Trichadenotecnum

Trichadenotecnum alexanderae, copyright Alain Hogue.


Belongs within: Psocinae.

Trichadenotecnum is a cosmopolitan genus of barklice with heavily pigmented fore wings.

See also: Trichadenotecnum: six spots and spiny terminalia.

Characters (from New & Lienhard 2007, Yoshizawa & Lienhard 2015): Fore wing markings including spots in cell C; opposing spots near base of cell r; proximal band fron stigmasac passing through M+Cu fork to distal third of cell cu2; submarginal row of six spots. Male with clunial arms usually present; hypandrium usually with median tongue; apophyses present on posterior margin of hypandrium.

<==Trichadenotecnum Enderlein 1909 YL15 [incl. Cryptopsocus Li 2002 YL15, Trichadenopsocus Roesler 1943 NL07, L02]
    |--+--+--T. bos YL15
    |  |  `--T. tambopatense YL15
    |  `--+--T. gonzalezi YL15
    |     `--T. peruense YL15
    `--+--+--T. corniculum YL15
       |  `--T. germinatum YL15
       |--T. marginatum group YL15
       |    |--+--T. sabahense Yoshizawa & Lienhard 2015 YL15
       |    |  `--T. sharkeyi Yoshizawa & Lienhard 2015 YL15
       |    `--+--T. cynostigmus (Li 2002) [=*Cryptopsocus cynostigmus] YL15
       |       |--T. marginatum New & Thornton 1976 YL15
       |       `--T. tigrinum Yoshizawa & Lienhard 2015 YL15
       `--+--T. alexanderae complex M93
          |    |--T. alexanderae (Sommerman 1948) YL15, L02 [=Trichadenopsocus alexanderae L03]
          |    |--T. castum Betz 1983 YL15, M93
          |    |--T. innuptum Betz 1983 M93
          |    `--T. merum Betz 1983 M93
          |--T. spiniserrulum group YL15
          |    |  i. s.: T. spiniserrulum Datta 1969 L02 [=Trichadenopsocus spiniserrulus L03]
          |    |--T. ianobidens YL15
          |    `--+--T. falx Yoshizawa 2001 YL15, M03
          |       `--T. furcalingum YL15
          `--+--T. iani YL15
             |--+--T. nothoapertum YL15
             |  `--T. yamatomajus Yoshizawa 2001 YL15, M03
             `--+--+--+--T. album YL15
                |  |  `--T. incognitum Roesler 1939 YL15, G74
                |  `--+--T. mixtum YL15
                |     `--+--T. latebrachium YL15
                |        `--T. medium Thornton 1961 YL15, L02 [=Loensia media L03, Thichorntontecnum medium L02]
                `--+--+--T. desolatum (Chapman 1930) YL15, M93 (see below for synonymy)
                   |  `--T. tuitense YL15
                   `--+--+--T. aconcinnum YL15
                      |  `--+--+--T. decui YL15
                      |     |  `--T. obrienorum YL15
                      |     `--+--T. miffy YL15
                      |        `--T. slossonae (Banks 1903) YL15, M93 (see below for synonymy)
                      `--+--+--T. chiapense YL15
                         |  `--T. erwini YL15
                         `--+--+--T. brevicornum YL15
                            |  `--T. quaesitum (Chapman 1930) YL15, M93 [=Psocus quaesitus M93]
                            `--+--T. concinnum YL15
                               `--T. nicaraguense YL15

Trichadenotecnum incertae sedis:
  *T. sexpunctatum (Linnaeus 1758) YL15 [=Hemerobius sexpunctatus YL15, Psocus sexpunctatus L02]
  T. adika Endang, Thornton & New 2002 NL07
  T. aduncatum (Li 2002) [=Trichadenopsocus aduncatus] L03
  T. alinguum Endang, Thornton & New 2002 NL07
  T. alobum Endang & New 2005 NL07
  T. alternatum (Li 2002) [=Trichadenopsocus alternatus] L03
  T. ampullaceum (Li 2002) [=Trichadenopsocus ampullaceus] L03
  T. angolensis Badonnel 1955 B55
  T. arciforme Thornton 1961 NL07
  T. baishanzuicum Li 1995 L02
  T. bicolor (Li 2001) [=Trichadenopsocus bicolor] L03
  T. bidens Thornton 1961 NL07 [=Loensia bidens L02]
  T. bidentatum Thornton 1984 NL07 [=Trichadenopsocus bidentatus L03]
  T. bitenatum (Li 1995) L03, L02 [=Trichadenopsocus biternatus (l. c.) L03]
  T. bromoense Endang, Thornton & New 2002 NL07
  T. bucciniforme Li 1993 L02 [=Trichadenopsocus bucciniformis L03]
  T. calycoideum Li 2002 L02
  T. cheahae Endang, Thornton & New 2002 NL07
  T. chinense Li 1997 L02
  T. cinnamonum Endang & New 2005 NL07
  T. circulare (Hagen 1859) NL07
  T. circularoides Badonnel 1955 NL07
  T. cornutum Endang & New 2005 NL07
  T. dactylinum (Li 2002) [=Trichadenopsocus dactylinus] L03
  T. digitatum (Li 2002) [=Trichadenopsocus digitatus] L03
  T. diplodurum Li 2002 L02
  T. emeishanense Li 2002 [=T. emeishanaense, T. emeishananse, T. emeishanse] L03
  T. felix Thornton 1961 L02 [=Trichadenopsocus felix L03]
  T. galihi Endang, Thornton & New 2002 NL07
  T. gallicum Lienhard 1986 L98
  T. germanicum Roesler 1939 G74
  T. godavarense New 1971 NL07
  T. gutianum Li 1995 L02
  T. hammani Endang, Thornton & New 2002 NL07
  T. himalayense Li & Yang 1987 L03, L02 [=Trichadenopsocus himalayaensis (l. c.) L03]
  T. imperatorium Li 1989 L02
  T. imrum New & Thornton 1976 NL07
  T. isocaulum Li 2002 L02
  T. jaculatorum (Li 2002) [=Trichadenopsocus jaculatorus] L03
  T. jambiense Endang & New 2005 NL07
  T. jinxiuense (Li 2002) [=Trichadenopsocus jinxiuensis] L03
  T. kalibiruense Endang & New 2005 NL07
  T. kerinciense Endang & New 2005 NL07
  T. krucilense Endang, Thornton & New 2002 NL07
  T. kunmingicum Li 2002 L02
  T. laticornutum Endang, Thornton & New 2002 NL07
  T. mamillatum Li 2002 L02
  T. maroccanum Baz 1989 L98
  T. mclachlani New 1973 NL07 [=Trichadenopsocus mclachlani L03]
  T. minisexmaculatum Li & Yang 1987 L03, L02 (see below for synonymy)
  T. minutum Enderlein 1926 NL07
  T. muaraense Endang & New 2005 NL07
  T. multangulare (Li 2002) [=Trichadenopsocus multangularis] L03
  T. multicuspidatum (Li 2002) [=Trichadenopsocus multicuspidatus] L03
  T. obsitum (Enderlein 1908) L02
  T. obsubulatum Li 2002 L02
  T. opiparipardale (Li 1995) L03, L02 [=Trichadenopsocus opiparipardalis L03]
  T. paradika Endang & New 2005 NL07
  T. pardoides Badonnel 1955 B55
  T. pardus Badonnel 1955 NL07 (see below for synonymy)
  T. paululum (Li 2002) [=Trichadenopsocus paululus] L03
  T. percussum Li 2002 L02
  T. pergracilum Li 2002 L02
  T. periphericum (Li 2001) [=Trichadenopsocus periphericus] L03
  T. pokhariense New 1983 L02
  T. proctum Endang & New 2005 NL07
  T. qingshuicum Li 2002 L02
  T. quadrispinosum Endang, Thornton & New 2002 NL07
  T. quadruplex (Li 2002) [=Trichadenopsocus quadruplex] L03
  T. rachimi Endang & Thornton 1992 NL07
  T. resupinum (Li 1997) L03, L02 [=Trichadenopsocus resupinus L03]
  T. rhomboides Li 2002 L02
  T. santosai Endang & Thornton 1992 NL07
  T. scoparium (Li 1995) L03, L02 [=Trichadenopsocus scoparius L03]
  T. sibolangitense Endang & New 2005 NL07
  T. soekarmanni Endang, Thornton & New 2002 NL07
  T. soenarti Endang, Thornton & New 2002 NL07
  T. spuristipiatum Li 1997 L02
  T. stipiatum Li 1992 L02
  T. stipulatum (Li 2002) [=Trichadenopsocus stipulatus] L03
  T. subrotundum (Li 2002) [=Trichadenopsocus subrotundus] L03
  T. subscalare (Li 2002) [=Trichadenopsocus subscalaris] L03
  T. sufflatum Li 1993 L02 [=Trichadenopsocus sufflatus L03]
  T. sumatrense Endang & New 2005 NL07
  T. tenuispinum Li 1995 L02
  T. thallodialum Li 2002 L02
  T. trichotomum (Li 2002) [=Trichadenopsocus trichotomus] L03
  T. trigonophyllum Li 1993 L02
  T. uncorne (Li 1995) L03, L02 [=Trichadenopsocus uncornis L03]
  T. uniforme (Li 2002) [=Trichadenopsocus uniformis] L03
  T. univittatum Li 2002 L02
  T. vaughani Endang, Thornton & New 2002 NL07
  T. waykambasense Endang & New 2005 NL07
  T. waykananense Endang & New 2005 NL07
  T. wuxiacum Li 1997 L02
  T. xizangicum Li 2002 L02
  T. majus group NL07
    |--T. apertum Thornton 1961 NL07, L02 [=Trichadenopsocus apertus L03]
    |--T. gombakense New & Lee 1992 NL07
    |--T. majus (Kolbe 1880) M93 (see below for synonymy)
    `--T. malayense New 1975 NL07 [=Trichadenopsocus malayaense (l. c.) L03]

