Poronoticae

Dorsal (left) and ventral views of Unduloribates hebes, from Norton & Behan-Pelletier (2009).

Belongs within: Brachypylina.
Contains: Oribatellidae, Anachipteria, Achipteria, Campachipteria, Tegoribatidae, Heterobelbidae, Tectocepheidae, Microtegeidae, Oripodoidea, Licneremaeoidea, Scapheremaeus, Eremaeozetoidea, Ameronothroidea, Galumnoidea, Ceratozetoidea, Phenopelopidae.

The Poronoticae are a group of oribatid mites possessing an octotaxic system, a series of glandular openings (most typically four pairs) on the notogaster in the form of porose areas or saccules. Monophyly of the group is uncertain; a number of taxa lacking the octotaxic system are otherwise similar in morphology to clearly poronotic species and it is probable that the octotaxic system has evolved more than once and/or been lost in some subgroups.

Members of the Poronoticae include the Achipterioidea, known from soil, litter, mosses and liverworts. Members of the Achipteriidae have characteristic spiral internal thickenings in the bothridium and a distinct long, thin, dorsally concave genu in leg IV (Norton & Behan-Pelletier 2009). The Epactozetidae are a Neotropical family with large, auriculate pteromorphs and foveate patterning on the notogaster (Norton & Behan-Pelletier 2009).

Poronoticae B65
    |  i. s.: Oribatellidae B65
    |--Achipterioidea PL17
    |    |--Epactozetidae NB-P09
    |    |    |--Epactozetes Grandjean 1930 S04
    |    |    |    |--*E. imitator Grandjean 1930 S04
    |    |    |    `--E. setosus Balogh & Mahunka 1969 S04
    |    |    `--Truncozetes Balogh & Mahunka 1969 S04
    |    |         |--*T. mucronatus Balogh & Mahunka 1969 S04
    |    |         |--T. rugosus Mahunka 1998 S04
    |    |         `--T. sturmi Balogh 1984 S04
    |    |--Achipteriidae S04
    |    |    |  i. s.: Hoffmanacarus Mahunka 1995 S04
    |    |    |           `--*H. virginianus Mahunka 1995 S04
    |    |    |         Cubachipteria Balogh & Mahunka 1979 S04
    |    |    |           |--*C. remota (Balogh & Mahunka 1979) [=Achipteria (*Cubachipteria) remota] S04
    |    |    |           `--C. bispina (Calugar 1990) [=Achipteria (Cubachipteria) bispina] S04
    |    |    |         Dentachipteria Nevin 1974 S04
    |    |    |           |--*D. ringwoodensis Newin 1974 S04
    |    |    |           `--D. highlandensis Nevin 1974 S04
    |    |    |         Novoribatella Engelbrecht 1986 NB-P09, S04
    |    |    |           |--*N. transcripta (Mahunka 1985) [=Tectoribates transcriptus] S04
    |    |    |           |--N. minutisetarum Engelbrecht 1986 S04
    |    |    |           `--N. pentasetarum Engelbrecht 1986 S04
    |    |    |         Tectoribates Berlese 1910 NB-P09, S04 [incl. Anoribatella Kunst 1962 S04]
    |    |    |           |--*T. proximus (Berlese 1910) (see below for synonymy) S04
    |    |    |           |--T. africanus Balogh 1959 S04
    |    |    |           |--T. deserticola (Balogh & Mahunka 1965) [=Anoribatella deserticola] S04
    |    |    |           |--T. mongolicus Bayartogtokh 1997 S04
    |    |    |           |--T. ornatus (Schuster 1958) S04 [=Anachipteria ornata S04, *Anoribatella ornata B65]
    |    |    |           `--T. parvus Sellnick 1931 S93
    |    |    |         Cerachipteria Grandjean 1935 [incl. Zygachipteria Mihelčič 1956] S04
    |    |    |           |--*C. digita Grandjean 1935 S04
    |    |    |           |    |--C. d. digita S04
    |    |    |           |    |--C. d. jugata Mihelčič 1956 [=C. (Zygachipteria) jugata] S04
    |    |    |           |    `--C. d. pyrenaica Pérez-Íñigo 1991 [incl. C. fusiger Mihelčič 1957 (n. d.)] S04
    |    |    |           |--C. franzi Willmann 1953 S04
    |    |    |           `--C. minuscula (Berlese 1902) [=Oribata minuscula] S04
    |    |    |         Parachipteria Hammen 1952 [incl. Pseudachipteria Travé 1960] S04
    |    |    |           |--*P. punctata (Nicolet 1855) [=Oribata punctata] S04
    |    |    |           |--P. agenjoi (Pérez-Íñigo 1976) [=Pseudachipteria agenjoi] S04
    |    |    |           |--P. floresiana (Pérez-Íñigo 1992) [=Pseudachipteria floresiana] S04
    |    |    |           |--P. insularis (Pérez-Íñigo 1992) [=Pseudachipteria insularis] S04
    |    |    |           |--P. magna (Sellnick 1928) S04 [=Notaspis magna S04, *Pseudachipteria magna BB92]
    |    |    |           |--P. nicoleti (Berlese 1883) S07
    |    |    |           `--P. ovalis (Koch 1835) (n. d.) [=Oribata ovalis] S04
    |    |    |--Anachipteria PL17
    |    |    `--+--Achipteria PL17
    |    |       `--Campachipteria PL17
    |    `--Tegoribatidae B-P01
    `--+--Heterobelbidae PL17
       `--+--+--Tectocepheidae PL17
          |  `--+--Microtegeidae PL17
          |     `--Oripodoidea PL17
          `--+--+--Licneremaeoidea SK10
             |  |--Scapheremaeus SK10
             |  |--Eremaeozetoidea ME09
             |  `--Ameronothroidea SK10
             `--+--Galumnoidea SK10
                `--+--Podacarus Grandjean 1955 SK10, S04 [Podacaridae]
                   |    `--*P. auberti Grandjean 1955 S04
                   |         |--P. a. auberti S04
                   |         `--P. a. occidentalis Wallwork 1966 S04
                   `--+--Ceratozetoidea PL17
                      `--Phenopelopoidea [Pelopoidea] NB-P09
                           |  i. s.: Plakoribates Popp 1960 NB-P09, S04
                           |           |--*P. multicuspidatus Popp 1960 S04
                           |           |--P. confluens Balogh 1970 S04
                           |           |--P. neotropicus Balogh & Mahunka 1978 S04
                           |           `--P. scutatus Hammer 1979 S04
                           |--Phenopelopidae PL17
                           `--Unduloribatidae [Unduloribatoidea] NB-P09
                                |--Koreoribates Choi 1994 S04
                                |    `--*K. foliatus Choi 1994 S04
                                `--Unduloribates Balogh 1943 S04
                                     |--*U. undulatus (Berlese 1914) [=Sphaerozetes (Tectoribates) undulatus] S04
                                     |--U. brevisetosus Nübel-Reidelbach & Woas 1992 S04
                                     |--U. hebes Aoki 1965 S04
                                     |--U. medusa Piffl 1972 S04
                                     `--U. parvus NB-P09

