Mnium hornum, from here.

Belongs within: Bryidae.

The Mniaceae are a family of mosses, with representatives known as thread-mosses and thyme-mosses, with their main diversity in the northern temperate zone. Members of Mniaceae can be distinguished from many (but not all) species of the similar Bryaceae by the absence in the former of comal tufts of leaves (Watson & Dallwitz). Members of the genus Mnium are usually tall and robust mosses with large leaves that often have a distinct border (Dixon 1924).

Characters (from Smith & Smith 2004): Dioicous or synoicous. All stems erect or fertile stems erect and sterile stems procumbent or arcuate; central strand present. Leaves pellucid, ovate to linear-lanceolate, acuminate to rounded; margins unbordered or with one- to several-stratose border, entire or dentate with single or double teeth; costa with stereid band in section; cells more or less quadrate to linear, smooth, sometimes in divergent rows. Female inflorescences bud-like, males discoid or not. Sporophytes sometimes more than one per perichaetium; setae long; capsules cernuous to pendulous, ovoid to cylindrical; peristome double; calyptrae cucullate.

    |--Orthomnion (Mitt.) Broth. 1905 SK02 [Orthomnieae W79]
    |    `--O. elimbatum (Nog.) Kop. 1980 [incl. Mnium rotundifolium] SK02
    |--Plagiomnium Kop. 1968 SK02 [Plagiomnieae W79]
    |    |--P. affine M08
    |    |--P. cuspidatum FHH01
    |    |--P. ellipticum SK02
    |    |--P. novae-zealandiae (Colenso) Kop. 1977 (see below for synonymy) SK02
    |    |--P. rhynchophorum J87
    |    |--P. rostratum [=Mnium rostratum; incl. M. longirostrum] SK02
    |    `--P. undulatum SF01
    `--+--Cinclidium D24 [Cinclidieae W79]
       |    `--C. stygium D24
       `--Mnium Hedw. 1801 SK02 [Mnieae W79]
            |--M. affine D24
            |    |--M. a. var. affine D24
            |    |--M. a. var. elatum [incl. M. seligeri] D24
            |    `--M. a. var. rugicum D24
            |--M. blyttii D24
            |--M. cinclidioides [=Bryum cinclidioides] D24
            |--M. cuspidatum [incl. M. silvaticum] D24
            |--M. hornum FHH01
            |--M. integrum D24
            |--M. japonicum N02
            |--M. leptolimbatum D03
            |--M. lycopodioides [=Bryum lycopodioides; incl. M. gracilentum] D24
            |--M. orthorrhynchum D24
            |--M. punctatum D24
            |    |--M. p. var. punctatum D24
            |    `--M. p. var. elatum non M. affine var. elatum D24
            |--M. riparium D24
            |--M. serratum [incl. M. marginatum] D24
            |--M. spinosum [=Bryum spinosum] D24
            |--M. spinulosum D24
            |--M. stellare D24
            |--M. subglobosum [incl. M. pseudopuncatum] D24
            `--M. undulatum D24

Mniaceae incertae sedis:
  Schizymenium Harv. ex Hooker 1840 SK02
    `--S. bryoides Harv. ex Hooker 1840 (see below for synonymy) SK02
  Mielichhoferia Nees & Hornsch. 1831 SK02
    |--M. forsythii Broth. 1916 SK02
    |--M. mielichhoferi N02
    |--M. spegazziniana D03
    |--M. sullivanii Müll.Hal. 1898 SK02
    `--M. tenuiseta M49
  Pohlia Hedw. 1801 SK02
    |--P. acuminata [=Webera acuminata] D24
    |--P. andalusica M08
    |--P. bulbifera (Warnst.) Warnst. 1904 SK02
    |--P. clavaeformis (Hampe) Broth. 1903 [=Bryum clavaeforme, Webera clavaeformis] SK02
    |--P. cruda (Hedw.) Lindb. 1879 SK02 (see below for synonymy)
    |--P. cuspidata Bartram 1951 SK02
    |--P. elongata [=Webera elongata; incl. W. ambigua] D24
    |--P. flexuosa N02
    |--P. longicolla N02
    |--P. lutescens M08
    |--P. melanodon M08
    |--P. mielichhoferia (Müll.Hal.) Broth. 1903 [=Bryum mielichhoferia, Webera mielichhoferia] SK02
    |--P. minor N02
    |--P. novae-seelandiae Dixon 1915 SK02
    |--P. nutans (Hedw.) Lindb. 1879 SK02 (see below for synonymy)
    |    |--P. n. var. nutans D24
    |    |--‘Webera’ n. var. bicolor D24
    |    `--‘Webera’ n. var. longiseta D24
    |--P. polymorpha [=Webera polymorpha] D24
    |    |--P. p. var. polymorpha D24
    |    `--‘Webera’ p. var. brachycarpa [=Pohlia brachycarpa] D24
    |--P. rigescens N02
    |--P. tenuifolia (Jaeger) Broth. 1903 SK02
    |--P. turgens (Broth.) Shaw 1987 [=Mielichhoferia turgens Broth. 1916] SK02
    `--P. wahlenbergii (Weber & Mohr) Andrews 1935 (see below for synonymy) SK02
         |--P. w. var. wahlenbergii D24
         `--‘Webera albicans’ var. glacialis [=Mnium glaciale] D24