Trichadenotecnum desolatum (Chapman 1930) YL15, M93 [=Psocus desolatus NL07, Trichadenotecnum (*Trichadenopsocus) desolatus M93, NL07]

Trichadenotecnum majus (Kolbe 1880) M93 [=Psocus sexpunctatus var. major L02, T. maijum (l. c.) L03, T. majum (l. c.) L03; incl. P. major var. ocellata Enderlein 1901 L02, P. major var. septentrionalis Enderlein 1910 L02]

Trichadenotecnum minisexmaculatum Li & Yang 1987 L03, L02 [=T. minsexmaculatum (l. c.) L03, T. minsexmaculatus (l. c.) L03]

Trichadenotecnum pardus Badonnel 1955 NL07 [=Trichadenopsocus pardus L03; incl. Trichadenotecnum pardidum Thornton 1961 L03, Trichadenopsocus pardidus L03]

Trichadenotecnum slossonae (Banks 1903) YL15, M93 [=Psocus slossonae M93; incl. T. unum Sommerman 1948 M93]

*Type species of generic name indicated

REFERENCES

[B55] Badonnel, A. 1955. Psocoptères de l'Angola. Diamang Publicações Culturais 26: 11–267.

[G74] Günther, K. K. 1974. Staubläuse, Psocoptera. Die Tierwelt Deutschlands 61: 1–315.

[L02] Li F. 2002. Psocoptera of China vol. 2. Science Press: Beijing.

[L98] Lienhard, C. 1998. Faune de France. France et Régions Limitrophes. 83. Psocoptères Euro-Méditerranéens. Fédération Française des Sociétés de Sciences Naturelles: Paris.

[L03] Lienhard, C. 2003. Nomenclatural amendments concerning Chinese Psocoptera (Insecta), with remarks on species richness. Revue Suisse de Zoologie 110 (4): 695–721.

[M93] Mockford, E. L. 1993. North American Psocoptera (Insecta). Sandhill Crane Press, Inc.

[M03] Mockford, E. L. 2003. New species and records of Psocoptera from the Kuril Islands. Mitt. Mus. Nat.kd. Berl., Dtsch. Entomol. Z. 50 (2): 191–230.

[NL07] New, T. R., & C. Lienhard. 2007. The Psocoptera of Tropical South-east Asia. Brill: Leiden.

[YL15] Yoshizawa, K., & C. Lienhard. 2015. Synonymy of Cryptopsocus Li with Trichadenotecnum Enderlein (Insecta: Psocodea: 'Psocoptera': Psocidae) and description of three new species. Zootaxa 3957 (4): 480–488.

Amphigerontiinae

Prospective Neoblaste sp., from here.


Belongs within: Psocetae.
Contains: Amphigerontia, Blaste.

The Amphigerontiinae are a group of barklice characterised by amorphous development of the male abdominal apex and a reduced phallosome (New & Lienhard 2007).

Characters (from New & Lienhard 2007): Fore wing veins R4+5 and M not connected or fused; vein M comprising distal border of discoidal cell usually convex; pterostigma relatively deep. Male hypandrial sclerites encompassing abdominal sternites 8 and 9 so apex of male abdomen appears 'encapsulated'; parameres open distally (no phallosome frame); apophysis of male paraproct broadly rounded. Female with apex of dorsal gonapophysis valve a slender tapered process.

<==Amphigerontiinae
    |--Stylatopsocus Li 2002 [Stylatopsocini] L02
    |    `--*S. biuncialis Li 2002 L02
    |--Amphigerontiini L02
    |    |--Amphigerontia NL07
    |    `--Metagerontia Li 2002 L02
    |         `--*M. tribulosa Li 2002 L02
    `--Blastini L02
         |--Blaste NL07
         |--Chilopsocus Li 2002 L02 [=Chilosopsocus L03]
         |    `--*C. macrochilus Li 2002 L03
         |--Disopsocus Li 2002 L02
         |    `--*D. megacheilus Li 2002 L02
         |--Epiblaste Li 1993 L02
         |    |--*E. glandacea Li 1993 L02
         |    `--E. huananiensis Li 2002 [=E. huananinsis] L03
         |--Pentablaste Li 2002 L02
         |    |--*P. obconica Li 2002 L02
         |    |--P. auctachila Li 2002 L02
         |    |--P. clavata Li 2002 L02
         |    |--P. flavae (Li 1995) [=Neoblaste flavae] L02
         |    |--P. flavida (Li 1989) L03, L02 (see below for synonymy)
         |    |--P. jinxiuica Li 2002 L02
         |    |--P. lanceolata Li 2002 L02
         |    |--P. longicaudata Li 2002 L02
         |    |--P. lushanensis Li 2002 L02
         |    |--P. minuscula Li 2002 L02
         |    |--P. pentasticha Li 2002 L03 (see below for synonymy)
         |    |--P. pini (Li 1990) [=Blastopsocidus pini] L02
         |    |--P. profunda (Li 1995) [=Neoblaste profunda] L02
         |    |--P. quinquedentata (Li & Yang 1988) [=Blostopsocidus quinquedentata] L02
         |    |--P. schizopetala (Li 1997) [=Neoblaste schizopetala] L02
         |    `--P. tetraedrica Li 2002 [=P. tetraedirca] L03
         `--Neoblaste Thornton 1960 L02
              |--*N. papillosa Thornton 1960 L02
              |--N. alticola Thornton 1984 NL07
              |--N. ancistroides Li 2002 L02
              |--N. brunnea Endang & Thornton 1992 NL07
              |--N. complexa Endang, Thornton & New 2002 NL07
              |--N. cubitalis (Enderlein 1919) NL07
              |--N. fujianensis (Li & Yang 1987) [=Blaste fujianensis] L02
              |--N. guangdongensis New 1991 L02
              |--N. javensis Endang, Thornton & New 2002 NL07
              |--N. octogona Li 2002 L02
              |--N. ovalis Li 2002 L02
              |--N. parasetosa Endang, Thornton & New 2002 NL07
              |--N. partibilis Li 2002 L02
              |--N. pinicola Li 1990 L02
              |--N. rectangula Li 2002 L02
              |--N. setosa Thornton 1960 L02
              |--N. soehardjani Endang, Thornton & New 2002 NL07
              |--N. timorensis Endang, Thornton & New 2002 NL07
              |--N. tricornis Li 2002 L02
              `--N. umbonalis Li 2002 L02

Amphigerontiinae incertae sedis:
  Chaetopsocidus Badonnel 1986 B86
    `--*C. sturmi Badonnel 1986 B86
  Blastopsocus Roesler 1843 M93
    |--*B. variabilis (Aaron 1883) M93 (see below for synonymy)
    |--B. angustus Smithers & Thornton 1981 NL07
    |--B. lithinus (Chapman 1930) [=Psocus lithinus, Blaste (Blastopsocus) lithinus] M93
    |--B. mockfordi Badonnel 1986 B86
    |--B. semistriatus (Walsh 1862) [=Psocus semistriatus] M93
    `--B. uncinata B86
  Euclismioides Smithers & Thornton 1981 NL07, L02
    `--*E. inocellata Smithers & Thornton 1981 NL07
  Neopsocopsis Badonnel 1935 G74
    `--N. hirticornis (Reuter 1893) (see below for synonymy) L98
  Anomaloblaste NL07
    |--*A. spinivalva Endang, Thornton & New 2002 NL07
    `--A. treubia Endang, Thornton & New 2002 NL07
  Indoblaste NL07
    |--*I. lienhardi Endang & Thornton 1992 NL07
    |--I. discoloba Endang, Thornton & New 2002 NL07
    |--I. lawuensis Endang, Thornton & New 2002 NL07
    `--I. sastrawani Endang & Thornton 1992 NL07
  Javablaste NL07
    |--*J. aidab Endang, Thornton & New 2002 NL07
    `--J. merapiensis Endang, Thornton & New 2002 NL07
  *Lasiopsocus michaelseni [=Blaste (*Lasiopsocus) michaelseni] NL07

*Blastopsocus variabilis (Aaron 1883) M93 [=Psocus variabilis NL07, Blaste (Blastopsocus) variabilis M93, Euclismia variabilis NL07; incl. P. medialis Banks 1907 M93]

Neopsocopsis hirticornis (Reuter 1893) [=Psocus hirticornis; incl. P. bastmannianus Enderlein 1918, Neopsocus pyrenaicus Badonnel 1935, *Neopsocopsis pyrenaicus] L98

Pentablaste flavida (Li 1989) L03, L02 [=Blastopsocidus flavidae L02, P. flavidae (l. c.) L03, P. flavivdae (l. c.) L03]

Pentablaste pentasticha Li 2002 L03 [=Neoblaste quinquedentata Li 1990 non Blostopsocidus quinquedentata Li & Yang 1988 L02, P. pentaticha L03]

*Type species of generic name indicated

REFERENCES

[B86] Badonnel, A. 1986. Psocoptères de Colombie (Insecta, Psocoptera). Spixiana 9 (2): 179–223.

