=Joppites Berthoumieu 1894, Joppoides Berthoumieu 1904; incl. Anisojoppa Cameron 1906, Celmis Tosquinet 1896, Henicophatnus Kriechbaumer 1894, Pseudojoppa Kriechbaumer 1898, Stenophorus Saussure 1892, Talimeda Cameron 1912] G84

Ctenochares bicolorus [=Ichneumon bicolorus; incl. C. apicalis, I. instructor, *Pseudojoppa instructor, Ctenochares rufator, Ichneumon xanthomelas, *C. xanthomelas, *Joppites xanthomelas, *Joppoides xanthomelas] G84

Ichneumon Linnaeus 1758 [incl. Brachypterus Gravenhorst 1829 non Kugelman 1794, Bureschias Heinrich 1936, Colobacis Cameron 1900, Coreojoppa Uchida 1926, Euichneumon Berthoumieu 1904, Matsumuraius Ashmead 1906, Pterocormus Foerster 1850, Tyanites Cameron 1903, Vabsaris Cameron 1903] G84

Ichneumon promissorius G84 [=Pterocormus promissorius F02; incl. I. albocinctus G84, I. australis G84, I. leucaniae G84, I. varilineatus G84]

Stenaoplus ornatitarsis (Cameron 1904) [=Hemiteles ornatitarsis; incl. H. caducus Schulz 1906, H. geniculatus Cameron 1904 non Thomson 1884] NI11

*Type species of generic name indicated

REFERENCES

[E12] Evenhuis, N. L. 2012. Publication and dating of the Exploration Scientifique de l'Algérie: Histoire Naturelle des Animaux Articulés (1846–1849) by Pierre Hippolyte Lucas. Zootaxa 3448: 1–61.

[F02] Foord, M. 2002. Entomological Society conference field trip 2002. Weta 24: 37–38.

[G84] Gauld, I. D. 1984. An Introduction to the Ichneumonidae of Australia. British Museum (Natural History).

[L00] Latreille, P. A. 1800. Sur une nouvelle espèce d'ichneumon. Bulletin des Sciences, par la Societé Philomathique de Paris 2 (42): 138.

[L02] Latreille, P. A. 1802. Histoire Naturelle, générale et particulière des crustacés et des insectes vol. 3. Familles naturelles des genres. F. Dufart: Paris.

[L58] Linnaeus, C. 1758. Systema Naturae per Regna Tria Naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata. Laurentii Salvii: Holmiae.

[MS01] Mocsáry, A., & V. Szépligeti. 1901. Hymenopterák [Hymenopteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 121–169. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[NI11] Norhafiza, A. F., & A. B. Idris. 2011. A catalogue of Ichneumoninae (Hymenoptera: Ichneumonidae) from Malaysia. Serangga 16 (1): 29–50.

[PK17] Peters, R. S., L. Krogmann, C. Mayer, A. Donath, S. Gunkel, K. Meusemann, A. Kozlov, L. Podsiadlowski, M. Petersen, R. Lanfear, P. A. Diez, J. Heraty, K. M. Kjer, S. Klopfstein, R. Meier, C. Polidori, T. Schmitt, S. Liu, X. Zhou, T. Wappler, J. Rust, B. Misof & O. Niehuis. 2017. Evolutionary history of the Hymenoptera. Current Biology 27 (7): 1013–1018.

[P93] Pittaway, A. R. 1993. The Hawkmoths of the Western Palaearctic. Harley Books: Colchester.

[R02] Riche, M. 1802. Description de l'Ichneumon-Hemipteron, espèce nouvelle. Bulletin des Sciences, par la Societé Philomathique de Paris 1 (Bulletin de la Société Philomathique, a ses correspondans): 1'.

[R93] Rossi, M. 1793. Observations de M. Rossi sur un nouveau genre d'insecte, voisin des ichneumons. Bulletin de la Société Philomatique, a ses Correspondans 23–24: 1.

Stizus

Stizus fasciatus, copyright Cor Zonneveld.

Belongs within: Stizini.

Stizus is a genus of medium-sized to large wasps found in Eurasia, North America and Africa that provision their nests with Orthoptera or mantids (Bohart & Menke 1976).

Characters (from Bohart & Menke 1976): Eyes converging slightly below, at least narrower below than above; a low and somewhat double convexity below mid-ocellus; mandible dentate subapically; exposed part of labrum moderately convex, usually a fourth to a third as long as broad; male antenna without a projection on flagellomere IX; scutellum typically (but not always) with a medial, pubescent depression in the female; propodeum not compressed into a ridge posterolaterally; midtibia with two apical spurs; forewing with submarginal cell II open in front; hindwing media diverging far before cu-a; median cell with two distal appendices; precoxal sulcus usually present.

