Belongs within: Proctotrupoidea.
The Proctotrupidae are a group of small wasps that are mostly endoparasites of Coleoptera larvae, rarely of Diptera. Most species have the wing venation reduced, with the radial cell reduced or almost obliterated, but a well-developed venation and radial cell is retained in the fore wing of the Austroserphini. The species Exallonyx trifoveatus is found worldwide as a parasite of Staphylinidae in compost heaps (Naumann 1991).
Characters (from Naumann 1991): Mostly small (3–15 mm); mesosoma without transscutal articulation; metasoma with large syntergum; trochantellus sometimes present on fore leg; T2 and S2 (petiole) and synsternum fused; metasoma in females usually posteriorly down-curved and telescopic; cerci digitiform or absent.
Proctotrupidae [Proctotrupinae, Serphidae]
|--Austroserphini [Austroserphidae] RC74
| |--Austroserphus Dodd 1933 MH11, R54
| | `--*A. albofasciatus Dodd 1933 R54
| `--Acanthoserphus Dodd 1915 RC74, R54
| `--*A. albicoxa Dodd 1915 R54
`--+--Disogmus Foerster 1856 MH11, R54
| |--D. areolator (Haliday 1839) [=Disogmus areolator] R54
| `--D. nigerrimus Dodd 1915 R54
`--+--Exallonyx trifoveatus MH11, N91
`--Proctotrupes Latreille 1796 MH11, R54
|--*P. brevipennis Latreille 1802 R54
|--P. collaris Szépligeti in Mocsáry & Szépligeti 1901 MS01
|--P. doddi Riek 1954 R54
|--P. gravidator MS01
|--P. janthinae Dodd 1915 [=Proctotrypes janthinae] R54
|--P. niger G20
|--P. nitens Dodd 1920 R54
|--P. splendidus Riek 1954 R54
|--P. syagrii R70
`--P. turneri Dodd 1920 R54
Proctotrupidae incertae sedis:
Codrus A71
Phaenoserphus Kieffer in André 1908 R54
|--*P. curtipennis (Haliday 1839) [=Proctotrupes curtipennis] R54
|--P. australiae (Kieffer 1907) [=Proctotrypes australiae] R54
|--P. calcar A71
`--P. viator A71
Labeo pusillus Szépligeti in Mocsáry & Szépligeti 1901 MS01
Prosacantha MS01
|--P. grandis MS01
`--P. spinosa Szépligeti in Moscáry & Szépligeti 1901 MS01
Isocybus MS01
|--I. grandis MS01
`--I. rossicus Szépligeti in Mocsáry & Szépligeti 1901 MS01
Pantoclis MS01
|--P. barycera MS01
`--P. rufipes Szépligeti in Mocsáry & Szépligeti 1901 MS01
Acropiesta MS01
Oxyserphus N91
Austrocodrus N91
Mischoserphus N91
Apoglypha N91
Serphonostus N91
Cryptoserphus Kieffer in André 1907 R54
|--*C. longicalcar Kieffer in André 1908 R54
|--C. nigriscutum Dodd 1915 R54
|--C. nitidus Dodd 1915 R54
`--C. sulcatus Riek 1954 R54
Paracodrus R54
Rasnitsynia huadongensis (Zhang 1985) [=Oligoneuroides huadongensis] RJ93
*Type species of generic name indicated
REFERENCES
[A71] Askew, R. R. 1971. Parasitic Insects. Heinemann Educational Books: London.
[G20] Goldfuss, G. A. 1820. Handbuch der Naturgeschichte vol. 3. Handbuch der Zoologie pt 1. Johann Leonhard Schrag: Nürnberg.
[MS01] Mocsáry, A., & V. Szépligeti. 1901. Hymenopterák [Hymenopteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 121–169. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.
[MH11] Munro, J. B., J. M. Heraty, R. A. Burks, D. Hawks, J. Mottern, A. Cruaud, J.-Y. Rasplus & P. Jansta. 2011. A molecular phylogeny of the Chalcidoidea (Hymenoptera). PLoS One 6 (11): e27023.
[N91] Naumann, I. D. 1991. Hymenoptera (wasps, bees, ants, sawflies). In: CSIRO. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 2 pp. 916–1000. Melbourne University Press: Carlton (Victoria).
[R54] Riek, E. F. 1954. Australian wasps of the family Proctotrupidae (Hymenoptera: Proctotrupoidea). Australian Journal of Zoology 3: 106–117.
[R70] Riek, E. F. 1970. Hymenoptera (wasps, bees, ants). In: CSIRO. The Insects of Australia: A textbook for students and research workers pp. 867–959. Melbourne University Press.
[RC74] Riek, E. F., & J. C. Cardale. 1974. Hymenoptera (wasps, bees, ants). In: CSIRO. The Insects of Australia: A textbook for students and research workers. Supplement 1974 pp. 107–111. Melbourne University Press.
[RJ93] Ross, A. J., & E. A. Jarzembowski. 1993. Arthropoda (Hexapoda; Insecta). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 363–426. Chapman & Hall: London.
Last updated: 18 March 2021.