[G74] Günther, K. K. 1974. Staubläuse, Psocoptera. Die Tierwelt Deutschlands 61: 1–315.

[L02] Li F. 2002. Psocoptera of China vol. 2. Science Press: Beijing.

[L98] Lienhard, C. 1998. Faune de France. France et Régions Limitrophes. 83. Psocoptères Euro-Méditerranéens. Fédération Française des Sociétés de Sciences Naturelles: Paris.

[L03] Lienhard, C. 2003. Nomenclatural amendments concerning Chinese Psocoptera (Insecta), with remarks on species richness. Revue Suisse de Zoologie 110 (4): 695–721.

[M93] Mockford, E. L. 1993. North American Psocoptera (Insecta). Sandhill Crane Press, Inc.

[NL07] New, T. R., & C. Lienhard. 2007. The Psocoptera of Tropical South-east Asia. Brill: Leiden.

Loensia

Loensia moesta, copyright Robert Webster.


Belongs within: Psocinae.

Loensia is a genus of barklice in which the fore wings are extensively marked with scattered blotches.

Characters (from Mockford 1993): Adults of both sexes macropterous. Five distal inner labral sensilla present. Fore wing marked with scattered spotting and blotching over most of wing surface with no particular pattern. Hypandrium with or without basal sclerites; asymmetrical medial strap present based on right side, usually trending leftward, bent upward and forward at apex, sometimes not bending leftward but a lobe or process on left side sometimes as long as or longer than strap. Phallosome with short anterior apodeme; distally with a single process, usually symmetrical. Male epiproct chair-shaped, extending anteriorly over clunium. Subgenital plate with distal process short and wide, about as long as, or slightly shorter than, its basal width; pigmented arms slender, directed forward, abruptly widening to form broad, quadrate lateral areas. Ovipositor valvulae with apical process of v2 long and slender; v3 relatively slender, transverse, with moderate to long median lobe.

<==Loensia Enderlein 1924 L02 [=Lensia (l. c.) L03]
    |--*L. fasciata (Fabricius 1787) M93 (see below for synonymy)
    |--L. bannaensis Li 2002 L02
    |--L. beijingensis Li 2002 L02
    |--L. bifurcata (Li 1992) [=Trichadenotecnum bifurcatum] L02
    |--L. binalis Li 2002 L02
    |--L. conspersa (Banks 1903) [=Psocus conspersus, Psocidus conspersus] M93
    |--L. dolabrata (Li & Yang 1987) [=Trichadenotecnum dolabratum] L02
    |--L. excrescens Li 1997 L02
    |--L. falcata Li 1997 L02
    |--L. folivalva Li 2002 L02
    |--L. fuscimacula Enderlein 1926 NL07
    |--L. glabridorsum Enderlein 1926 NL07
    |--L. guangdongica (Li 1993) L02 [=Trichadenotecnum guangdongicum L03, T. guandongicum (l. c.) L03]
    |--L. hengshanica Li 2002 L02
    |--L. infundibularis (Li 1989) [=Trichadenotecnum infundibularis] L02
    |--L. maculosa (Banks 1908) [=Myopsocus maculosus, Psocus maculosus, Trichadenotecnum (Loensia) maculosum] M93
    |--L. moesta (Hagen 1861) [=Psocus moestus, Amphigerontia moestus, Trichadenotecnum moestum] M93
    |--L. octogona Li 2002 L02
    |--L. pearmani Kimmins 1941 (see below for synonymy) L98
    |--L. pycnacantha Li 2002 L02
    |--L. rectangula (Li 1992) [=Trichadenotecnum rectangulum] L02
    |--L. scrobicularis (Li 1992) [=Trichadenotecnum scrobicularum] L02
    |--L. spicata Li 1995 L02
    |--L. spissa Li 1995 L02
    |--L. stigmatoidea (Li 1993) L03, L02 [=Trichadenotecnum stigmatoideum L02, L. stigmatoides (l. c.) L03]
    |--L. taeniana (Li & Yang 1987) [=Trichadenotecnum taenianum] L02
    |--L. teretiuscula (Li & Yang 1987) [=Trichadenotecnum teretiusculum] L02
    `--L. variegata (Latreille 1799) (see below for synonymy) L98

*Loensia fasciata (Fabricius 1787) M93 [=Hemerobius fasciatus M93, Amphigerontia fasciata M93, Psocus fasciatus L02, Trichadenotecnum (Loensia) fasciatum M93; incl. P. pilicornis Latreille 1799 M93, Loensia fasciata ab. psocoides Enderlein 1910 L98]

Loensia pearmani Kimmins 1941 [incl. Trichadenotecnum macilentum Roesler 1943, Psocus picicornis Stephens 1836 non Hemerobius picicornis Fabricius 1793] L98

Loensia variegata (Latreille 1799) [=Psocus variegatus; incl. P. atomarius Stephens 1836, P. flavonimbatus Rostock 1879] L98

*Type species of generic name indicated

REFERENCES

[L02] Li F. 2002. Psocoptera of China vol. 2. Science Press: Beijing.

[L98] Lienhard, C. 1998. Faune de France. France et Régions Limitrophes. 83. Psocoptères Euro-Méditerranéens. Fédération Française des Sociétés de Sciences Naturelles: Paris.

[L03] Lienhard, C. 2003. Nomenclatural amendments concerning Chinese Psocoptera (Insecta), with remarks on species richness. Revue Suisse de Zoologie 110 (4): 695–721.

[M93] Mockford, E. L. 1993. North American Psocoptera (Insecta). Sandhill Crane Press, Inc.

[NL07] New, T. R., & C. Lienhard. 2007. The Psocoptera of Tropical South-east Asia. Brill: Leiden.

Blaste

Blaste quieta, copyright Robert Webster.


Belongs within: Amphigerontiinae.

Blaste is a cosmopolitan genus of barklice, the definition of which remains debated.

Characters (from Mockford 1993): Adults macropterous or brachypterous, rarely micropterous. Fore wing with veins Rs and M joined at a point, or by a short fusion, or by a short crossvein. Hypandrium distally symmetrical with a pointed process medially (occasionally absent), flanked by two processes of variable shape and length. Male parameres fused at base. Male epiproct with straight anterior margin, not overlapping clunial margin. Subgenital plate with egg guide relatively short, usually somewhat arrow-head shaped; pigmented arms directed anterolaterally, only slightly expanded at distal ends, no anterior median projection from basal junction of arms.

<==Blaste Kolbe 1883 SN08
    |--B. (Blaste) NL07
    |    |--B. (B.) quieta (Hagen 1861) M93 (see below for synonymy)
    |    `--B. (B.) angolensis Badonnel 1955 B55
    |--B. (Blastopsis Badonnel 1955) B55
    |    `--B. (*B.) triangularum Badonnel 1955 B55
    |--B. (Blastopsocidus Badonnel 1955) B55
    |    `--B. (*B.) maculatus Badonnel 1955 B55
    |--B. (Euclismia Enderlein 1925) NL07, G74
    |    |--B. (*E.) quadrimaculata (Latreille 1794) NL07, G74 (see below for synonymy)
    |    |--B. (E.) arabica New 1979 L98
    |    |--B. (B.) balli (Badonnel 1945) [=Euclismia balli] L98
    |    |--B. (E.) conspurcata (Rambur 1842) [=Psocus conspurcatus; incl. P. hilaris Navás 1907] L98
    |    |--B. (E.) cyprica Lienhard 1995 L98
    |    |--B. (E.) diyma Lienhard 1986 L98
    |    |--B. (E.) dundoensis Badonnel 1955 B55
    |    |--B. (E.) lepesmei B55
    |    |--B. (E.) membranosa Lienhard & Halperin 1988 L98
    |    |--B. (E.) osella Lienhard 1987 L98
    |    `--B. (E.) sarda Lienhard 1986 L98
    `--B. (Euclismiopsis Badonnel 1955) B55
         |--B. (*E.) machadoi Badonnel 1955 B55
         |--B. (E.) medleri New 1975 [incl. Neopsocopsis aegyptiacus Meinander 1981, B. muiri Smithers 1979] L98
         |--B. (E.) pauliani B55
         `--B. (E.) vilhenai Badonnel 1955 B55