<==Stizus Latreille 1802-1803 [incl. Megalostizus Schulz 1906, Megastizus Patton 1879, Stizolarra Saussure 1887] BM76
    |--S. ruficornis (Forster 1771) (see below for synonymy) BM76
    |    |--S. r. ruficornis BM76
    |    |--S. r. eremicus Bytinski-Salz 1955 BM76
    |    `--S. r. strigatus Mochi 1939 BM76
    |--S. adelphus Kirby 1900 BM76
    |--S. aegyptius Lepeletier 1845 BM76
    |--S. aestivalis Mercet 1906 BM76
    |    |--S. a. aestivalis BM76
    |    `--S. a. merceti Beaumont 1951 BM76
    |--S. anchoratus Mantero 1917 BM76
    |--S. anchorites Turner 1920 BM76
    |--S. annulatus (Klug 1845) (see below for synonymy) BM76
    |--S. apicalis Guérin-Méneville 1844 BM76
    |--S. arenarum Handlirsch 1892 BM76
    |--S. arnoldi Mochi 1939 BM76
    |--S. atrox (Smith 1856) [=Larra atrox; incl. L. flavomaculata Smith 1873, Stizus pentheres Handlirsch 1895] BM76
    |--S. aurifluus Turner 1916 BM76
    |--S. basalis Guérin-Méneville 1844 [incl. Larra infuscata Klug 1845] BM76
    |--S. baumanni Handlirsch 1901 BM76
    |--S. beccarii Mantero 1917 BM76
    |--S. bensoni Arnold 1951 BM76
    |--S. berlandi Arnold 1945 BM76
    |--S. biclipeatus (Christ 1791) (see below for synonymy) BM76
    |--S. bipunctatus (Smith 1856) [=Larra bipunctata] BM76
    |--S. bizonatus Spinola 1838 [incl. Larra sphegiformis Klug 1845] BM76
    |--S. brevipennis Walsh 1869 [=*Megastizus brevipennis; incl. Larra brendeli Taschenberg 1875] BM76
    |--S. breyeri Arnold 1936 BM76
    |--S. chrysorrhoeus Handlirsch 1892 BM76
    |--S. coloratus Nurse 1903 BM76
    |--S. combustus (Smith 1856) [=Larra combusta; incl. L. fuliginosa Klug 1845 non Dahlbom 1843] BM76
    |--S. continuus (Klug 1835) [=Larra continua] BM76
    |--S. delessertii Guérin-Méneville 1844 BM76
    |--S. deserticus Giner Marí 1945 BM76
    |--S. dewitzii Handlirsch 1892 [incl. S. erythraspis Cameron 1910, S. propodealis Bischoff 1915] BM76
    |--S. dispar Morawitz 1888 BM76
    |--S. elegans Dahlbom 1845 BM76
    |--S. ellenbergeri Arnold 1929 BM76
    |--S. emir Handlirsch 1901 BM76
    |--S. erythraeensis Mantero 1917 BM76
    |--S. erythrocephalus (Taschenberg 1880) [=Larra erythrocephala] BM76
    |--S. euchromus Handlirsch 1892 BM76
    |--S. eugeniae Gussakovskij 1935 BM76
    |--S. excellens Lohrmann 1943 [incl. S. eximius Arnold 1936 non Morawitz 1894] BM76
    |--S. eximius Morawitz 1894 BM76
    |--S. fasciatus (Fabricius 1781) (see below for synonymy) BM76
    |--S. fedtschenkoi Radoszkowski 1877 BM76
    |--S. ferrandii Magretti 1898 BM76
    |--S. ferrugineus (Smith 1856) [=Larra ferruginea; incl. S. ritsemae Handlirsch 1895] BM76
    |--S. franzi Turner 1916 [incl. S. congoensis Mantero 1917, S. neavei Kohl 1913 non Turner 1912] BM76
    |--S. fuscatus Bingham 1897 BM76
    |--S. fuscipennis (Smith 1856) [=Larra fuscipennis] BM76
    |--S. gobiensis Tsuneki 1970 BM76
    |--S. gracilipes Handlirsch 1892 BM76
    |--S. hamatus Arnold 1929 BM76
    |--S. handlirschi Radoszkowski 1893 BM76
    |--S. hebraeus Balthasar 1954 BM76
    |--S. hispanicus Mocsáry 1883 [incl. S. villosus Costa 1887] BM76
    |--S. histrio Morawitz 1888 BM76
    |--S. hugelii Handlirsch 1892 BM76
    |--S. hyalipennis Handlirsch 1892 BM76
    |--S. imperator Nurse 1903 BM76
    |--S. imperialis Handlirsch 1892 [incl. S. tulbaghensis Arnold 1936] BM76
    |    |--S. i. imperalis BM76
    |    `--S. i. conspicuus Arnold 1952 BM76
    |--S. iridis Dow 1942 BM76
    |--S. jordanicus Lohmann 1942 BM76
    |--S. kaszabi Tsuneki 1971 BM76
