Conchifera

Isospira bucanioides, frontal view, from Knight et al. (1960).


Belongs within: Spiralia.
Contains: Stenothecoida, Monoplacophora, Cephalopoda, Bivalvia, Rostroconchia, Scoliostomatidae, Caseolus, Leiostyla, Clisospiroidea, Bellerophontoidea, Neomphaliones, Cyclobranchia, Fissobranchia, Neritimorpha, Loxonematoidea, Heterobranchia, Peruneloidea, Cyclophoroidea, Ampullariidae, Viviparoidea, Orthonematoidea, Subulitoidea, Soleniscidae, Meekospiridae, Cerithiimorpha, Hypsogastropoda.

The Conchifera are a major clade containing the majority of molluscs, ancestrally characterised by the presence of a single dorsal shell (though this shell has become reduced or lost in several derived subgroups). Relationships between the several conchiferan classes have historically been contentious; almost every possible arrangement has been proposed at one time or another.

The Gastropoda are the largest of the commonly-recognised molluscan classes. Ancestrally, gastropods have a dorsal, unchambered, often helically coiled, shell (lost in numerous derived groups) and a head with eyes and cephalic tentacles. Gastropods are also distinguished (at least in the crown group) by the process of torsion where, during development, the gut becomes U-shaped and the initially posterior anus is moved to an anterior position (Cox 1960). Whether some early fossil groups such as the Bellerophontacea with coiled but bilaterally symmetrical shells underwent torsion has been subject to some debate; consequently, some authors would dispute the classification of such groups as gastropods. The effects of torsion have also become reduced in some derived subgroups of the gastropods. Earlier classifications united those groups exhibiting full torsion through to maturity as the 'Prosobranchia', a taxon that has now been abandoned as paraphyletic. Also now regarded as paraphyletic are the 'Archaeogastropoda', a previously recognised group of gastropods with the gill ctenidia bilaterally filamented and terminally free, including essentially all living gastropods outside the Apogastropoda, in which the ctenidia become lost, or pectinate with filaments on one side only with the axis adherent to the pallial wall (Knight et al. 1960). Living 'archaeogastropods' have been divided between the Patellogastropoda, Neritimorpha and Vetigastropoda; there are also some small groups, such as the Cocculiniformia, for which there are conflicting opinions as to whether they form a clade with the Vetigastropoda or whether they represent independent lineages.

On the basis of their possession of a radula with a central tooth flanked by a small number of lateral and marginal teeth, similar to those found in some other mollusc classes, the Patellogastropoda have been suggested as the sister group to other living gastropods, though this has been contested by recent molecular analyses (Bouchet et al. 2017). The remaining living gastropods commonly have a more longitudinally flexible radula with a higher number of more differentiated lateral and marginal teeth (though this has been further modified, and sometimes secondarily reduced, in some derived lineages). The Neritimorpha and Apogastropoda are further united by the loss of a number of organs on the right side of the body, such as the right kidney. Further such losses characterise the Apogastropoda alone, such as the loss of the right auricle (Simone 2011).