Blaste incertae sedis:
  B. anuaudi (Badonnel 1955) L02
  B. binotata (Enderlein 1926) NL07
  B. bistriata Schmidt & Thornton 1993 SN08
  B. capricornuta Mockford 1974 M74
  B. cockerelli (Banks 1904) [=Psocus cockerelli] M93
  B. cyclota Li 2002 L02
  B. euryphylla Li & Yang 1987 L02
  B. falcifer Smithers 1979 SN08
  B. fasciata Mockford 1974 M74
  B. garciorum Mockford 1984 M93
  B. harpophylla (Li 1993) [=Neoblaste harpophylla] L02
  B. immobilis Mockford 1974 M74
  B. lignicola (Enderlein 1906) [=Psocus lignicola, B. (Euclismia) lignicola] SN08
  B. longicauda Mockford 1974 M74
  B. longipennis (Banks 1918) [=Psocus longipennis] M93
  B. longivalva New 1975 NL07
  B. nana New 1975 NL07
  B. octofaria Li 2002 L02
  B. opposita (Banks 1907) [=Psocus oppositus; incl. P. interruptus Banks 1920] M93
  B. oregona (Banks 1900) [=Psocus oregonus; incl. P. californicus Banks 1905] M93
  B. osceola Mockford 1984 M93
  B. panops Smithers 1979 SN08
  B. persimilis (Banks 1908) [=Psocus persimilis] M93
  B. phrynae Li 2002 L02
  B. posticata (Banks 1905) [=Psocus posticatus, Psocidus posticatus] M93
  B. pusilla M93
  B. serrata New 1973 NL07
  B. smilivirgata Li 1989 L02
  B. subapterous (Chapman 1930) [=Psocus subapterous] M93
  B. subquieta (Chapman 1930) [=Psocus subquietus] M93
  B. taylori New 1974 SN08
  B. tillyardi Smithers 1969 SN08
  B. verticalis Li 2002 L02
  B. villei Badonnel 1967 L02
  B. yigongensis Li & Yang 1987 L02

Blaste (Blaste) quieta (Hagen 1861) M93 [=Psocus quietus M93; incl. P. bifasciatus Walsh 1863 non Latreille 1799 M93, P. inornatus Aaron 1883 M93, *Blaste juvenilis Kolbe 1883 SN08, M93, P. stigmosalis Banks 1914 M93]

Blaste (*Euclismia) quadrimaculata (Latreille 1794) NL07, G74 [=Psocus quadrimaculatus NL07; incl. P. maculipennis Stephens 1836 G74, P. subnebulosus Stephens 1836 G74]

*Type species of generic name indicated

REFERENCES

[B55] Badonnel, A. 1955. Psocoptères de l'Angola. Diamang Publicações Culturais 26: 11–267.

[G74] Günther, K. K. 1974. Staubläuse, Psocoptera. Die Tierwelt Deutschlands 61: 1–315.

[L02] Li F. 2002. Psocoptera of China vol. 2. Science Press: Beijing.

[L98] Lienhard, C. 1998. Faune de France. France et Régions Limitrophes. 83. Psocoptères Euro-Méditerranéens. Fédération Française des Sociétés de Sciences Naturelles: Paris.

[M74] Mockford, E. L. 1974. Records and descriptions of Cuban Psocoptera. Entomologica Americana 48 (2): 103–215.

[M93] Mockford, E. L. 1993. North American Psocoptera (Insecta). Sandhill Crane Press, Inc.

[NL07] New, T. R., & C. Lienhard. 2007. The Psocoptera of Tropical South-east Asia. Brill: Leiden.

[SN08] Schmidt, E. R., & T. R. New. 2008. The Psocoptera (Insecta) of Tasmania, Australia. Memoirs of Museum Victoria 65: 71–152.

Amphigerontia

Amphigerontia contaminata, copyright Mick Talbot.


Belongs within: Amphigerontiinae.

Amphigerontia is a genus of barklice found in Europe and the Americas with further species of uncertain affinities assigned to the genus from other parts of the world (New & Lienhard 2007).

Characters (from New & Lienhard 2007): Ocelli present. Fore wing veins Rs and M connected by a crossvein; discoidal cell wing narrowed posteriorly, externally concave. Male hypandrium with hind margin in three lobes.

<==Amphigerontia Kolbe 1880 L02
    |--*A. bifasciata (Latreille 1799) M93 (see below for synonymy)
    |--A. anchorae Li 1989 [=A. anchonae (l. c.), A. ancorne] L03
    |--A. contaminata (Stephens 1836) [=Psocus contaminatus; incl. P. megastigmus Stephens 1836] M93
    |--A. diffusa Navas 1933 L02
    |--A. feai Ribaga 1908 NL07
    |--A. ficivorella Okamoto 1908 L02
    |--A. guiyangica Li 1990 L02
    |--A. incerta Ribaga 1908 NL07
    |--A. infernicola (Chapman 1930) [=Psocus infernicolus] M93
    |--A. intermedia (Tetens 1891) [=Psocus intermedius] G74
    |--A. jezoensis Okamoto 1907 M03
    |--A. lata Enderlein 1926 NL07
    |--A. lhasaensis Li & Yang 1987 [=A. lhasana, A. lhsasna] L03
    |--A. montivaga (Chapman 1930) [=Psocus montivagus] M93
    |--A. namiana Navás 1920 NL07
    |--A. petiolata (Banks 1918) [=Psocus petiolatus] M93
    |--A. shanxiensis Li 2002 L02
    |--A. sicyoides Li 2002 L02
    |--A. ukingana Enderlein 1902 E07
    |--A. unacrodonta Li 2002 L02
    `--A. voeltzkowi B55

*Amphigerontia bifasciata (Latreille 1799) M93 [=Psocus bifasciatus NL07; incl. P. additus Banks 1918 M93, P. confraternus Banks 1905 M93, Amphigerontia confraterna M93, P. moderatus Banks 1907 M93, A. pearmani Roesler 1943 G74, P. subfasciatus Zetterstedt 1840 M93, A. subnebulosa Kolbe 1880 M93]

*Type species of generic name indicated

REFERENCES

[B55] Badonnel, A. 1955. Psocoptères de l'Angola. Diamang Publicações Culturais 26: 11–267.

[E07] Enderlein, G. 1907. Wissenschaftliche Ergebnisse der Schwedischen Zoologischen Expedition nach dem Kilimandjaro, dem Meru und den Umgebenden Massaisteppen Deutsch-Ostafrikas 1905–1906 vol. 15. Corrodentia pt 2. Copeognatha. Almqvist & Wiksells Boktryckeri-A. B.: Uppsala.

[G74] Günther, K. K. 1974. Staubläuse, Psocoptera. Die Tierwelt Deutschlands 61: 1–315.

[L02] Li F. 2002. Psocoptera of China vol. 2. Science Press: Beijing.

[L03] Lienhard, C. 2003. Nomenclatural amendments concerning Chinese Psocoptera (Insecta), with remarks on species richness. Revue Suisse de Zoologie 110 (4): 695–721.

[M93] Mockford, E. L. 1993. North American Psocoptera (Insecta). Sandhill Crane Press, Inc.

[M03] Mockford, E. L. 2003. New species and records of Psocoptera from the Kuril Islands. Mitt. Mus. Nat.kd. Berl., Dtsch. Entomol. Z. 50 (2): 191–230.

[NL07] New, T. R., & C. Lienhard. 2007. The Psocoptera of Tropical South-east Asia. Brill: Leiden.

Hemineura

Elipsocus hyalinus, copyright Ken Schneider.


Belongs within: Elipsocidae.

Hemineura is a genus of barklice found in the western Palaearctic, suggested by phylogenetic analysis to be paraphyletic to the more widespread genus Elipsocus (Schmidt & New 2004).

Characters (from Schmidt & New 2004, for Elipsocus + Hemineura): Antenna with thirteen segments, two placoid sensilla at apex of flagellomere 10, three at base of flagellomere 1; lacinia truncate; setae on fore wing posterior margin emerging from centre of costa; marginal wing scales triangular, not equilateral, with distal apices sharply pointed; hind wing vein Cu1 meeting posterior margin at acute angle; tarsi three-segmented; abdomen ovoid, abdominal vesicles absent; sides of phallosome frame parallel; dorsal gonapophysis valve spine reduced on mesial margin of lobe, valve apically broadening from base, ventral valve normal; subgenital plate anterior diverging arms of uniform width or apically wide.