    |--S. kiseritzkii Gussakovskij 1928 BM76
    |--S. koenigi Morawitz 1888 [incl. S. tages Kirby 1889] BM76
    |--S. kohlii Mocsáry 1883 BM76
    |--S. lacteipennis Mocsáry 1883 BM76
    |--S. lepidus (Klug 1845) [=Larra lepida] BM76
    |--S. lineatus (Fabricius 1793) [=Bembex lineatus] BM76
    |--S. lohrmanni Bohart in Bohart & Menke 1976 (see below for synonymy) BM76
    |--S. lughensis Magretti 1898 BM76
    |--S. luteotaeniatus Gussakovskij 1933 BM76
    |--S. lutescens Radoszkowski 1877 BM786
    |--S. marnonis Handlirsch 1892 BM76
    |--S. marshalli Turner 1912 BM76
    |--S. marthae Handlirsch 1892 [incl. S. cheops Morice 1897, S. cheops Bingham 1897 non Morice 1897] BM76
    |--S. melanoxanthus (Smtih 1856) [=Larra melanoxantha] BM76
    |--S. melanurus Handlirsch 1892 BM76
    |--S. melleus (Smith 1856) [=Larra mellea] BM76
    |--S. mocsaryi Handlirsch 1895 BM76
    |--S. mongolicus Tsuneki 1971 BM76
    |--S. multicolor Turner 1916 BM76
    |--S. nadigi Roth 1933 BM76
    |--S. neavei Turner 1912 BM76
    |--S. nigriventris Arnold 1951 BM76
    |--S. niloticus Handlirsch 1892 BM76
    |--S. occidentalis Parker 1929 BM76
    |--S. ocellatus Gistel 1857 BM76
    |--S. orientalis Cameron 1890 BM76
    |--S. pauli Mantero 1917 BM76
    |--S. perrisii Dufour 1838 BM76
    |    |--S. p. perrisii BM76
    |    `--S. p. ibericus Beaumont 1962 BM76
    |--S. pictus Dahlbom 1845 BM76
    |--S. pluschtschevskii Radoszkowski 1888 BM76
    |--S. praestans Morawitz 1893 BM76
    |--S. pulcherrimus (Smith 1856) [=Larra pulcherrima] BM76
    |--S. pygidialis Handlirsch 1892 BM76
    |--S. quartinae (Gribodo 1884) [=Sphecius quartinae] BM76
    |--S. raddei Handlirsch 1889 BM76
    |--S. rapax Handlirsch 1892 BM76
    |--S. richardsi Arnold 1959 BM76
    |--S. rubellus Turner 1912 BM76
    |--S. rubroflavus Turner 1916 BM76
    |--S. rufescens (Smith 1856) [=Larra rufescens] BM76
    |--S. rufipes (Fabricius 1804) [=Liris rufipes; incl. S. gigas Walsh & Riley 1868, S. grandis Lepeletier 1845] BM76
    |--S. rufiventris Radoszkowski 1877 [incl. S. sarmaticus Morawitz 1891] BM76
    |    |--S. r. rufiventris BM76
    |    `--S. r. compar Handlirsch 1892 BM76
    |--S. rufocinctus Dahlbom 1845 BM76
    |--S. rufoniger Mochi 1939 BM76
    |--S. saharae Roth 1934 BM76
    |--S. saussurei Handlirsch 1895 BM76
    |--S. savignyi Spinola 1838 [incl. Larra succinea Klug 1845] BM76
    |--S. schmiedeknechti Handlirsch 1898 BM76
    |--S. scoliaeformis Arnold 1929 BM76
    |--S. scolioides Kokujev 1902 BM76
    |--S. sexfasciatus (Fabricius 1793) [=Bembex sexfasciatus] BM76
    |--S. spectrum Handlirsch 1901 BM76
    |--S. spinulosus Radoszkowski 1876 BM76
    |--S. stevensoni Arnold 1936 BM76
    |--S. storeyi Turner 1920 BM76
    |--S. tenuicornis (Smith 1856) [=Larra tenuicornis] BM76
    |--S. texanus Cresson 1872 BM76
    |--S. transcaspicus Radoszkowski 1893 BM76
    |--S. tricolor Handlirsch 1892 BM76
    |--S. tridens MS01
    |--S. tridentatus [=Bembex tridentata] L02
    |--S. tunetanus Costa 1893 BM76
    |--S. vespiformis (Fabricius 1775) [=Sphex vespiformis, *Stizolarra vespiformis] BM76
    |--S. vespoides (Walker 1871) [=Larra vespoides; incl. L. argentea Taschenberg 1875, L. magnifica Smith 1873] BM76
    |--S. wheeleri Arnold 1959 BM76
    |--S. zhelochovtzevi Gussakovskij 1952 BM76
    |--S. zimini Gussakovskij 1928 BM76
    |--S. zonatus (Klug 1845) [=Larra zonata; incl. L. dimidiata Taschenberg 1875 non Panzer 1806-1809] BM76
    `--S. zonosoma Handlirsch 1895 BM76