Conchifera (see below for synonymy) SB12
    |--+--Monoplacophora SW11
    |  `--Cephalopoda SW11
    `--Pleistomollusca [Cyclomya] KC11
         |--Bivalvia SW11
         `--+--Rostroconchia SW11
            `--Gastropoda (see below for synonymy) SW11
                 |  i. s.: Spiniplatyceras FBL02
                 |         Serpentubina Horný 1964 FBL02
                 |         Kiviasukkaan Peel 1975 FBL02
                 |         Pragoserpulina Frýda 1998 [Pragoserpulinidae] BR05
                 |           `--*P. tomasi Frýda 1998 BR17
                 |         Luoguispira Pan & Erwin 2002 PE02
                 |           |--*L. micra Pan & Erwin 2002 PE02
                 |           |--L. lineata Pan & Erwin 2002 PE02
                 |           `--L. multilinea Pan & Erwin 2002 PE02
                 |         Luoguella Pan & Erwin 2002 PE02
                 |           |--*L. elegantula Pan & Erwin 2002 [=Gyrospira elegantula] PE02
                 |           `--L. diana Pan & Erwin 2002 [=Gyrospira diana] PE02
                 |         Propupa PE02
                 |         Extendilabrum PE02
                 |         Tetrabubispira PE02
                 |         Kalchreuthia Gründel & Nützel 1998 GK02
                 |         Paladilha (Lartella Cossmann 1921) GK02
                 |         Pseudotaphrus (Pezantia Cossmann 1896) GK02
                 |         Chlamydarion Pi02
                 |         Derjuginella rufofasciata GAS03
                 |         Erheria vincta GAS03
                 |         Melanochlamis GAS03
                 |           |--M. cylindrica HS01
                 |           `--M. diomedea GAS03
                 |         Yokoyamia ornatissima GAS03
                 |         Epheria turrita KBC03
                 |         Tersiella plicosa KBC03
                 |         Ausola angustata KBC03
                 |         Tegella C79
                 |           |--T. aquilirostris KBC03
                 |           |--T. nigrans KBC03
                 |           `--T. verrucosa C79
                 |         Hinia cuvieri M62
                 |         Neritula donovani M62
                 |         Oxytrema silicula B96
                 |         Hystrivasum horridum Heilprin 1886 HP04
                 |         Benthonella tenella (Jeffreys 1869) OT04
                 |         Chlupacispira spinosa Blodgett & Rohr 1989 FB04
                 |         Eulepetopsis vitrea GO06
                 |         Purkynespira Frýda & Bandel 1997 FB01
                 |           `--P. reticulata FB01
                 |         Streptopelma henchmani Marwick 1926 F27a
                 |         Latirofusus Cossmann 1889 F27c
                 |           |--*L. funiculosus F27c
                 |           |--L. aciformis [=Fusus aciformis] F27c
                 |           |--L. acus [=Fusus acus] F27c
                 |           |--L. lancea [=Fusus lancea] F27c
                 |           |--L. optatus M. & M. 1923 F27a
                 |           `--L. spiceri (Tenison-Woods 1877) [=Fusus spiceri; incl. L. nigrofuscus Tate 1891] F27c
                 |         Buccinulus F27a
                 |           |--B. albus F27a
                 |           |--B. huttoni Kirk 1882 F27a
                 |           `--B. kirki Hutton 1873 [incl. B. gracilis Kirk 1882] F27a
                 |         Triploca Tate 1894 F27a
                 |           |--T. ligata F27a
                 |           `--T. waihaoensis Marshall & Murdoch 1923 F27a
                 |         Leucotina F27a
                 |           |--L. casta (Adams 1853) [=Monoptgyma casta] H09
                 |           |--L. helva Hedley 1900 H09
                 |           `--L. pura (Adams 1855) F27a
                 |         Sublacuna Cossmann 1899 [=Strebloramphus Tate 1898 (preoc.)] F27c
                 |           `--*S. mirula (Tate 1898) [=*Strebloramphus mirulus] F27c
                 |         Coptochetus asperulus [=Fusus asperulus] F27c
                 |         Trifora Deshayes in Blainville 1828 O27b
                 |           |--*T. perversa O27b
                 |           |--T. callipyrga Bartsch 1907 O27b
                 |           |--T. carpenteri Bartsch 1903 O27b
                 |           |--T. catalinensis Bartsch 1907 O27b
                 |           |--T. montereyensis Bartsch 1907 O27b
                 |           |--T. pedroana Bartsch 1907 O27b
                 |           |--T. peninsularis Bartsch 1907 O27b
                 |           `--T. stearnsi Bartsch 1907 O27b
                 |         Barocospira Horný 1964 FBL02
                 |         Ibergia Blodgett & Frýda 1999 FBL02
                 |         Crassimarginatidae BR05
                 |           |--Yukonoconcha Frýda, Blodgett & Lenz 2002 FBL02
                 |           |    `--*Y. pedderi Frýda, Blodgett & Lenz 2002 FBL02
                 |           `--Crassimarginata Jhaveri 1969 BR05
                 |                |--*C. crassicosta Jhaveri 1969 FBL02
                 |                `--C. tenuicosta Jhaveri 1969 FBL02
                 |         Scoliostomatidae BR05
                 |         Levihelix pusilla Gründel 2000 GK02
                 |         Sinuginella pygmaea HS01
                 |         Adelphotectonica reevi HS01
                 |         Bouchetriphora pallida HS01
                 |         Limulatys reliquus HS01
                 |         Pisinna HS01
                 |           |--P. manawatawhia HS01
                 |           |--P. olivacea HS01
                 |           |    |--P. o. olivacea HS01
                 |           |    `--P. o. impressa HS01
                 |           |--P. rekohuana HS01
                 |           |    |--P. r. rekohuana HS01
                 |           |    `--P. r. lactorubra HS01
                 |           |--P. semiplicata HS01
                 |           `--P. zosterophila HS01
                 |         Sagenotriphora ampulla HS01
                 |         Camita rotellina (Gould 1860) [=Trochus (Monodonata) rotellinus] H08
                 |         Turcia H09
                 |           |--T. maculata (Brazier 1877) [=Thalotia maculata] H09
                 |           |--T. monilifera Adams 1855 H09
                 |           `--T. montrouzieri (Fischer 1878) [=Tectaria montrouzieri] H09
                 |         Mecoliotia spinosa Hedley 1902 H09
                 |         Quoyia decollata (Quoy & Gaimard 1833) [=Planaxis decollatus] H09
                 |         Eglisia tricarinata Adams & Reeve 1840 H09
                 |         Couthouyia aculeata Hedley 1900 H09
                 |         Oscilla tasmanica (Ten.-Woods 1877) [=Parthenia tasmanica] H09
                 |         Elusa subulata (Adams 1855) [=Pyramidella subulata] H09
                 |         Scalenostoma striatum Hedley 1900 H09
                 |         Distortrix decipiens (Reeve 1844) [=Triton decipiens] H09
                 |         Radius angasi (Reeve 1865) [=Ovulum angasi] H09
                 |         Laimodonta conica Pease 1862 H09
                 |         Paraplysia piperata (Smith 1884) [=Aplysia piperata] H09
                 |         Euselenops luniceps (Cuvier 1830) [=Pleurobranchus luniceps] H09
                 |         Placomopherus insignis Smith 1884 H09
                 |         Adiozoptyxis Dietrich 1925 [Adiozoptyxinae, Adiozoptyxisinae] BR05
                 |           `--*A. polymorpha (Gemmellaro 1865) [=Nerinea polymorpha] BR17
                 |         ‘Cavolinia’ Bruguière 1791 non Abilgaard 1791 BR05
                 |         Crenea Risso 1826 BR05
                 |         Cypraeogemmula Vredenburg 1920 [Cypraeogemmulinae, Cypraeogemmulini] BR05
                 |           `--*C. scabriuscula (Koenen 1890) BR17 (see below for synonymy)
                 |         Incrispella Tasch 1963 [Incrispellidae] BR05
                 |           `--*I. rectotortis Tasch 1963 BR17
                 |         Japeuthria Iredale 1918 [Japeuthriinae] BR05
                 |           `--*J. ferrea (Reeve 1847) [=Buccinum ferreum] BR17
                 |         Kanamarua Kuroda 1951 [Kanamariidae, Kanamaruidae] BR05
                 |           `--*K. adonis (Dall 1919) BR17 [=Colus adonis BR17, C. (Anomalosipho) adonis O27a]
