Mnium hornum, from here.

Belongs within: Bryidae.

The Mniaceae are a family of mosses, with representatives known as thread-mosses and thyme-mosses, with their main diversity in the northern temperate zone. Members of Mniaceae can be distinguished from many (but not all) species of the similar Bryaceae by the absence in the Mniaceae of comal tufts of leaves (Watson & Dallwitz).

Characters (from Smith & Smith 2004): Dioicous or synoicous. All stems erect or fertile stems erect and sterile stems procumbent or arcuate; central strand present. Leaves pellucid, ovate to linear-lanceolate, acuminate to rounded; margins unbordered or with one- to several-stratose border, entire or dentate with single or double teeth; costa with stereid band in section; cells more or less quadrate to linear, smooth, sometimes in divergent rows. Female inflorescences bud-like, males discoid or not. Sporophytes sometimes more than one per perichaetium; setae long; capsules cernuous to pendulous, ovoid to cylindrical; peristome double; calyptrae cucullate.

    |--Orthomnion (Mitt.) Broth. 1905 SK02 [Orthomnieae W79]
    |    `--O. elimbatum (Nog.) Kop. 1980 [incl. Mnium rotundifolium] SK02
    |--Plagiomnium Kop. 1968 SK02 [Plagiomnieae W79]
    |    |--P. cuspidatum FHH01
    |    |--P. ellipticum SK02
    |    |--P. novae-zealandiae (Colenso) Kop. 1977 (see below for synonymy) SK02
    |    |--P. rhynchophorum J87
    |    |--P. rostratum [=Mnium rostratum; incl. M. longirostrum] SK02
    |    `--P. undulatum SF01
    `--+--Cinclidieae W79
       `--Mnium Hedw. 1801 SK02 [Mnieae W79]
            |--M. cuspidatum N02
            |--M. hornum FHH01
            |--M. integrum N02
            |--M. japonicum N02
            |--M. leptolimbatum D03
            `--M. pseudopunctatum N02

Mniaceae incertae sedis:
  Schizymenium Harv. ex Hooker 1840 SK02
    `--S. bryoides Harv. ex Hooker 1840 (see below for synonymy) SK02
  Mielichhoferia Nees & Hornsch. 1831 SK02
    |--M. forsythii Broth. 1916 SK02
    |--M. mielichhoferi N02
    |--M. spegazziniana D03
    |--M. sullivanii Müll.Hal. 1898 SK02
    `--M. tenuiseta M49
  Pohlia Hedw. 1801 SK02
    |--P. bulbifera (Warnst.) Warnst. 1904 SK02
    |--P. clavaeformis (Hampe) Broth. 1903 [=Bryum clavaeforme, Webera clavaeformis] SK02
    |--P. cruda (Hedw.) Lindb. 1879 SK02 (see below for synonymy)
    |--P. cuspidata Bartram 1951 SK02
    |--P. elongata N02
    |--P. flexuosa N02
    |--P. longicolla N02
    |--P. mielichhoferia (Müll.Hal.) Broth. 1903 [=Bryum mielichhoferia, Webera mielichhoferia] SK02
    |--P. minor N02
    |--P. novae-seelandiae Dixon 1915 SK02
    |--P. nutans (Hedw.) Lindb. 1879 (see below for synonymy) SK02
    |--P. rigescens N02
    |--P. tenuifolia (Jaeger) Broth. 1903 SK02
    |--P. turgens (Broth.) Shaw 1987 [=Mielichhoferia turgens Broth. 1916] SK02
    `--P. wahlenbergii (Weber & Mohr) Andrews 1935 (see below for synonymy) SK02

Nomina nuda: Leptochlaena microdonta Mitt. ex Muell. 1880 [=Mielichhoferia microdonta Mitt. 1882] SK02
             Mnium alpestre Müll.Hal. 1901 SK02
             Pohlia austrocarnea Broth. ex Ramsay 1984 SK02

Plagiomnium novae-zealandiae (Colenso) Kop. 1977 [incl. Mnium novae-valesiae Müll.Hal. 1876 (n. n.), M. tasmanicum Watts & Whitel. 1902 (nom. inv.)] SK02

Pohlia cruda (Hedw.) Lindb. 1879 SK02 [=Webera cruda SK02; incl. W. erythrocaulis SK02, Bryum erythrocaule (preoc.) SK02, Pohlia erythrocaulis SK02, W. synoico-cruda D03]

Pohlia nutans (Hedw.) Lindb. 1879 [=Webera nutans, Bryum nutans; incl. Pohlia austronutans (n. n.), B. beccarii, P. beccarii, Webera beccarii, Bryum leptopelmatum, Pohlia leptopelmata, Webera leptopelmata, W. longifolia, Bryum longifolium (preoc.), Pohlia longifolia, B. montanum, P. montana, Webera montana] SK02

Pohlia wahlenbergii (Weber & Mohr) Andrews 1935 [=Mniobryum wahlenbergii; incl. Bryum albicans (nom. illeg.), M. albicans (nom. illeg.), Pohlia albicans (nom. illeg.), Webera albicans (nom. illeg.), Mniobryum wahlenbergii var. tasmanicum, Pohlia albicans var. tasmanica, P. cruda var. tasmanica, Webera tasmanica] SK02

Schizymenium bryoides Harv. ex Hooker 1840 [=Mielichhoferia bryoides (Harv. ex Hooker) Wijk & Margad. 1962; incl. M. australis, M. ecklonii] SK02

*Type species of generic name indicated


[D03] Dusén, P. 1903. Patagonian and Fuegian mosses. In Reports of the Princeton University Expeditions to Patagonia, 1896-1899, vol. 8 – Botany (W. B. Scott, ed.) pp. 63-126. The University: Princeton (New Jersey).

[FHH01] Frey, W., M. Hofmann & H. H. Hilger. 2001. The gametophyte-sporophyte junction: Unequivocal hints for two evolutionary lines of archegoniate land plants. Flora 196: 431-445.

[J87] Judd, W. S. 1987. Floristic study of Morne La Visite and Pic Macaya National Parks, Haiti. Bulletin of the Florida State Museum – Biological Sciences 32 (1): 1-136.

[M49] Martin, W. 1949. Distribution of the mosses indigenous to New Zealand (supp. no. 1). Transactions and Proceedings of the Royal Society of New Zealand 77 (3): 355-360.

[N02] Negi, H. R. 2002. Abundance and diversity of moss communities of Chopta-Tunganath in the Garhwal Himalaya. Journal of the Bombay Natural History Society 99 (3): 418-433.

Smith, A. J. E., & R. Smith. 2004. The Moss Flora of Britain and Ireland, 2nd ed. Cambridge University Press.

[SF01] Stech, M., & W. Frey. 2001. CpDNA-relationship and classification of the liverworts (Hepaticophytina, Bryophyta). Nova Hedwigia 72 (1-2): 45-58.

[SK02] Streimann, H., & N. Klazenga. 2002. Catalogue of Australian Mosses. Flora of Australia Supplementary Series 17. Australian Biological Resources Study: Canberra.

[W79] Wiley, E. O. 1979. An annotated Linnean hierarchy, with comments on natural taxa and competing systems. Systematic Zoology 28 (3): 308-337.

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