Stizus annulatus (Klug 1845) [=Larra annulata; incl. S. eversmanni Radoszkowski 1877, S. kisilkumii Radoszkowski 1877, S. picticornis Morawitz 1891, Larra subapicalis Walker 1871] BM76

Stizus biclipeatus (Christ 1791) [=Vespa biclipeata; incl. Larra cingulata Fabricius 1798, L. nubilipennis Smith 1856, Tiphia ruficornis Fabricius 1787 nec Vespa ruficornis Forster 1771 nec Bembex ruficornis Fabricius 1787] BM76

Stizus fasciatus (Fabricius 1781) [=Bembex fasciatus; incl. B. integra Fabricius 1804, Stizus terminalis Dahlbom 1845] BM76

Stizus lohrmanni Bohart in Bohart & Menke 1976 [=S. arnoldi Lohrmann 1943 non Mochi 1939; incl. S. pubescens Arnold 1929 non Klug 1835] BM76

Stizus ruficornis (Forster 1771) [=Vespa ruficornis; incl. S. distinguendus Handlirsch 1901, V. gaditana Gmelin 1790, S. komarovi Radoszkowski 1888, S. ornatus Dahlbom 1845, Larra pubescens Klug 1835, Bembex ruficornis Fabricius 1787 non Vespa ruficornis Forster 1771, *Stizus ruficornis, S. scutellaris Kirby 1900] BM76

*Type species of generic name indicated

REFERENCES

[BM76] Bohart, R. M., & A. S. Menke. 1976. Sphecid Wasps of the World. University of California Press: Berkeley.

[L02] Latreille, P. A. 1802. Histoire Naturelle, générale et particulière des crustacés et des insectes vol. 3. Familles naturelles des genres. F. Dufart: Paris.

[MS01] Mocsáry, A., & V. Szépligeti. 1901. Hymenopterák [Hymenopteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 121–169. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

Pompilus

Pompilus cinereus, copyright B. G. Mot.

Belongs within: Pompilidae.

Pompilus is one of a number of genera of spider wasps with black integument overlain with grey pubescence (Elliott 2007).

Pompilus Fabricius 1798 [incl. Chionopompilus Priesner 1955] E07
    |--P. cinereus (Fabricius 1775) (see below for synonymy) E07
    |--P. aterrimus MS01
    |--P. audax Westwood 1842 E07
    |--P. caerulescens Dalla Torre 1897 [=P. caeruleus Smith 1879 non Taschenberg 1869] E07
    |--P. campestris MS01
    |--P. csikianus Kohl in Mocsáry & Szépligeti 1901 MS01
    |--P. flavicornis Smith 1855 E07
    |--P. fumipennis MS01
    |--P. infandus Smith 1868 E07
    |--P. lugubris Smith 1868 E07
    |--P. lunulatus (Saussure 1867) (see below for synonymy) E07
    |--P. niveus Saunders 1901 [incl. *Chionopompilus rabinovitchi Priesner 1955] E07
    |--P. pictifrons Smith 1855 E07
    |--P. sanguinicolor Dalla Torre 1897 [=P. sanguinolentus Smith 1879 non Fabricius 1793] E07
    |--P. tripunctatus MS01
    |--P. trivialis MS01
    `--P. viaticus MS01

Pompilus cinereus (Fabricius 1775) [=Sphex cinerea; incl. Pompilus ami Tsuneki 1989, Pompilus argyrostoma Stephens 1829 (n. n.), Sericopompilus bivittatus Banks 1934, Pompilus bivittatus, Pompilus bunun Tsuneki 1989, Pompilus chevrieri Tournier 1889, Pompilus plumbeus chevrieri, Aporoideus clarus Banks 1941, Pompilus plumbeus gotlandicus Wolf 1972, Pompilus ithonus Cameron 1908, Psammochares latilabris Arnold 1937, Pompilus leprosus Dalla Torre 1897, Pompilus plumbeus lusitanicus Wolf & Dinz 1970, Pompilus plumbeus lucitanicus (l. c.), Psammochares plumbeus maroccana Nadig & Nadig 1935 (n. n.), Pompilus pulcher placidus Priesner 1968 (nom. inv.), Sphex plumbea Fabricius 1787, Ferreola plumbea, Pepsis plumbea, Pompilius (l. c.) plumbeus, Pompilus plumbea, Pompilus plumbeus, Psammochares plumbeau (l. c.), Ps. plumbeus, Pompilus plumbeicolor Dalla Torre 1897, Pompilus pruinosus Smith 1879 non Smith 1855, *Pompilus pulcher Fabricius 1798, Pompilus plumbeus pulcher, Psammochares plumbeus pulchra, Pompilus sericeibalteatus Cameron 1910, Pompilus tsou Tsuneki 1989] E07

Pompilus lunulatus (Saussure 1867) [=Pogonius lunulatus, Agenia lunulata, Deuteragenia lunulata, Pseudagenia lunulata; incl. Pompilus spectrum Kohl 1886] E07

*Type species of generic name indicated

REFERENCES

[E07] Elliott, M. G. 2007. Annotated catalogue of the Pompilidae (Hymenoptera) of Australia. Zootaxa 1428: 1–83.