                 |         Ladamarekia Horný 1992 [Ladamarekiidae] BR05
                 |           `--*L. miranda Horný 1992 BR17
                 |         Megalostominae BR05
                 |           |--Cataulus BR05
                 |           `--Nicida BR05
                 |         Melanioptyxis Cossmann 1896 [Melanioptyxinae] BR05
                 |           `--*M. altararis (Cossmann 1885) [=Nerinea altararis] BR17
                 |         Neoptyxis Wenz 1940 [Neoptyxidae] BR05
                 |           `--*N. astrachanica (Rebinder 1902) [=Nerinea astrachanica] BR17
                 |         Oligoptyxis Pchelintsev 1953 [Oligoptyxidae] BR05
                 |           `--*O. turricula Pchelintsev 1953 BR17
                 |         Yuopisthonema Nützel in Bouchet, Rocroi et al. 2017 BR17 (see below for synonymy)
                 |           `--*Y. undulatum (Yu 1974) [=*Opisthonema undulatum] BR17
                 |         Palaeocyclophorus Wenz 1923 BR05
                 |         Pentaptyxis Pchelintsev 1965 [Pentaptyxidae] BR05
                 |           `--*P. staszycii (Zeuschner 1849) [=Acteon staszycii] BR17
                 |         Plesioplocus Pchelintsev 1953 [Plesioplocidae] BR05
                 |           `--*P. grandis Pchelintsev 1953 BR17
                 |         Polyodonte Fischer 1807 BR05
                 |         Polyodonta Megerle 1811 BR05
                 |         Protoconchoididae [Protoconchioididae] BR05
                 |           |--Patelliconus Horný 1961 BR17, BR05 [Patelliconidae]
                 |           |    `--*P. primulus (Perner 1903) [=Palaeacmaea primula] BR17
                 |           `--Protoconchoides Shaw 1962 [=Protoconchioides (l. c.)] BR05
                 |                |--*P. hermitensis (Resser 1945) [=Scenella hermitensis] BR17
                 |                `--P. conula TS11
                 |         Pseudobythinella Melville 1956 BR05
                 |         Rhytidopilus Cossmann 1895 [=Rhytidophilus (l. c.); Rhytidophilidae, Rhytidopilidae] BR05
                 |           `--*R. humbertinus (Buvignier 1852) [=Patella humbertina] KC60
                 |         Styliferina Adams 1860 [Styliferinidae] BR05
                 |           `--*S. goniochila Adams 1860 BR17 [=Alaba goniochila H09]
                 |         Stylocheilus Gould 1852 [Stylocheilinae, Stylochiniae] BR05
                 |           `--*S. lineolatus (Gould 1852) [=Aplysia lineolatus] BR17
                 |         Transovula de Gregorio 1880 [Transovulini] BR05
                 |           `--*T. schefferi (de Gregorio 1880) [=Ovula schefferi] BR17
                 |         Papustyla pulcherrima USDI77
                 |         Acilia franciscae TE01
                 |         Anisus L06
                 |           |--A. spirorbis (Linnaeus 1758) L06
                 |           `--A. vortex (Linnaeus 1758) L06
                 |         Bathyomphalus contortus (Linnaeus 1758) L06
                 |         Caseolus CC06
                 |         Leiostyla CC06
                 |         Eucobresia media (Lowe 1854) CC06
                 |         Heterostoma Filippi 1837 CC06, LT64 [=Heterostomum Diesing 1850 LT64]
                 |           `--H. paupercula (Lowe 1831) [incl. Steenbergia duplex] CC06
                 |         Spirorbula CC06
                 |           |--S. depauperata (Lowe 1831) [incl. S. latinea (Paiva 1866)] CC06
                 |           |--S. obtecta CC06
                 |           `--S. squalida CC06
                 |         Plagyrona placida (Shuttleworth 1852) CC06
                 |         Staurodon saxicola CC06
                 |         Actinella CC06
                 |           |--A. arcinella CC06
                 |           |    |--A. a. arcinella CC06
                 |           |    `--A. a. papillosa CC06
                 |           |--A. crassiuscula CC06
                 |           |--A. effugiens CC06
                 |           |--A. morenensis CC06
                 |           `--A. promontoriensis CC06
                 |         Pseudocampylaea CC06
                 |           |--P. lowei CC06
                 |           `--P. portosanctana CC06
                 |         Amphorella CC06
                 |           |--A. cimensis CC06
                 |           |--A. gracilis CC06
                 |           |--A. melampoides CC06
                 |           |--A. oryza CC06
                 |           |--A. terebella CC06
                 |           |--A. triticea CC06
                 |           `--A. tuberculata CC06
                 |         Cylichnidia ovuliformis CC06
                 |         Lemniscia michaudi CC06
                 |         Eufenella pupoides (Adams 1860) BD86
                 |         Clathrofenella reticulata (Adams 1860) BD86
                 |         Heliosoma trivolvis AS07
                 |         Allocharopa belli SC07
                 |         Probaicalia ZZ88
                 |           |--P. gerassimovi ZZ88
                 |           |--P. rammelmeyeri ZZ88
                 |           `--P. vitimensis ZZ88
                 |         Brotiopsis wakinoensis ZZ88
                 |         Tulotomoides talatziensis ZZ88
                 |         Gyrodema etheridgei F71
                 |         Magelia bidens Tenison-Woods 1878 F71
                 |         Odontostomia cretacea Etheridge 1907 F71
                 |         Lindoliva spengleri Petuch 1988 P89
                 |         Armiger crista (Linnaeus 1758) P88
                 |         Stagnicola P88
                 |           |--S. caperata (Say 1829) P88
                 |           |--S. catascopium (Say 1817) P88
                 |           |--S. palustris Muller 1774 P88
                 |           `--S. reflexa (Say 1821) P88
                 |         Probythinella lacustris (Baker 1928) P88
                 |         Bulimnea megasoma (Say 1824) P88
                 |         Japonia AA88
                 |           |--J. ciliata (Sowerby 1843) AA88
                 |           `--J. stephanophora (Moellendorff 1895) AA88
                 |         Gymnentome valida H75
                 |         Ptychostylus guimarotensis Bandel 1991 B01
                 |         Melampoides jurassicus Bandel 1991 B01
                 |         Ittibittium Houbrick 1993 BC01
                 |           `--I. turriculum (Usticke 1969) (see below for synonymy) BC01
                 |         Torrella Henderson & Bartsch 1920 BC01
                 |           `--*T. torreiana (Gundlach ex Arango y Molina 1878) [=Ctenopoma torreianum] BC01
                 |         Pseudotorinia retifera (Dall 1892) [incl. Heliacus crystallina Usticke 1969] BC01
                 |         Bullata Jousseaume 1875 BC01
                 |           `--B. matthewsi (van Mol & Tursch 1967) [=Marginella (Prunum) matthewsi] BC01
                 |         Elimia Adams & Adams 1854 BC01
                 |           `--E. laeta (Jay 1839) [=Melania laeta] BC01
                 |         Boonea impressa (Say 1822) [incl. Odostomia (Menestho) beauforti Jacot 1921] BC01
                 |         Fargoa bushiana (Bartsch 1909) [incl. Odostomia charlottae Usticke 1969] BC01
                 |         Mesophora Laseron 1958 BC01
                 |           `--M. novem (Usticke 1969) [=Triphora novem] BC01
                 |         Eutriphora bermudensis (Bartsch 1911) [incl. Triphora turrissimilis Usticke 1969] BC01
                 |         Aroapyrgus Baker 1931 BC01
                 |           `--A. dubiosus (Adams 1852) [=Truncatella dubiosa] BC01
                 |         Chippewaella patellitheca W01
                 |         Moerkeia Böhm 1895 NE02
                 |         Greggelix indigena C90
                 |         Torquilla frumentum D56
                 |         Australorbis odhner NG13
                 |         Sinopupoides hubeiensis NG13
                 |         Bulgarica thessalonica (Olivier 1801) O03
                 |         Palleseria longwelli C60
                 |         Clavilithes rugosus C60
                 |         Partulida spiralis C60
                 |         Ampullella bulimoides C60
                 |         Cepatia cepacea C60
                 |         Palaeocyclotus exaratus C60
                 |         Pseudorhytidopilus Cox in Knight, Cox et al. 1960 KC60