<==Hemineura Tetens 1891 [incl. Actenotarsus Enderlein 1907] SN04
    |  i. s.: H. blascoi Baz 1994 L98
    |         H. clunialis Lienhard 1995 L98
    |         H. hispanica (Enderlein 1907) [=Actenotarsus hispanicus; incl. H. francescae Badonnel 1989] L98
    |         H. trudiae Lienhard & Halperin 1988 L98
    |         H. turanica Vishnyakova in Vishnyakova & Kaplin 1980 L98
    |         H. wittmeri Badonnel 1981 L98
    |--H. sclerophallina Lienhard 1986 SN04, L98
    `--+--+--*H. dispar Tetens 1891 G74, SN04 [incl. H. fusca Reuter 1904 L98]
       |  `--H. bigoti Badonnel 1970 SN04, L98 [=Elipsocus bigoti L98]
       `--Elipsocus Hagen 1866 SN04 [incl. Cabarer Navás 1908 M93; Elipsocini]
            |--E. pumilis (Hagen 1861) M93 (see below for synonymy)
            |--E. abdominalis Reuter 1904 SN04 (see below for synonymy)
            |--E. annulatus Roesler 1954 G74
            |--E. azoricus Meinander 1975 L98
            |--E. brincki Badonnel 1963 L98
            |--E. coloripennis Lienhard 1996 L98
            |--E. fasciatus (Navás 1908) L98 [=*Cabarer fasciatus NL07]
            |--E. guentheri Mockford 1980 M93
            |--E. hyalinus (Stephens 1836) M93 (see below for synonymy)
            |--E. labralis Lienhard 1996 L98
            |--E. lanceloticus Baz 1991 L98
            |--E. marplatensis Williner 1943 SN04
            |--E. moebiusi Tetens 1891 (see below for synonymy) L98
            |--E. nuptialis Roesler 1954 G74
            |--E. obscurus Mockford 1980 M93
            |--E. oligotrichus Thornton 1959 NL07
            |--E. pallidus Jentsch 1938 G74
            |--E. pusillus Lienhard 1996 L98
            |--E. rubrostigma Navás 1934 SN04
            `--E. viridimicans Enderlein 1900 SN04

Elipsocus abdominalis Reuter 1904 SN04 [=E. hyalinus var. abdominalis M93; incl. E. mclachlani Kimmins 1941 M93, E. occidentalis Banks 1907 M93]

Elipsocus hyalinus (Stephens 1836) M93 [=Psocus hyalinus M93; incl. E. abietis Kolbe 1880 M93, P. bipunctatus Stephens 1836 (preoc.) M93, E. bipunctatus G74, E. abietis ab. tharandtensis Enderlein 1901 G74]

Elipsocus moebiusi Tetens 1891 [incl. E. balmesi Navás 1910, E. brevistylus Reuter 1893, E. pallidus Jentsch 1938] L98

*Elipsocus pumilis (Hagen 1861) M93 [=Psocus pumilis M93; incl. P. quadrimaculatus Westwood 1840 nec Stephens 1836 nec Latreille 1794 M93, Elipsocus quadrimaculatus M93, *E. westwoodii McLachlan 1867 L98]

*Type species of generic name indicated

REFERENCES

[G74] Günther, K. K. 1974. Staubläuse, Psocoptera. Die Tierwelt Deutschlands 61: 1–315.

[L98] Lienhard, C. 1998. Faune de France. France et Régions Limitrophes. 83. Psocoptères Euro-Méditerranéens. Fédération Française des Sociétés de Sciences Naturelles: Paris.

[M93] Mockford, E. L. 1993. North American Psocoptera (Insecta). Sandhill Crane Press, Inc.

[NL07] New, T. R., & C. Lienhard. 2007. The Psocoptera of Tropical South-east Asia. Brill: Leiden.

[SN04] Schmidt, E. R., & T. R. New. 2004. A systematic and phylogenetic revision of the family Elipsocidae (Insecta: Psocoptera), with the erection of two new families: Lesneiidae and Sabulopsocidae. Invertebrate Systematics 18: 157–213.

Ectopsocopsis

Ectopsocopsis cryptomeriae, from Takipororin.


Belongs within: Lachesilloidea.

Ectopsocopsis is a genus of ectopsocid barklice found in Eurasia, Africa and North America. Males have a complex clunial ornamentation, often with a row of teeth and/or extensive asymmetrical sclerotisation; females have the gonapophyses represented only by external valves and usually have a median posterior lobe on the subgenital plate (New & Lienhard 2007).

<==Ectopsocopsis Badonnel 1955 [=Ectopsocoides (l. c.)] L98
    |--*E. balli (Badonnel 1949) B55 [=Ectopsocus balli NL07]
    |--E. annulata Badonnel 1955 M93, B55
    |--E. anurus Badonnel 1955 B55
    |--E. badonneli [=Ectopsocus badonneli] B55
    |--E. beijingensis Li 2002 L02
    |--E. biporosus Li 1992 L02
    |--E. biunciatus Li 2002 L02
    |--E. brevimanubrius Li 2002 L02
    |--E. bulbiapicius Li 2002 L02
    |--E. clavellatus Li 2002 L02
    |--E. cognata Thornton & Wong 1968 NL07
    |--E. corbiformis Li 1993 L02
    |--E. cryptomeriae (Enderlein 1907) (see below for synonymy) L98
    |--E. curtinervis Li 2002 L02
    |--E. daqingshanensis Li 2002 L02
    |--E. decorata (Thornton & Wong 1968) NL07
    |--E. flavipedia Li 2002 [=E. flavipedius, E. flavpedius] L03
    |--E. gannanensis Li 2002 L02
    |--E. guangdongensis Li 1991 L02
    |--E. jiangxiensis Li 2002 L02
    |--E. luteolicapitus Li 2002 L02
    |--E. mozambicus (Badonnel 1931) B55, B49 [=Ectopsocus mozambicus B55]
    |--E. phaeostictous Li 2002 L02
    |--E. pinisugus Li 2002 L02
    |--E. pumilis (Banks 1920) L02 (see below for synonymy)
    |--E. sarmentiformis Li 2002 L02
    |--E. septentrionalis Li 2002 [=E. septentrionalus] L03
    |--E. spathulatus [=Ectopsocus spathulatus] B55
    |--E. spatiosimanubrus Li 1999 L02
    |--E. tenuimanubrius Li 2002 L02
    |--E. terricolis Badonnel 1955 B55
    |--E. trimaculatus Li 2002 L02
    `--E. xerophylla Vishnyakova 1970 [=Ectopsocopsis xerophyllus, Ectopsocoides (l. c.) xerophylla] L98

Ectopsocopsis cryptomeriae (Enderlein 1907) [=Ectopsocus cryptomeriae, Ectopsocoides (l. c.) cryptomeriae; incl. Ectopsocus lepnevae Danks 1955] L98

Ectopsocopsis pumilis (Banks 1920) L02 [=Peripsocus pumilis M93, Ectopsocus pumilis M93; incl. Ectopsocus josephi Galil 1984 M93]

*Type species of generic name indicated

REFERENCES

[B49] Badonnel, A. 1949. Psocoptères de la Cote d'Ivoire. Mission Paulian-Delamare (1945). Revue Française d'Entomologie 16: 20–46.

[B55] Badonnel, A. 1955. Psocoptères de l'Angola. Diamang Publicações Culturais 26: 11–267.

[L02] Li F. 2002. Psocoptera of China vol. 1. Science Press: Beijing.

[L98] Lienhard, C. 1998. Faune de France. France et Régions Limitrophes. 83. Psocoptères Euro-Méditerranéens. Fédération Française des Sociétés de Sciences Naturelles: Paris.

[L03] Lienhard, C. 2003. Nomenclatural amendments concerning Chinese Psocoptera (Insecta), with remarks on species richness. Revue Suisse de Zoologie 110 (4): 695–721.

[M93] Mockford, E. L. 1993. North American Psocoptera (Insecta). Sandhill Crane Press, Inc.

[NL07] New, T. R., & C. Lienhard. 2007. The Psocoptera of Tropical South-east Asia. Brill: Leiden.

Ectopsocus

Ectopsocus briggsi, copyright Christophe Quintin.


Belongs within: Lachesilloidea.

Ectopsocus is a genus of barklice including some of the most widespread psocid species not directly associated with humans. Males have a clunium bearing a row of teeth or other ornamentation; females have all three pairs of gonapophyses present and the subgenital plate is usually bilobed and setose (New & Lienhard 2007).