[MS01] Mocsáry, A., & V. Szépligeti. 1901. Hymenopterák [Hymenopteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 121–169. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

Cryptocheilus

Cryptocheilus australis, from here.

Belongs within: Pompilidae.

Cryptocheilus is a genus of pompilid wasps characterised by a marginal cell on the forewing that is broadly rounded apically, with vein Rs meeting the anterior margin of the wing almost at a right angle (Wahis 2008).

Cryptocheilus Panzer 1806 (see below for synonymy) E07
    |--*C. annulatus (Fabricius 1798) [=Sphex annulata, *Calicurgus annulatus] E07
    |--C. australasiae Haupt 1935 non Mygnimia australasiae Smith 1873 (not preoc. if in dif. gen.) E07
    |--C. australis (Guérin-Méneville 1838) (see below for synonymy) E07
    |--C. avidus (Nurse 1903) [=Salius avidus] W08
    |--C. basipennis (Turner 1910) [=Calicurgus basipennis] E07
    |    |--C. b. basipennis E07
    |    `--C. b. agnatus (Turner 1910) [=Calicurgus basipennis agnatus] E07
    |--C. birkmanni Banks 1926 [=*Chilochares birkmanni] E07
    |--C. darwinii Turner 1910 E07
    |--‘Sphex’ discolor Fabricius 1793 [=*Homonotus discolor, *Trachyocheilus discolor] E07
    |--C. distinctus (Smith 1868) [=Pompilus distinctus] E07
    |--C. fulvidorsalis Turner 1910 E07
    |--‘Salius’ fuscus MS01
    |--C. geminus (Bingham 1896) [=Salius (Priocnemis) geminus] W08
    |--‘Calicurgus’ hyalinatus H37
    |--C. ichneumonoides (Costa 1874) [=Priocnemis icheumonoides, C. (*Ichneumonocheilus) ichneumonoides] E07
    |--‘*Stenoclavelia’ mirabilis Arnold 1932 E07
    |--C. modestus (Smith 1855) [=Calicurgus modestus] E07
    |--C. momus (Bingham 1896) [=Salius (Priocnemis) momus] W08
    |--C. praepositus Haupt 1935 [=Cryptochilus praepositus] E07
    |--C. properus (Bingham 1896) [=Salius (Priocnemis) properus] W08
    |--C. rubellus (Eversmann 1864) [=Pompilus rubellus, C. (*Cyphonocheilus) rubellus] E07
    |--C. sathanas Turner 1910 E07
    |--‘Salius’ schenckii MS01
    |--‘Sphex’ sexpunctata Fabricius 1804 [=*Adonta sexpunctata, *Salius sexpunctatus] E07
    |--‘Salius’ spectabilis Morawitz 1892 [=*Chlorocheilus spectabilis] E07
    |--C. stehliki Wolf 1968 [=C. (*Priocnemiocheilus) stehliki] E07
    |--‘*Adirostes’ tolteca Banks 1946 E07
    `--C. valentulus (Bingham 1896) [=Salius (Priocnemis) valentulus] W08

Cryptocheilus Panzer 1806 [=Calicurgus Brullé 1833, Cryptochilus Rafinesque 1815; incl. Adirostes Banks 1946, Adonta Billberg 1820, Chilochares Banks 1941, Chlorocheilus Wolf 1965 (n. n.), Cyphonocheilus Wolf 1970, Homonotus Dahlbom 1845 non Dahlbom 1843, Ichneumonocheilus Wolf 1970, Priocnemiocheilus Wolf 1970, Salius Fabricius 1804 non Schrank 1798, Stenoclavelia Arnold 1932, Trachyocheilus Wolf 1965 (n. n.)] E07

Cryptocheilus australis (Guérin-Méneville 1838) [=Pompilus australis, Cryptochilus (Prionocnemis) australis, Heterodontonyx australis] E07

*Type species of generic name indicated

REFERENCES

[E07] Elliott, M. G. 2007. Annotated catalogue of the Pompilidae (Hymenoptera) of Australia. Zootaxa 1428: 1–83.

[H37] Haupt, H. 1937. Beitrag zur Kenntnis der Psammochariden-Fauna Australiens (Spinnenfangende Wegwespen). Subf. Claveliinae Hpt. Trib. Calicurgini nov. Zeitschrift für Naturwissenschaften 91: 121–136.

[MS01] Mocsáry, A., & V. Szépligeti. 1901. Hymenopterák [Hymenopteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 121–169. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[W08] Wahis, R. 2008. Contribution à la connaissance des pompilides d'Australie (Hymenoptera: Pompilidae). 2. Sur quelques spécimens récoltés par G. Else (Natural History Museum, London) avec descriptions de deux espèces nouvelles des genres Auplopus et Ctenostegus. Faunistic Entomology—Entomologie Faunistique 61 (1–2): 23–31.

Nomia

Pearly-banded bee Nomia strigata, from here.

Belongs within: Halictidae.