                 |           `--*P. lennieri Cox in Knight, Cox et al. 1960 (see below for synonymy) KC60
                 |         Atlantobellerophon Trechmann 1930 KC60
                 |           `--*A. zealandicus Trechmann 1930 KC60
                 |         Burdikinia Knight 1937 [=Polyamma Etheridge 1917 non Kriechbaumer 1894] KC60
                 |           `--*B. burdikiensis (Etheridge 1917) [=*Polyamma burdikiensis] KC60
                 |         Cinctaspira Powell 1933 KC60
                 |           `--*C. conica Powell 1933 KC60
                 |         ‘Conchula’ Steininger 1849 non Herrmannsen 1847 (n. d.) KC60
                 |           `--*C. cylindracea Steininger 1849 (n. d.) KC60
                 |         ‘Geinitzia’ Dietz 1911 nec Gemmellaro 1892 nec Handlirsch 1906 KC60
                 |           `--*G. carinata Dietz 1911 KC60
                 |         Kebina Vologdin 1955 KC60
                 |           `--*K. pulchra Vologdin 1955 KC60
                 |         Ozarkina Ulrich & Bridge 1931 [=Ozarkispira Ulrich & Bridge 1931 non Walcott 1924] KC60
                 |           `--*O. typica (Ulrich & Bridge 1931) [=*Ozarkispira typica] KC60
                 |         Ozarkocanus Heller 1956 KC60
                 |           `--*O. prearcuatus Heller 1956 KC60
                 |         Pondia Oder 1932 KC60
                 |           `--*P. powelli Oder 1932 KC60
                 |         Pythmenema Lamont & Gilbert 1945 KC60
                 |           `--*P. praenantium (Phillips 1848) [=Euomphalus praenantius] KC60
                 |         Trochotremaria Ryckholt 1860 KC60
                 |         Umbotrochus Perner 1903 [=Umbonitrochus Cossmann 1918] KC60
                 |           `--*U. aspersus Perner 1903 KC60
                 |         Acevina Rusconi 1952 KC60
                 |           `--*A. cuyunchensis (Rusconi 1952) [=Helcionella (*Acevina) cuyunchensis] KC60
                 |         Odontosagda V18
                 |           |--O. abbotti Vanatta 1918 V18
                 |           |--O. blandi V18
                 |           `--O. hilli V18
                 |         Aplysiopsis smithi (Marcus 1961) B77
                 |         Volvaria ficoides H86
                 |         Cassidaria H86
                 |           |--C. senex [=Struthiolaria senex] H86
                 |           `--C. sulcata H86
                 |         Sorapisella Bandel 1993 B00
                 |         Triadoskenea Bandel 1993 B00
                 |           `--T. ampezzana Bandel 1993 B02
                 |         Rolandomphalus Schwardt 1992 B00
                 |         Macrophragmus tokyoensis K10
                 |         Fascioplex neozelanica K95
                 |         Pseudoaclisina Yoo 1994 B02
                 |           |--*P. turgida [=Aclisina turgida] B02
                 |           `--P. microspirulata B02
                 |         Palaeoalvania Yoo 1994 B02
                 |           `--*P. talenti (Yoo 1994) [=Aclisina talenti] B02
                 |         Sinistriconcha Heidelberger & Bandel 1999 B02
                 |           |--*S. lierli Heidelberger & Bandel 1999 B02
                 |           `--S. minutissima (Yoo 1988) B02
                 |         Iberus E99
                 |           |--I. dupuydelomei E99
                 |           `--I. marmoratus C84
                 |         Carinifex (Carinifex) newberryi LK84
                 |         Amycla corniculum PP64
                 |         Ocinebra PP64
                 |           |--O. aciculata PP64
                 |           `--O. edwardsi PP64
                 |         Scabrella Sacco 1890 H22
                 |           `--*S. scabra [=Columbella scabra] H22
                 |         Pseudaulicina Chavan 1948 BR17
                 |           `--*P. musicalis (Lamarck 1803) [=Voluta musicalis, Volutilithes musicalis] BR17
                 |         Kimina Yoo 1994 B02
                 |           |--*K. globosa Yoo 1994 B02
                 |           |--K. australis Yoo 1994 B02
                 |           |--K. minor Yoo 1994 B02
                 |           `--K. yooi Frýda & Bandel 1997 B02
                 |         Harperispira Bandel 2002 BR17
                 |         Kiringella Rozov 1968 W01, RJ76
                 |           |--K. kultavasaensis RJ76
                 |           `--K. pyramidalis W01
                 |         Palaeoniso brazieri SS16
                 |         Burdekina JB12
                 |         Natria JB12
                 |         Peraubium JB12
                 |         Stusakia pulchra F12
                 |         Mesolanistes V-RR93
                 |           |--M. bajanohongorensis V-RR93
                 |           |--M. debilicostata V-RR93
                 |           `--M. shuwalovi V-RR93
                 |        Severynella trochlearis Hynda 1983 D94
                 |        Lacunina Kittl 1899 TTE93
                 |        Bullopsis Conrad 1858 TTE93
                 |        Serradonta Okutani, Tsuchida & Fujikara 1992 AJH07
                 |        Pleuroprocta silvatica PB27
                 |        Trichotoxon PB27
                 |          |--T. maculatum PB27
                 |          |    |--T. m. maculatum PB27
                 |          |    `--T. m. perforatum PB27
                 |          |--T. pardus PB27
                 |          `--T. ruwenzoriense PB27
                 |         Bocageia PB27
                 |           |--B. germaini PB27
                 |           |--B. interioris PB27
                 |           |--B. liocephala PB27
                 |           `--B. runssorina PB27
                 |         Africarion PB27
                 |           |--A. copiosus PB27
                 |           |--A. spatiosus PB27
                 |           `--A. tenebrosus PB27
                 |         Leucochroopsis E99
                 |           |--L. apicalis E99
                 |           |--L. dufrenoyi E99
                 |           |--L. kleini E99
                 |           |--L. leptoloma E99
                 |           |--L. pisum E99
                 |           `--L. valentinensis E99
                 |                |--L. v. valentinensis E99
                 |                `--L. v. sermenazensis E99
                 |         Wenzia ramondi E99
                 |         Megalotachea E99
                 |           |--M. christoli E99
                 |           |--M. delphinensis E99
                 |           |    |--M. d. delphinensis E99
                 |           |    `--M. d. tersannensis E99
                 |           |--M. gualinoi E99
                 |           `--M. turonensis E99
                 |         Tudorella E99
                 |           |--T. baudoni E99
                 |           `--T. draparnaudi E99
                 |                |--T. d. draparnaudi E99
                 |                `--T. d. minor E99
                 |         Pseudoleacina (Paraglandina) E99
                 |           |--P. (P.) christoliana E99
                 |           `--P. (P.) oligostropha E99
                 |         Pseudochloritis E99
                 |           |--P. incrassatus E99
                 |           `--P. mollonensis E99
                 |         Galactochilus E99
                 |           |--G. leobersdorfensis E99
                 |           `--G. locardi E99
                 |         Negulus E99
                 |           |--N. bleicheri E99
                 |           |--N. raricostatus E99
                 |           |--N. suturalis E99
                 |           |    |--N. s. suturalis E99
                 |           |    `--N. s. gracilis E99
                 |           |--N. truci E99
                 |           `--N. villafranchianus E99
                 |         Eostrobilops E99
                 |           |--E. aloisii E99
                 |           |--E. fischeri E99
                 |           `--E. patuliformis E99
                 |         Macrozonites E99
                 |           |--M. casteti E99
                 |           `--M. collongeoni E99
                 |         Fortuna seringi E99
                 |         Truciella ballesioi E99
                 |         Protodrepanostoma E99