<==Ectopsocus McLachlan 1899 SN08 (see below for synonymy)
    |--*E. briggsi McLachlan 1899 SN08 [incl. E. limbatus Navás 1909 M93]
    |--E. adelphos Thornton & Wong 1968 NL07
    |--E. aethiops (Hagen 1859) NL07
    |--E. amblyura Thornton & Wong 1968 NL07
    |--E. amphithrix Thornton & Wong 1968 NL07
    |--E. anisodontus Li 2002 L02
    |--E. argus Thornton & Wong 1968 NL07
    |--E. axillaris (Smithers 1969) [=Interpsocus axillaris] SN08
    |--E. baliosus Thornton & Wong 1968 NL07
    |--E. barlowi New & Lee 1992 NL07
    |--E. basalis (Banks 1937) NL07
    |--E. berlesii Ribaga 1900 (n. d.) L98
    |--E. bicaudatus B55
    |--E. bifurcatus Vaughan, Thornton & New 1991 NL07
    |--E. brinchangensis New & Lee 1992 NL07
    |--E. brunneus (Edwards 1950) SN08 [=*Interpsocus brunneus NL07]
    |--E. californicus (Banks 1903) SN08 (see below for synonymy)
    |--E. cetratus SN08
    |--E. cinctus Thornton 1962 [=Estipulaceus cinctus] NL07
    |--E. cirratus Thornton & Wong 1968 NL07
    |--E. cluniatus Vaughan, Thornton & New 1989 NL07
    |--E. coccophilus Ball 1943 NL07
    |--E. columbianus Badonnel 1986 B86
    |--E. comitus Thornton & Wong 1968 [=Estipulaceus comitus] NL07
    |--E. coyae Schmidt & New 2008 SN08
    |--E. crinitus Thornton & Wong 1968 NL07
    |--E. cristatus Thornton & Wong 1968 NL07
    |--E. decenipunctatus Li 1993 [=E. decemipunctatus (l. c.), E. decenipnctatus] L03
    |--E. denervus Thornton & Wong 1968 NL07
    |--E. denotatus Thornton & Wong 1968 NL07
    |--E. dicroglossus Thornton & Wong 1968 NL07
    |--E. drepanus Thornton 1984 NL07
    |--E. edwardsi New 1973 SN08
    |--E. equidentus Li 1999 [=E. equicornutus] L03
    |--E. erosus (Enderlein 1903) NL07
    |--E. ewarti Vaughan, Thornton & New 1989 NL07
    |--E. femestratus Thornton & Wong 1968 L02
    |--E. fumidus Thornton & Wong 1968 NL07
    |--E. furcatus Thornton & Wong 1968 NL07
    |--E. gentingensis New & Lee 1992 NL07
    |--E. ghesquierei Ball 1943 L02
    |--E. graminus Schmidt & New 2008 SN08
    |--E. gyroancistrus Li 2002 L02
    |--E. heurni (Navás 1924) NL07
    |--E. hickmani Schmidt & New 2008 SN08
    |--E. hirsutus Thornton 1962 NL07 [=Ectotrichus hirsutus L03]
    |--E. hypandrus Thornton 1984 NL07
    |--E. inclinans Li 2002 L02
    |--E. innotatus Thornton & Wong 1968 NL07
    |--E. insularis Smithers & Thornton 1974 L02
    |--E. intersitus Thornton & Wong 1968 NL07
    |--E. isodentus Li 1997 L02
    |--E. kepongensis New & Lee 1992 NL07
    |--E. leteusistrius Li 2002 L02
    |--E. longitudinalis Li 2002 L02
    |--E. machadoi Badonnel 1955 B55
    |--E. macrotaenious Li 2002 L02
    |--E. maindroni Badonnel 1935 [=Estipulaceus maindroni] NL07
    |--E. marginatus M93
    |--E. meridionalis Ribaga 1904 NL07 [=E. briggsi var. meridionalis M93]
    |--E. musae (Kunstler & Chaine 1902) L98
    |--E. myrmecophilus (Enderlein 1903) NL07
    |--E. nerens (Hickman 1934) [=Micropsocus nerens] SN08
    |--E. nidicolus Thornton & Wong 1968 NL07
    |--E. nyingchiensis Li 2002 [=E. nyingchinensis] L02
    |--E. ornatus Thornton 1962 NL07 [=Ectotrichus ornatus L03]
    |--E. paraplesius Thornton & Wong 1968 NL07
    |--E. pearmani MG56
    |--E. petersi Smithers 1978 [incl. E. australis Schmidt & Thornton 1993] SN08
    |--E. pictus Mockford 1974 M74
    |--E. pilosus Badonnel 1967 NL07
    |--E. predanus Vaughan, Thornton & New 1989 NL07
    |--E. proctus New & Lee 1992 NL07
    |--E. punctatus M83
    |--E. quadratiapicius Li 2002 [=E. quadratiapicis] L03
    |--E. quadrisetus Li 2002 L02
    |--E. richardsi (Pearman 1929) L03 [=*Chaetopsocus richardsi NL07, Ectotrichus richardsi L03]
    |--E. ridderi Vaughan, Thornton & New 1989 NL07
    |--E. rileyae Schmidt & Thornton 1993 SN08
    |--E. risdonensis Schmidt & New 2008 SN08
    |--E. salpinx Thornton & Wong 1968 NL07
    |--E. setulosus Li 1997 L03, L02 [=*Ectotrichus setulosus L03]
    |--E. similis Badonnel 1955 B55
    |--E. speciosus Thornton & Wong 1968 NL07
    |--E. spilocephalus Li 1993 L02
    |--E. spilotus M93
    |--E. sprenti Schmidt & New 2008 SN08
    |--E. stictus Thornton & Wong 1968 NL07
    |--E. strauchi Enderlein 1906 [incl. Peripsocus opulentus Navás 1908] M93
    |--E. strictifoliatus Li 2002 L02
    |--E. sumatrensis Vaughan, Thornton & New 1991 NL07
    |--E. tenellus Thornton & Wong 1968 NL07
    |--E. thibaudi Badonnel 1979 M93
    |--E. thoi New & Lee 1992 NL07
    |--E. thysanus Thornton & Wong 1968 NL07
    |--E. tinctus Navás 1924 NL07
    |--E. titschacki Jentsch 1939 NL07 [incl. E. gabelensis Badonnel 1955 M93]
    |--E. triangulus Thornton & Wong 1968 NL07
    |--E. tridentatus Thornton 1962 L02 [=E. meridionalis tridentatus L98]
    |--E. uncinatus M93
    |--E. vachoni Badonnel 1945 NL07 [incl. E. dimorphus Mockford & Gurney 1956 M93]
    |--E. vacuus New & Lee 1992 NL07
    |--E. vannus Thornton & Wong 1968 NL07
    |--E. venosus Lienhard 2002 NL07
    |--E. vilhenai Badonnel 1955 NL07
    |--E. vishnyakovae Schmidt 1993 (see below for synonymy) L98
    |--E. waterstradti (Enderlein 1901) [=*Micropsocus waterstradti] NL07
    |--E. yongi New & Lee 1992 NL07
    `--E. zayuensis Li 2002 L02

Ectopsocus McLachlan 1899 SN08 [incl. Chaetopsocus Pearman 1929 L03, Ectotrichus Li 2002 L03, Interpsocus Edward 1950 L02, Micropsocus Enderlein 1901 L02]

Ectopsocus californicus (Banks 1903) SN08 [=Peripsocus californicus SN08; incl. E. congener Tillyard 1923 M93, E. froggatti Enderlein 1906 SN08]

Ectopsocus vishnyakovae Schmidt 1993 [incl. E. brunneus Vishnyakova in Svadzhyan et al. 1963 non Interpsocus brunneus Edwards 1950] L98

*Type species of generic name indicated

REFERENCES

[B55] Badonnel, A. 1955. Psocoptères de l'Angola. Diamang Publicações Culturais 26: 11–267.

[B86] Badonnel, A. 1986. Psocoptères de Colombie (Insecta, Psocoptera). Spixiana 9 (2): 179–223.

[L02] Li F. 2002. Psocoptera of China vol. 1. Science Press: Beijing.

[L98] Lienhard, C. 1998. Faune de France. France et Régions Limitrophes. 83. Psocoptères Euro-Méditerranéens. Fédération Française des Sociétés de Sciences Naturelles: Paris.

[L03] Lienhard, C. 2003. Nomenclatural amendments concerning Chinese Psocoptera (Insecta), with remarks on species richness. Revue Suisse de Zoologie 110 (4): 695–721.

[M83] Martin, N. A. 1983. Miscellaneous observations on a pasture fauna: an annotated species list. DSIR Entomology Division Report 3: 1–98.

[M74] Mockford, E. L. 1974. Records and descriptions of Cuban Psocoptera. Entomologica Americana 48 (2): 103–215.

[M93] Mockford, E. L. 1993. North American Psocoptera (Insecta). Sandhill Crane Press, Inc.

[MG56] Mockford, E. L., & A. B. Gurney. 1956. A review of the psocids, or book-lice and bark-lice, of Texas (Psocoptera). Journal of the Washington Academy of Sciences 46 (11): 353–368.

[NL07] New, T. R., & C. Lienhard. 2007. The Psocoptera of Tropical South-east Asia. Brill: Leiden.

[SN08] Schmidt, E. R., & T. R. New. 2008. The Psocoptera (Insecta) of Tasmania, Australia. Memoirs of Museum Victoria 65: 71–152.

Pseudocaeciliinae

Heterocaecilius lachlani, from Insects of Tasmania.


Belongs within: Pseudocaecilioidea.