Nomia is a genus of bees whose species excavate nests in soil (Michener 1965).

Characters (from Michener 1965): First flagellar segment shorter than scape; proboscis with apices of galeae in repose not reaching posterior end of proboscidial fossa; mesepisternum with distinct pre-episternal groove extending but little below scrobal groove; stigma distinct, longer than broad; three submarginal cells present, second submarginal cell on posterior side shorter than first;, third submarginal cell at least three-fourths as long as first and more than twice as long as second; posterior tibial spurs present; hind tibia without corbicula.

Nomia Latreille 1804 M65
    |  i. s.: N. albocincta Lucas 1848 E12
    |         N. caerulescens Friese 1909 M65
    |         N. clavata (Smith 1853) [=Halictus clavatus] M65
    |         N. femoralis MS01
    |         N. flavilabris Lucas 1848 E12
    |         N. melanderi WP99
    |         N. perforata Lucas 1848 E12
    |--N. (Nomia) M65
    |    |--*N. (N.) curvipes (Fabricius 1781) [=Andrena curvipes] M65
    |    |--N. (N.) eburneifera M65
    |    `--N. (N.) viridilimbata M65
    |--N. (Austronomia Michener 1965) M65
    |    |--N. (*A.) australica Smith 1875 [=N. australiaca Dalla Torre 1896] M65
    |    |--N. (A.) adelaidella Cockerell 1910 M65
    |    |--N. (A.) aenea Smith 1875 M65
    |    |--N. (A.) aenescens Friese 1912 M65
    |    |--N. (A.) aerata Smith 1875 M65
    |    |--N. (A.) alboscopacea Friese 1917 M65
    |    |--N. (A.) analis Friese 1924 M65
    |    |--N. (A.) antennata M65
    |    |--N. (A.) argentifrons Smith 1862 M65
    |    |--N. (A.) aurata M65
    |    |--N. (A.) babindensis Cockerell 1930 M65
    |    |--N. (A.) brisbanensis Cockerell 1913 M65
    |    |--N. (A.) cheesmanae Michener 1965 (see below for synonymy) M65
    |    |--N. (A.) clypeata M65
    |    |--N. (A.) cyanella Cockerell 1913 M65
    |    |--N. (A.) dentiventris Smith 1875 M65
    |    |--N. (A.) dimissa Cockerell 1921 M65
    |    |--N. (A.) doddi Cockerell 1905 M65
    |    |--N. (A.) erimae Friese 1909 M65
    |    |--N. (A.) excellens Cockerell 1929 M65
    |    |--N. (A.) ferricauda Cockerell 1913 M65
    |    |--N. (A.) flavoviridis Cockerell 1905 M65
    |    |--N. (A.) fortior Cockerell 1929 M65
    |    |--N. (A.) frenchi Cockerell 1912 M65
    |    |--N. (A.) fruhstorferi M65
    |    |--N. (A.) fulvoanalis Friese 1924 M65
    |    |--N. (A.) generosa Smith 1875 M65
    |    |--N. (A.) geophila Cockerell 1930 M65
    |    |--N. (A.) gilberti Cockerell 1905 M65
    |    |--N. (A.) gracilipes Smith 1875 [incl. N. haemorrhoidalis Westwood 1875] M65
    |    |--N. (A.) grisella Cockerell 1913 M65
    |    |--N. (A.) hippophila Cockerell 1910 M65
    |    |--N. (A.) hypodonta Cockerell 1905 M65
    |    |--N. (A.) kankauana M65
    |    |--N. (A.) kurandina Cockerell 1910 M65
    |    |--N. (A.) latetibialis Friese 1924 M65
    |    |--N. (A.) latitarsis Friese 1924 M65
    |    |--N. (A.) latiuscula Friese 1924 M65
    |    |--N. (A.) maai Michener 1965 M65
    |    |--N. (A.) macularis Friese 1924 M65
    |    |--N. (A.) melanodonta Cockerell 1926 M65
    |    |--N. (A.) melanoptera Cockerell 1910 M65
    |    |--N. (A.) melvilliana Cockerel 1929 M65
    |    |--N. (A.) miranda Rayment 1954 M65
    |    |--N. (A.) moerens Smith 1875 M65 [=Lipotriches moerens ZS10]
    |    |--N. (A.) muscosa Cockerell 1910 M65
    |    |--N. (A.) musgravei Cockerell 1929 M65
    |    |--N. (A.) nana Smith 1875 M65
    |    |--N. (A.) nuda Rayment 1939 [=N. australica nuda] M65
    |    |--N. (A.) pennata Friese 1924 M65
    |    |--N. (A.) perlucida M65
    |    |--N. (A.) phanerusa Cockerell 1913 M65
    |    |--N. (A.) philippinensis M65
    |    |--N. (A.) pilipes M65
    |    |--N. (A.) purnongensis Cockerell 1913 M65
    |    |--N. (A.) reginae Cockerell 1905 M65
    |    |--N. (A.) regis Cockerell 1910 M65
    |    |--N. (A.) ‘rubra’ Rayment 1931 non Friese 1904 M65
    |    |--N. (A.) rufocognita Cockerell 1905 M65
    |    |--N. (A.) satelles Cockerell 1912 M65
    |    |--N. (A.) semiaurea Cockerell 1905 M65
    |    |--N. (A.) semipallida Cockerell 1905 M65
    |    |--N. (A.) sicheli Vachal 1897 M65
    |    |--N. (A.) smithella Gribodo 1894 [incl. N. ruficornis Smith 1875 non Spinola 1838] M65
    |    |--N. (A.) stalkeri Cockerell 1910 M65
    |    |--N. (A.) subaustralica Cockerell 1910 M65
    |    |--N. (A.) submoerens Cockerell 1914 M65
    |    |--N. (A.) tenuihirta Cockerell 1905 M65
    |    |--N. (A.) testaceipes Friese 1924 M65
    |    |--N. (A.) thor Cockerell 1930 M65
    |    |--N. (A.) turneri Friese 1924 M65
    |    |--N. (A.) ulongensis Cockerell 1929 M65
    |    |--N. (A.) victoriae Cockerell 1910 M65
    |    |--N. (A.) willeyi Cameron 1905 [incl. N. gazellae Friese 1909, N. metallica Cameron 1901 non Smith 1863] M65
    |    `--N. (A.) wilmattae Cockerell 1929 M65
    |--N. (Curvinomia Michener 1944) (see below for synonymy) M65