                 |           |--P. bernardii E99
                 |           |--P. nordsiecki E99
                 |           `--P. plioauriculatum E99
                 |         Monoptychia monoptyx E99
                 |         Serruluna anodon E99
                 |         Cladopoda H79
                 |           |--C. directa H79
                 |           `--C. monilifera H79
                 |--Clisospiroidea F12
                 |--Bellerophontacea [Belleromorpha, Bellerophontina, Fissidorsata, Planspiralia, Prorhipidoglossa] KC60
                 |    |  i. s.: Patellostium Waagen 1880 (n. d.) KC60
                 |    |           `--*P. macrostoma (Roemer 1844) [=Bellerophon macrostoma] KC60
                 |    |         Euphemitella Tasch 1953 (n. d.) KC60
                 |    |           `--*E. emrichi Tasch 1953 KC60
                 |    |--Bellerophontoidea BR17
                 |    `--Isospira Koken 1897 KC60 [Isospiridae, Isospiroidea BR05]
                 |         `--*I. bucanioides Koken 1897 BR17
                 `--+--Neomphaliones BR17
                    `--+--+--Cyclobranchia BR17
                       |  `--Fissobranchia S11
                       `--Adenogonogastropoda [Branchiata, Ectophthalma, Teleohydrophila] S11
                            |--Neritimorpha S11
                            `--Apogastropoda (see below for synonymy) PC08
                                 |  i. s.: Protostylus Mansuy 1914 (n. d.) KC60
                                 |           `--*P. lantenoisi Mansuy 1914 KC60
                                 |         ‘Cyclora’ imbricata Whiteaves 1881 G88
                                 |         ‘Cyclora’ valvatiformis Whiteaves 1881 G88
                                 |--Loxonematoidea PC08
                                 `--+--Heterobranchia PC08
                                    `--Caenogastropoda (see below for synonymy) PC08
                                         |  i. s.: Rhabdocolpus (Infacerithium) klebyensis Gründel 1974 GK02
                                         |         Gaeatrivia millardi Cate 1979 M03
                                         |         Serratovolva dondani Cate 1964 M03
                                         |         Decorochilina Frýda 1999 F01
                                         |         Plicatus Pan & Erwin 2002 [Plicatusidae] BR05
                                         |           |--*P. scalaris Pan & Erwin 2002 PE02
                                         |           `--P. multifilaris Pan & Erwin 2002 PE02
                                         |         Spanionema Whidborne 1891 [Spanionematidae, Spanionematoidea] BR05
                                         |           `--*S. scalaroides (Whidborne 1889) [=Loxonema scalaroides] KC60
                                         |         Spirochrysalis Kittl 1894 NE02
                                         |           `--S. protomicra Pan 1982 NE02
                                         |         Lampanella minima PC08
                                         |         Pragoscutula Frýda 1998 PC08, BR05 [Pragoscutulidae]
                                         |           `--P. wareni Frýda 1998 FB01
                                         |         Dendropupoidea (see below for synonymy) KC60
                                         |           |--Maturipupa Pilsbry 1926 KC60
                                         |           |    `--*M. vermilionensis (Bradley 1872) (see below for synonymy) KC60
                                         |           |--Anthracopupa Whitfield 1881 (see below for synonymy) BR05
                                         |           |    |--*A. ohioensis Whitfield 1881 KC60
                                         |           |    `--A. britannica Cox 1926 TTE93
                                         |           `--Dendropupa Owen 1859 BR17 (see below for synonymy)
                                         |                |--*D. vetusta (Dawson 1859) [=Pupa vetusta] KC60
                                         |                |--D. grandaeva (Dawson 1880) TTE93
                                         |                `--D. primaeva (Matthew 1895) TTE93
                                         |         Acanthonema Grabau in Sherzer & Grabau 1909 BR05 (see below for synonymy)
                                         |             `--A. newberryi (Meek 1871) TTE93 (see below for synonymy)
                                         |         Ampezzanildidae BR17
                                         |           |--Cassianilda Bandel 1995 NE02
                                         |           |--Stuorilda Bandel 1995 NE02
                                         |           `--Ampezzanilda Bandel 1994 BR05
                                         |                `--*A. aialensis (Zardini 1980) [=Promathildia aialensis] BR17
                                         |         Kittlidiscus Haas 1953 BR05 (see below for synonymy)
                                         |           `--K. bronni (Klipstein 1845) KC60 (see below for synonymy)
                                         |         Spirostylidae [Spirostylinidae] TE01
                                         |           |--Spirostylus Kittl 1894 TE01
                                         |           |    |--*S. subcolumnaris (Münster 1841) (see below for synonymy) TE01
                                         |           |    `--S. linctus (Böhm 1895) (see below for synonymy) TE01
                                         |           `--Climacina TTE93
                                         |                |--C. catharinae Gemmellaro 1876 TTE93
                                         |                `--C. mariae Gemmellaro 1876 TTE93
                                         |         Peruneloidea BR17
                                         |--Cyclophoroidea S11
                                         `--Hydrogastropoda S11
                                              |--Ampullariidae S11
                                              `--Epiarthroidea [Teleobranchia, Vivipariformes, Vivipariformii] S11
                                                   |--Viviparoidea S11
                                                   `--Sorbeoconcha (see below for synonymy) S11
                                                        |  i. s.: Globocornus Espinosa & Ortea 2010 [Globocornidae] BR17
                                                        |           `--*G. darwini Espinosa & Ortea 2010 BR17
                                                        |         Acteonina d’Orbigny 1850 BR05 (see below for synonymy)
                                                        |           |--*A. carbonaria (de Koninck 1843) (see below for synonymy) BR05
                                                        |           |--A. acuta (d’Orbigny 1841) [=Tornatina acuta] BR17
                                                        |           `--A. obesa Stoliczka 1868 TTE93
                                                        |         Orthonematoidea BR17
                                                        |         Subulitina [Subulitata] GS75
                                                        |           |--Subulitoidea GS75
                                                        |           `--Soleniscoidea BR17
                                                        |                |--Soleniscidae BR17
                                                        |                |--Meekospiridae BR17
                                                        |                `--Anozygidae BR17
                                                        |                     |--Anozyga Hoare 1980 [Anozyginae] BR05
                                                        |                     |    `--*A. bulla Hoare 1980 BR17
                                                        |                     `--Tmetonema Longstaff 1912 (see below for synonymy) BR05
                                                        |                          `--*T. subsulcatum Longstaff 1912 KC60
                                                        |         Pluviostilla BR17
                                                        |--Cerithiimorpha PC08
                                                        `--Hypsogastropoda PC08