Nomina nuda: Leptochlaena microdonta Mitt. ex Muell. 1880 [=Mielichhoferia microdonta Mitt. 1882] SK02
             Mnium alpestre Müll.Hal. 1901 SK02
             Pohlia austrocarnea Broth. ex Ramsay 1984 SK02

Plagiomnium novae-zealandiae (Colenso) Kop. 1977 [incl. Mnium novae-valesiae Müll.Hal. 1876 (n. n.), M. tasmanicum Watts & Whitel. 1902 (nom. inv.)] SK02

Pohlia cruda (Hedw.) Lindb. 1879 SK02 [=Mnium crudum D24, Webera cruda SK02; incl. W. erythrocaulis SK02, Bryum erythrocaule (preoc.) SK02, Pohlia erythrocaulis SK02, W. synoico-cruda D03]

Pohlia nutans (Hedw.) Lindb. 1879 SK02 [=Webera nutans SK02, Bryum nutans SK02; incl. Pohlia austronutans (n. n.) SK02, B. beccarii SK02, P. beccarii SK02, Webera beccarii SK02, Distichium capillaceum var. brevifolium D24, Swartzia montana var. compacta D24, Bryum leptopelmatum SK02, Pohlia leptopelmata SK02, Webera leptopelmata SK02, W. longifolia SK02, Bryum longifolium (preoc.) SK02, Pohlia longifolia SK02, B. montanum SK02, P. montana SK02, Swartzia montana D24, Webera montana SK02]

Pohlia wahlenbergii (Weber & Mohr) Andrews 1935 [=Mniobryum wahlenbergii; incl. Bryum albicans (nom. illeg.), M. albicans (nom. illeg.), Pohlia albicans (nom. illeg.), Webera albicans (nom. illeg.), Mniobryum wahlenbergii var. tasmanicum, Pohlia albicans var. tasmanica, P. cruda var. tasmanica, Webera tasmanica] SK02

Schizymenium bryoides Harv. ex Hooker 1840 [=Mielichhoferia bryoides (Harv. ex Hooker) Wijk & Margad. 1962; incl. M. australis, M. ecklonii] SK02

*Type species of generic name indicated


[D24] Dixon, H. N. 1924. The Student's Handbook of British Mosses 3rd ed. V. V. Sumfield: Eastbourne.

[D03] Dusén, P. 1903. Patagonian and Fuegian mosses. In: Scott, W. B. (ed.) Reports of the Princeton University Expeditions to Patagonia, 1896–1899 vol. 8. Botany pp. 63–126. The University: Princeton (New Jersey).

[FHH01] Frey, W., M. Hofmann & H. H. Hilger. 2001. The gametophyte-sporophyte junction: unequivocal hints for two evolutionary lines of archegoniate land plants. Flora 196: 431–445.

[J87] Judd, W. S. 1987. Floristic study of Morne La Visite and Pic Macaya National Parks, Haiti. Bulletin of the Florida State Museum—Biological Sciences 32 (1): 1–136.

[M08] Marstaller, R. 2008. Moosgesellschaften am Südrand des Kyffhäusergebirges bei Bad Frankenhausen (Kyffhäuserkreis). 130. Beitrag zur Moosvegetation Thüringens. Mauritiana 20 (2): 289–348.

[M49] Martin, W. 1949. Distribution of the mosses indigenous to New Zealand (supplement no. 1). Transactions and Proceedings of the Royal Society of New Zealand 77 (3): 355–360.

[N02] Negi, H. R. 2002. Abundance and diversity of moss communities of Chopta-Tunganath in the Garhwal Himalaya. Journal of the Bombay Natural History Society 99 (3): 418–433.

Smith, A. J. E., & R. Smith. 2004. The Moss Flora of Britain and Ireland, 2nd ed. Cambridge University Press.

[SF01] Stech, M., & W. Frey. 2001. CpDNA-relationship and classification of the liverworts (Hepaticophytina, Bryophyta). Nova Hedwigia 72 (1–2): 45–58.

[SK02] Streimann, H., & N. Klazenga. 2002. Catalogue of Australian Mosses. Flora of Australia Supplementary Series 17. Australian Biological Resources Study: Canberra.

[W79] Wiley, E. O. 1979. An annotated Linnean hierarchy, with comments on natural taxa and competing systems. Systematic Zoology 28 (3): 308–337.

Last updated: 28 December 2017.

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