    |    |--N. (C.) tetrazona Cockerell 1910 (see below for synonymy) M65
    |    |--N. (C.) albofasciata M65
    |    |--N. (C.) aurantifer Cockerell 1910 M65
    |    |--N. (C.) borneana M65
    |    |--N. (C.) chalybeata Smith 1875 [=*Paranomia chalybeata, *Paranomina chalybeata] M65
    |    |--N. (C.) elegans M65
    |    |--N. (C.) formosa M65
    |    |--N. (C.) fulvata (Fabricius 1804) [=Megilla fulvata, Amegilla fulvata] M65
    |    |--N. (C.) iridescens Smith 1853 M65
    |    |--N. (C.) longitarsis M65
    |    |--N. (C.) lusoria M65
    |    |--N. (C.) luteofasciata Friese 1917 M65
    |    |--N. (C.) megasoma M65
    |    |--N. (C.) mimosae M65
    |    |--N. (C.) opulenta Smith 1864 M65
    |    |--N. (C.) penangensis M65
    |    |--N. (C.) quadridentata Smith 1875 M65
    |    |--N. (C.) ridleyi M65
    |    |--N. (C.) subpurpurea M65
    |    |--N. (C.) swainsoniae Cockerell 1921 M65
    |    |--N. (C.) tuberculifrons M65
    |    `--N. (C.) viridicincta M65
    |--N. (Hoplonomia Ashmead 1904) M65
    |    |--N. (*H.) quadrifasciata (Ashmead 1904) [=*Hoplonomia quadrifasciata] M65
    |    |--N. (H.) amboinensis Cockerell 1907 M65
    |    |--N. (H.) austrovagans Cockerell 1905 M65 [=N. (H.) pulchribalteata var. austrovagans R35]
    |    |--N. (H.) callichlora M65
    |    |--N. (H.) cuneata M65
    |    |--N. (H.) darwinorum Cockerell 1910 M65
    |    |--N. (H.) ellioti Smith 1875 M65
    |    |--N. (H.) flavipennis Friese 1909 M65
    |    |--N. (H.) froggatti Cockerell 1911 M65
    |    |--N. (H.) incerta M65
    |    |--N. (H.) lyonsiae Cockerell 1912 M65
    |    |--N. (H.) papuana Cockerell 1929 M65
    |    |--N. (H.) parvula M65
    |    |--N. (H.) pulchra M65
    |    |--N. (H.) pulchribalteata Cameron 1901 M65
    |    |--N. (H.) punctulata M65
    |    |--N. (H.) rhododonta M65
    |    |--N. (H.) rubroviridis Cockerell 1905 M65
    |    |--N. (H.) strigata (Fabricius 1793) [=Andrena strigata] M65
    |    |--N. (H.) tridentata M65
    |    `--N. (H.) westwoodi M65
    |--N. (Mellitidia Guérin-Méneville 1830) M65
    |    |--N. (*M.) australis (Guérin-Méneville 1830) [=Andrena australis] M65
    |    |--N. (M.) cincta Smith 1859 M65
    |    |--N. (M.) dentata Smith 1859 M65
    |    |--N. (M.) gressitti Michener 1965 M65
    |    |--N. (M.) horvarthi Friese 1909 M65
    |    |--N. (M.) kochi Friese 1909 M65
    |    |--N. (M.) longicornis Smith 1859 M65
    |    |--N. (M.) luteiventris Friese 1909 M65
    |    |--N. (M.) metallica Smith 1863 M65
    |    |--N. (M.) simillima Smith 1863 M65
    |    |--N. (M.) simplicinotum Michener 1965 M65
    |    `--N. (M.) tomentifera Friese 1909 M65
    |--N. (Pseudapis Kirby 1900) [incl. Nomiapis Cockerell 1919] M65
    |    |--N. (*P.) anomala (Kirby 1900) [=*Pseudapis anomala] M65
    |    |--N. (P.) diversipes Latreille 1806 [=*Nomiapis diversipes] M65
    |    `--N. (P.) lepidota Cockerell 1905 M65
    |--N. (Ptilonomia Michener 1965) M65
    |    `-N. (*P.) plumosa Michener 1965 M65
    |--N. (Reepenia Friese 1909) [incl. Megaloptodes Moure 1958] M65
    |    |--N. (*R.) variabilis Friese 1909 M65
    |    |--N. (R.) bidentata Smith 1863 M65
    |    |--N. (R.) bituberculata (Smith 1853) (see below for synonymy) M65
    |    |--‘Reepenia’ brevicornis R35
    |    |--N. (R.) florea Smith 1863 M65
    |    |--N. (R.) fuliginosa Friese 1909 M65
    |    |--N. (R.) fulviventris Cameron 1901 M65
    |    |--N. (R.) ‘fulviventris’ Friese 1909 non Cameron 1901 M65
    |    |--N. (R.) ‘triangulifera’ Friese 1909 non Vachal 1897 M65
    |    `--N. (R.) zebrae Friese 1909 M65
    `--N. (Rhopalomelissa Alfken 1926) M65
         |--N. (*R.) xanthogastra (Alfken 1926) [=*Rhopalomelissa xanthogaster] M65
         |--N. (R.) abuensis M65
         |--N. (R.) andrennina M65
         |--N. (R.) aureohirta M65
         |--N. (R.) aurifrons M65
         |--N. (R.) basipicta M65
         |--N. (R.) basutorum M65
         |--N. (R.) bicarinata M65
         |--N. (R.) burmica M65
         |--N. (R.) carinicollis M65
         |--N. (R.) ceratina M65
         |--N. (R.) comberi M65
         |--N. (R.) elongata M65
         |--N. (R.) elongatula M65
         |--N. (R.) lautula M65
         |--N. (R.) levicauda M65
         |--N. (R.) fausta M65
         |--N. (R.) floralis M65
         |--N. (R.) fulvinerva M65
         |--N. (R.) halictella Cockerell 1905 M65
         |--N. (R.) kangrae M65
         |--N. (R.) levicauda M65
         |--N. (R.) mediorufa M65
         |--N. (R.) minutula Friese 1909 M65
         |--N. (R.) modesta Smith 1862 [incl. N. clavata Smith 1862 non Halictus clavatus Smith 1853] M65
         |--N. (R.) morata M65
         |--N. (R.) notabilis M65
         |--N. (R.) palavanica M65
         |--N. (R.) patellifera M65
         |--N. (R.) phenacura M65
         |--N. (R.) pseudoceratina Cockerell 1910 M65
         |--N. (R.) rubella M65
         |--N. (R.) triangularis Cockerell 1905 M65
         `--N. (R.) williamsi Cockerell 1930 M65