Conchifera incertae sedis:
  Canopoconus EL11
  Stenothecoida R96
  Aeolidiopsis Pruvot-Fol 1956 [Aeolidiopsidae] BR17
  Archaeotremaria Yu 1979 (n. d.) [Archaeotremariidae, Archaeotremarioidea] BR17
    `--*A. polytremata Yu 1979 (n. d.) BR17
  Vitrella Clessin 1877 BR17
  Crossopoma Martens 1891 BR17
    `--*C. planorbulum (Lamarck 1816) (nom. cons) (see below for synonymy) BR17
  ‘Electra’ Albers 1850 non Lamouroux 1816 BR17
    `--*E. ceylanica (Pfeiffer 1845) [=Achatina ceylanica] BR17
  Granoconus Yu 1979 (n. d.) [Granoconidae] BR17
    `--*G. trematus Yu 1979 (n. d.) BR17
  *Xerosecta explanata (Müller 1774) [=Helix explanata] BR17
  Peruinia Polinski 1922 [Peruiniinae, Peruiniini] BR17
    `--*P. peruana (Troschel 1847) [=Clausilia peruana] BR17
  Eucycloidea pulchella (d’Orbigny 1850) [=Purpurina pulchella] BR17
  Thoralispira laevis W01
  Kepinospira Yü 1963 RJ76
  Pseudotheca Redlich 1899 RJ76
  Siluracmaea Horny 1963 RJ76
  Sinkiangodiscus Yü 1963 RJ76
  Sinobellerophon Yü 1962 RJ76
  Sinosinuites Yü 1963 RJ76