The Pseudocaeciliinae are a group of barklice with two-segmented tarsi, known predominantly from south-east Asia and the Pacific (New & Lienhard 2007).

<==Pseudocaeciliinae [Meplerinae, Pseudoscottiinae] NL07
    |--*Allopsocus marginalis Banks 1920 NL07
    |--Scottiella NL07
    |--Platyocaecilius Li 2002 L02b
    |    `--*P. parallelivenius Li 2002 L02b
    |--Fashengocaecilius Lienhard 2003 L03 [=Calocaecilius Li 2002 non Mockford 1974 L02b]
    |    `--*F. biaduncus (Li 1992) [=Allocaecilius biaduncus, *Calocaecilius biaduncus] L03
    |--Phyllocaecilius Li 2002 L02b
    |    `--*P. atrichus Li 2002 L02b
    |--Chorocaecilius Li 2000 L02a
    |    |--C. brunellus (Tillyard 1923) (see below for synonymy) SN08
    |    `--C. decemdentatus Li 2000 L02a
    |--Phallocaecilius Lee & Thornton 1967 L02b
    |    |--*P. hirsutus (Thornton 1961) [=Pseudocaecilius hirsutus] NL07
    |    `--P. sentosus Li 1999 L02a
    |--Thelocaecilius Li 2002 L02b
    |    |--*T. mecokeratus Li 2002 L02b
    |    `--T. papillatus (Li 1995) [=Heterocaecilius papillatus] L02b
    |--Neocaecilius Li 2000 L02a
    |    |--*N. campanula [=Heterocaecilius campanula] NL07
    |    |--N. granulatus (Lee & New 1992) NL07
    |    |--N. mutatus (Lee & New 1992) NL07
    |    `--N. qianshanensis Li 2000 L02a
    |--Mesocaecilius Okamoto 1910 NL07, L02a [Mesocaeciliinae]
    |    |--*M. quadrimaculatus Okamoto 1910 L02a
    |    |--M. bellus Li 2002 L02b
    |    |--M. decorosus Li 2002 L02b
    |    |--M. elegans Li 1997 L02b
    |    |--M. euryopterus Li 2002 L02b
    |    |--M. lepidus Li 2002 L02b
    |    `--M. paraornatus (New 1977) [=Pseudocaecilius paraornatus] L02b
    |--Ophiodopelma Enderlein 1908 L02a
    |    |--*O. ornatipenne Enderlein 1908 NL07, L02a
    |    |--O. anocella Li 2002 L02b
    |    |--O. fluctosa Li 1995 L02a
    |    |--O. glyptocephalum Lienhard 1985 NL07
    |    |--O. hieroglyphicum (Enderlein 1903) NL07
    |    |--O. multipunctatum (Hagen 1859) NL07
    |    |--O. permaculatum Lee & Thornton 1967 NL07
    |    |--O. pictipenna Lee & Thornton 1967 NL07
    |    |--O. polyspila Li 1993 L02b
    |    `--O. semiceps Lee & Thornton 1967 NL07
    |--Levucaecilius Smithers 1995 NL07, S95
    |    |--*L. greenwoodi [=Pseudocaecilius greenwoodi, Orbocaecilius greenwoodi] NL07
    |    |--L. albicrus (Thornton 1981) [=Heterocaecilius albicrus] S95
    |    |--L. apicalis (Thornton 1981) [=Heterocaecilius apicalis] S95
    |    |--L. dardanus (Lee & Thornton 1967) S95 [=Heterocaecilius dardanus NL07, Neocaecilius dardanus NL07]
    |    |--L. hilli Smithers 1995 NL07
    |    |--L. panicus (Lee & Thornton 1967) [=Heterocaecilius panicus] S95
    |    |--L. pictus (Thornton 1981) [=Heterocaecilius pictus] S95
    |    |--L. simplex (Lee & Thornton 1967) NL07, L02a [=Heterocaecilius simplex NL07]
    |    |--L. veitchi [=Heterocaecilius veitchi] S95
    |    `--L. volatus (Lee & Thornton 1967) [=Heterocaecilius volatus] S95
    |--Lobocaecilius Lee & Thornton 1967 L02a
    |    |--*L. cynara Lee & Thornton 1967 L02a
    |    |--L. arcuatus Vaughan, Thornton & New 1989 NL07
    |    |--L. bifurcus Li 1993 L02a
    |    |--L. drepanus New & Thornton 1976 NL07
    |    |--L. ligur Lee & New 1992 NL07
    |    |--L. longifurca Lee & Thornton 1967 NL07
    |    |--L. pahangensis Lee & New 1992 NL07
    |    |--L. perductivirgus Li 2002 L02b
    |    |--L. pilosus Lee & New 1992 NL07
    |    |--L. quadripartitus Li 1995 L02b
    |    |--L. thorntoni Lee & New 1992 NL07
    |    `--L. viridis Lee & New 1992 NL07
    |--Allocaecilius Lee & Thornton 1967 L02a
    |    |--*A. heterothrax Lee & Thornton 1967 L02a
    |    |--A. albidorsualis Li 1993 L02a
    |    |--A. arcuatus Li 2002 L02b
    |    |--A. elongatus Lee & Thornton 1967 NL07
    |    |--A. fusciceps Lee & Thornton 1967 NL07
    |    |--A. gombakensis Lee & New 1992 NL07
    |    |--A. guangdongicus Li 1993 L02a
    |    |--A. maculatus Lee & Thornton 1967 NL07
    |    |--A. selangorensis New & Thornton 1975 NL07
    |    |--A. sinensis Lee & Thornton 1967 L02a
    |    |--A. smithersi Lee & New 1992 NL07
    |    `--A. tenuilongus Li 1995 L02a
    |--Mepleres Enderlein 1926 SN08 [incl. Meniscopsocus Li 1993 L02b, Pseudoscottiella Badonnel 1946 L02b]
    |    |--*M. maeandricus Enderlein 1926 SN08
    |    |--M. cornutus (Lee & New 1992) NL07 [=Pseudoscottiella cornuta L02b]
    |    |--‘Pseudoscottiella’ decolor Badonnel 1955 B55
    |    |--‘Pseudoscottiella’ hyalina Badonnel 1955 B55
    |    |--‘Pseudoscottiella’ immaculata Badonnel 1955 B55
    |    |--M. indicus (Lee & New 1992) NL07
    |    |--M. limbatus (Enderlein 1908) L02a
    |    |--M. longicellus Li 2002 L02b
    |    |--M. longitudinalis Li 2002 [=M. longitudianlis] L03
    |    |--‘Pseudoscottiella’ maculata Broadhead & Richard 1982 L02b
    |    |--*Pseudoscottiella’ megops NL07
    |    |--M. nigroguttatus (Karny 1925) NL07 [=Pseudoscottiella nigraguttata L02b]
    |    |--M. ocellatus Li 2002 L02b
    |    |--M. oresbius Li 2002 L02b
    |    |--M. parvicellus Li 2002 L02b
    |    |--M. pisiformis (Lee & New 1992) NL07
    |    |--M. plenimacularis Li 2002 L02b [=Pseudoscottiella plenimacularis L02a]
    |    |--M. proboscideus Li 2002 L02b
    |    |--M. procurrens Li 2002 L02b
    |    |--M. radialis (Thornton 1984) NL07
    |    |--M. raphalis (Lee & New 1992) NL07
    |    |--M. rotundatus (New 1974) [=Pseudoscottiella rotundata, Meniscopsocus rotundatus] SN08
    |    |--M. similis (Lee & New 1992) NL07
    |    |--M. sinicus (Li 1993) L02b [=*Meniscopsocus sinicus NL07]
    |    |--M. solenocerus (Lee & New 1992) NL07
    |    |--M. suella (Thornton 1984) NL07 [=Pseudoscottiella suella L02b]
    |    |--M. tanei (Smithers 1977) [=Pseudoscottiella tanei, Meniscopsocus tanei] SN08
    |    |--M. transversus (Banks 1937) NL07
    |    |--‘Pseudoscottiella’ tuberculata Badonnel 1955 B55
    |    |--M. ulterior Li 2002 L02b
    |    |--M. unitus Li 2002 L02b
    |    |--M. urbanus (Vaughan, Thornton & New 1991) NL07
    |    |--M. vilus (Lee & New 1992) NL07
    |    |--‘Pseudoscottiella’ yeni New 1974 L02b
    |    `--M. yunnanicus Li 2002 L02b
    |--Pseudocaecilius Enderlein 1903 Y02 [incl. Hageniella Enderlein 1903 (preoc.) L02a]
    |    |--P. citricola (Ashmead 1879) NL07 (see below for synonymy)
    |    |--P. angustus (Enderlein 1903) NL07
    |    |--P. bibulbus Li 1993 L03, L02a [=P. bibalbus (l. c.) L03]
    |    |--P. bicostatus Li 2002 L02b
    |    |--P. ceratocercus Li 2002 L02b
    |    |--P. claggi (Banks 1937) NL07
    |    |--P. clunialis Vaughan, Thornton & Neo 1991 NL07
    |    |--P. cornutus Lee & Thornton 1967 NL07
    |    |--P. danaus Lee & New 1992 NL07
    |    |--P. euryocercus Li 2002 [=P. eurocerus] L03
    |    |--P. formosanus (Banks 1937) L02a
    |    |--P. galactozonalis Li 2002 [=P. galactozonacis] L03
    |    |--P. helicoides Lee & Thornton 1967 NL07
    |    |--P. immaculatus Li 2002 L02b
    |    |--P. inaequalis (Banks 1916) NL07
    |    |--P. innotatus Banks 1937 NL07
    |    |--P. kagoshimensis Y02
    |    |--P. largicellus Li 2002 L02b
    |    |--P. machadoi Badonnel 1955 B55
    |    |--P. monotaeniatus Li 2002 [=P. monotaenitus, P. monotaenius] L03
    |    |--P. nitoris (Banks 1937) NL07
    |    |--P. octomaculatus (Li 1995) [=Heterocaecilius octomaculatus] L02b
    |    |--P. otiosus (Banks 1937) NL07
    |    |--P. papillaris Li 2002 L02b
    |    |--P. plagiozonalis Li 1995 L02a
    |    |--P. productus New & Thornton 1976 NL07
    |    |--P. pusillus (Banks 1931) NL07
    |    |--P. ranus Thornton 1984 NL07
    |    |--P. serratus Lee & New 1992 NL07
    |    |--P. setifer Vaughan, Thornton & New 1991 NL07
    |    |--P. sexdentatus Li 2002 L02b
    |    |--P. tahitiensis (Karny 1926) [=Epipsocus tahitiensis] M93
    |    |--P. tenellus Enderlein 1926 NL07
    |    |--P. testaceus Enderlein 1903 NL07
    |    |--P. undecimimaculatus Li 1995 L02b
    |    |--P. utricularis Li 1999 L02a
    |    |--P. venimaculatus Li 2002 L02b
    |    |--P. wellsae Lee & New 1992 NL07
    |    `--‘Psocus’ zonatus [=*Hageniella zonata] NL07
    `--Heterocaecilius Lee & Thornton 1967 L02a [incl. Orbocaecilius Li 1999 L03, Obrocaecilius (l. c.) L03]
         |--*H. minutus Lee & Thornton 1967 SN08 [=H. minotus NL07]
         |--H. abacutidens Li 2002 L02b
         |--H. analis Vaughan, Thornton & New 1991 NL07
         |--H. anareolatus New 1993 NL07
         |--H. anomalus (Thornton 1961) Y02, L02b [=Orbocaecilius anomolus L02b]
         |--H. argutus (Li 2002) [=Obrocaecilius (l. c. for Orbocaecilius) argutus] L03
         |--H. bicruris (Li 2002) [=Obrocaecilius (l. c. for Orbocaecilius) bicruris] L03
         |--H. bidigitatus Li 1992 L02a [=Orbocaecilius bidigitatus L03]
         |--H. bifarius (Li 2002) [=Obrocaecilius (l. c. for Orbocaecilius) bifarius] L03
         |--H. brachystigmus (Li 2002) [=Obrocaecilius (l. c. for Orbocaecilius) brachystigmus] L03
         |--H. brevifurca Thornton 1984 NL07
         |--H. challengerius Lee & New 1992 NL07
         |--H. circulicellus Li 1995 L02b [=Orbocaecilius circulicellus L03]
         |--H. dasoceratus (Li 2002) [=Obrocaecilius (l. c. for Orbocaecilius) dasoceratus] L03
         |--H. deloneurous Li 2002 L02b
         |--H. dentalis Li 2002 L02b
         |--H. duodecidentus (Li 2002) [=Obrocaecilius (l. c. for Orbocaecilius) duodecidentus] L03
         |--H. fasciatus Lee & Thornton 1967 NL07
         |--H. flaviformis Lee & New 1992 NL07
         |--H. fumosus Thornton 1984 NL07
         |--H. fuscipalpus Lee & Thornton 1967 NL07
         |--H. fuscus Y02
         |--H. gressitti Lee & Thornton 1967 NL07
         |--H. hepaticivenus Li 2002 L02b
         |--H. hoplites Lee & New 1992 NL07
         |--H. huananensis Li 2002 L02b
         |--H. huaxiensis Li 2002 L02b
         |--H. imparilis (Li 2002) [=Obrocaecilius (l. c. for Orbocaecilius) imparilis] L03
         |--H. jinghongicus Li 1995 L02a
         |--H. kobus Lee & Thornton 1967 NL07
         |--H. lachlani (Enderlein 1903) [=Pseudocaecilius lachlani] SN08
         |--H. leei New 1993 NL07
         |--H. liangi Li 1991 L02b
         |--H. loiaoi New 1991 L02a
         |--H. longicornis (Li 2002) [=Obrocaecilius (l. c. for Orbocaecilius) longicornis] L03
         |--H. longipenna Lee & Thornton 1967 NL07
         |--H. longispinus Lee & Thorton 1967 NL07
         |--H. maculans Li 1991 L02b
         |--H. maculifornis (Thornton 1961) L02b [=Pseudocaecilius maculifrons L02a]
         |--H. maculosus Lee & New 1992 NL07
         |--H. marginatus Lee & New 1992 NL07
         |--H. mecotrichus Li 2002 L02b
         |--H. odontothelus Li 2002 L02b
         |--H. orientalis Lee & New 1992 NL07
         |--H. ornatus (Enderlein 1903) [=Pseudocaecilius ornatus] NL07
         |--H. papillosus Vaughan, Thornton & New 1991 NL07
         |--H. paulicellus (Li 1999) NL07, L02a [=*Orbocaecilius paulicellus L02a]
         |--H. platokeratis Li 1993 L02a
         |--H. pseudoanomalus Li 1993 L02b [=Obrocaecilius (l. c. for Orbocaecilius) pseudoanomalus L03]
         |--H. quadrangulus Li 1992 L02b
         |--H. reticulatus Lee & New 1992 NL07
         |--H. rhadnus Thornton 1984 NL07
         |--H. sedecimidentus (Li 2002) [=Obrocaecilius (l. c. for Orbocaecilius) sedecimidentus] L03
         |--H. stenokeratis Li 1993 L02a
         |--H. stiliger Lee & Thornton 1967 NL07
         |--H. taipingensis Garcia Aldrete 1999 NL07
         |--H. tartareus Li 2002 L02b
         |--H. vulturius (Li 2002) [=Obrocaecilius (l. c. for Orbocaecilius) vulturius] L03
         `--H. xihuicus (Li 2002) [=Obrocaecilius (l. c. for Orbocaecilius) xihuicus] L03