Nomia (Austronomia) cheesmanae Michener 1965 [=N. nuda Cheesman 1953 non N. australica nuda Rayment 1939] M65

Nomia (Curvinomia Michener 1944) [incl. Paranomia Friese 1897 non Conrad 1860, Paranomina Michener 1944 non Hendel 1907] M65

Nomia (Curvinomia) tetrazona Cockerell 1910 [incl. N. californiensis Michener 1937, *Curvinomia californiensis] M65

Nomia (Reepenia) bituberculata (Smith 1853) [=Megalopta bituberculata, *Megaloptodes bituberculata; incl. Nomia (Reepenia) eboracina Cockerell 1912, Tetralonia testacea Smith 1854] M65

*Type species of generic name indicated

REFERENCES

[E12] Evenhuis, N. L. 2012. Publication and dating of the Exploration Scientifique de l'Algérie: Histoire Naturelle des Animaux Articulés (1846–1849) by Pierre Hippolyte Lucas. Zootaxa 3448: 1–61.

[M65] Michener, C. D. 1965. A classification of the bees of the Australian and South Pacific regions. Bulletin of the American Museum of Natural History 130: 1–362.

[MS01] Mocsáry, A., & V. Szépligeti. 1901. Hymenopterák [Hymenopteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 121–169. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[R35] Rayment, T. 1935. A Cluster of Bees: Sixty essays on the life-histories of Australian bees, with specific descriptions of over 100 new species. Endeavour Press: Sydney.

[WP99] Walter, D. E., & H. C. Proctor. 1999. Mites: Ecology, Evolution and Behaviour. CABI Publishing: Wallingford (UK).

[ZS10] Zborowski, P., & R. Storey. 2010. A Field Guide to Insects in Australia 3^rd ed. Reed New Holland: Sydney.