Acanthonema Grabau in Sherzer & Grabau 1909 BR05 [Acanthonematidae BR17, Acanthonematinae]

Acanthonema newberryi (Meek 1871) TTE93 [=Orthonema newberryi KC60; incl. *A. holopiforme Grabau 1909 KC60, BR17]

Acteonina d’Orbigny 1850 BR05 [=Actaeonina (l. c.) BR05; Acteoninidae, Acteonininae, Acteoninoidea BR17]

*Acteonina carbonaria (de Koninck 1843) [=Chemnitzia carbonaria] BR05

Anthracopupa Whitfield 1881 [Anthracopupidae, Anthracopupinae] BR05

Apogastropoda [Anaclodonta, Architectonicoida, Chalazaeata, Ctenobranchia, Davisianoidei, Dioecia, Discopoda, Ectobranchia, Entobranchia, Entomotaeniata, Gymnoglossa, Heterogastropoda, Heterostropha, Holonephridia, Homoestropha, Hydrocenoidei, Iniophthalma, Loxonematacea, Monopleurobranchia, Monotocardia, Murchisoniata, Musioglossata, Oligopteria, Orbacea, Paludinimorpha, Pectinibranchia, Pectinibranchiata, Peristomacea, Peristomania, Peristomidae, Potamophila, Procaenogastropoda, Ptenobranchiata, Pyramidellida, Pyramidellimorpha, Pyramidelliones, Pyramidelloida, Rissoellina, Solenisciformes, Spartaebranchia, Subulitiformes, Taenioglossa, Trachelobranchia, Volumina] PC08