Chorocaecilius brunellus (Tillyard 1923) [=Caecilius brunellus, Heterocaecilius brunellus, Pseudocaecilius brunellus; incl. H. diogenes Lee & Thornton 1967, *Chorocaecilius diogenes] SN08

Pseudocaecilius citricola (Ashmead 1879) NL07 [=Psocus citricola M93; incl. Pse. elutus var. africanus Badonnel 1931 L02a, Elipsocus criniger Perkins 1899 M93, Kilauella criniger M93, Pseudocaecilius criniger M93, *Pse. elutus Enderlein 1903 NL07, L02b, Trichopsocus (*Pseudocaecilius) elutus L02a, Trichopsocus indicatus Navás 1934 M93, Pse. medianicostatus Li 1993 L02a, Caecilius pretiosus Banks 1920 M93, Pse. pretiosus M93, Pse. wolcotti Banks 1924 M93]

*Type species of generic name indicated

REFERENCES

[B55] Badonnel, A. 1955. Psocoptères de l'Angola. Diamang Publicações Culturais 26: 11–267.

[L02a] Li F. 2002a. Psocoptera of China vol. 1. Science Press: Beijing.

[L02b] Li F. 2002b. Psocoptera of China vol. 2. Science Press: Beijing.

[L03] Lienhard, C. 2003. Nomenclatural amendments concerning Chinese Psocoptera (Insecta), with remarks on species richness. Revue Suisse de Zoologie 110 (4): 695–721.

[M93] Mockford, E. L. 1993. North American Psocoptera (Insecta). Sandhill Crane Press, Inc.

[NL07] New, T. R., & C. Lienhard. 2007. The Psocoptera of Tropical South-east Asia. Brill: Leiden.

[SN08] Schmidt, E. R., & T. R. New. 2008. The Psocoptera (Insecta) of Tasmania, Australia. Memoirs of Museum Victoria 65: 71–152.

[S95] Smithers, C. N. 1995. Psocoptera (Insecta) of Christmas Island. Invertebrate Taxonomy 9: 529–561.

[Y02] Yoshizawa, K. 2002. Phylogeny and higher classification of suborder Psocomorpha (Insecta: Psocodea: ‘Psocoptera’). Zoological Journal of the Linnean Society 136: 371–400.