Caenogastropoda [Architaenioglossa, Architaenioglossae, Basiophthalma, Cyclophoracea, Cyclophoroidei, Fusiformia, Globularioidei, Mesogastropoda, Mesogastropodida, Opisthophthalma, Palaeocyclophoridae, Procyclophoridae, Procyclophoroidea, Rostrifera, Teleogeophila] PC08

Conchifera [Ancyropoda, Cephalaea, Cyrtonellea, Cyrtonellina, Cyrtosoma, Diasoma, Ganglioneura, Loboconcha, Perocepala, Rhacopoda, Tubicolae, Uberes, Visceroconcha] SB12

*Crossopoma planorbulum (Lamarck 1816) (nom. cons) [=Cyclostoma planorbulum, Cyclotus planorbulus, Cyclotus planorbutus (l. c.); incl. Crossopoma cornuvenatorium (Gmelin 1791)] BR17

*Cypraeogemmula scabriuscula (Koenen 1890) BR17 [=Trivia scabriuscula BR17, Cypraea scabriuscula H09]

Dendropupa Owen 1859 BR17 [incl. Strophella Dawson 1895 KC60, Strophites Dawson 1880 non Deshayes 1832 KC60; Dendropupidae]

Dendropupoidea [Anthracopupoidea, Dendropupinae, Procyclophoroida] KC60

Gastropoda [Abranchiata, Alata, Anentomostomata, Anisobranchia, Anisopleura, Aponotoneura, Apterygia, Archaeogastropoda, Archaeoprosobranchia, Arthrocochlides, Asiphonobranchia, Asiphonata, Aspidobranchia, Auxogastropoda, Chiastoneura, Chismobranchiata, Clypeaceae, Cocculinacea, Cocculiniformia, Cochliostraca, Conivalvia, Cricostomata, Ctenobranchiata, Dicoeca, Dicranobranchia, Diotocardia, Eogastropoda, Fissuracea, Fornices, Galeroconcha, Gasteropoda, Gasteropodophora, Gasterozoa, Gastraeopoda, Gastropodea, Goniostomata, Gonyostomata, Gymnopoda, Helcionellina, Hemicyclostoma, Holostomata, Hyperstrophina, Lepadophora, Lepetellina, Limaces, Neurobranchia, Operculata, Orthogastropoda, Otidea, Paracephalophora, Paragastropoda, Patellina, Peristomatacea, Peristomatidae, Platyceratacea, Podophthalma, Prosobranchia, Prosobranchiata, Pselaphocephala, Raphidoglossa, Rhipidoglossa, Riphidoglossa, Scutibranchia, Scutiformia, Sinuata, Sinuopeoidei, Solenostomata, Streptoneura, Syncephala, Tectipeda, Triforidoidei, Trochina, Trochomorpha, Trochomorphi, Turbinimorpha, Xenophoroidei] SW11

Ittibittium turriculum (Usticke 1969) [=Bittium varium f. turricula, Alabina turricula, B. turriculum, Cerithiodes turricula] BC01

Kittlidiscus Haas 1953 BR05 [=Schizodiscus Kittl 1891 non Hall & Clarke 1888 KC60; Kittlidiscidae BR17]

*Kittlidiscus bronni (Klipstein 1845) KC60 [=Pleurotomaria bronni KC60, Schizodiscus bronni KC60; incl. P. plana Klipstein 1845 non Münster 1844 KC60, *Kittlidiscus planus BR17, *Schizodiscus planus BR17]

*Maturipupa vermilionensis (Bradley 1872) [=Pupa vermilionensis] KC60

*Pseudorhytidopilus lennieri Cox in Knight, Cox et al. 1960 [incl. Helcion castellana Lennier 1868 non Patella castellana Thurmann & Étallon 1861] KC60

Sorbeoconcha [Anandria, Aulobranchiata, Brevicommisurata, Eriophthalma, Neotaenioglossa, Omalaxoidea, Palaeocaenogastropoda, Siphonobranchei, Siphonobranchia, Siphonobranchiata, Tubispirantia]

Spirostylus linctus (Böhm 1895) [=Omphaloptycha lincta; incl. O. porrecta Böhm 1895] TE01

*Spirostylus subcolumnaris (Münster 1841) [=Melania subcolumnaris] TE01

Tmetonema Longstaff 1912 [Tmetoneminae] BR05

Yuopisthonema Nützel in Bouchet, Rocroi et al. 2017 BR17 [=Opisthonema Yu 1974 non Gill 1862 BR05; Opisthonematidae, Yuopisthonematidae]

*Type species of generic name indicated

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