*Tectoribates proximus (Berlese 1910) [=Sphaerozetes (*Tectoribates) proximus; incl. T. punctatus Sellnick 1943] S04

*Type species of generic name indicated

REFERENCES

[B65] Balogh, J. 1965. A synopsis of the world oribatid (Acari) genera. Acta Zoologica Academiae Scientiarum Hungaricae 11 (1–2): 5–99.

[BB92] Balogh, J., & P. Balogh. 1992. The Oribatid Mites Genera of the World vol. 1. Hungarian Natural History Museum: Budapest.

[B-P01] Behan-Pelletier, V. M. 2001. Phylogenetic relationships of Hypozetes (Acari: Tegoribatidae). In: Halliday, R. B., D. E. Walter, H. C. Proctor, R. A. Norton & M. J. Colloff (eds) Acarology: Proceedings of the 10^th International Congress pp. 50–57. CSIRO Publishing: Melbourne.

[ME09] Maraun, M., G. Erdmann, G. Schulz, R. A. Norton, S. Scheu & K. Domes. 2009. Multiple convergent evolution of arboreal life in oribatid mites indicates the primacy of ecology. Proceedings of the Royal Society of London Series B—Biological Sciences 276: 3219–3227.

[NB-P09] Norton, R. A., & V. M. Behan-Pelletier. 2009. Suborder Oribatida. In: Krantz, G. W., & D. E. Walter (eds) A Manual of Acarology 3^rd ed. pp. 430–564. Texas Tech University Press.

[PL17] Pachl, P., A. C. Lindl, A. Krause, S. Scheu, I. Schaefer & M. Maraun. 2017. The tropics as an ancient cradle of oribatid mite diversity. Acarologia 57 (2): 309–322.

[SK10] Schäffer, S., S. Koblmüller, T. Pfingstl, C. Sturmbauer & G. Krisper. 2010. Ancestral state reconstruction reveals multiple independent evolution of diagnostic morphological characters in the "Higher Oribatida" (Acari), conflicting with current classification. BMC Evolutionary Biology 10: 246.

[S07] Schatz, H. 2007. Biogeography of oribatid mites (Acari, Oribatida) from the Cordillera de Talamanca, Costa Rica and Panama. In: Morales-Malacara, J. B., V. M. Behan-Pelletier, E. Ueckermann, T. M. Pérez, E. G. Estrada-Venegas & M. Badii (eds) Acarology XI: Proceedings of the International Congress pp. 151–167. Instituto de Biología and Faculdad de Ciencias, Universidad Nacional Autónoma de México, Sociedad Latinoamericana de Acarología: México.

[S93] Selden, P. A. 1993. Arthropoda (Aglaspidida, Pycnogonida and Chelicerata). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 297–320. Chapman & Hall: London.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

Austrachipteria

Austrachipteria sp., photographed by S. E. Thorpe.

Belongs within: Hydrozetoidea.

Austrachipteria is a genus of oribatid mites that is most diverse in Australasia though a single species A. pulla is known from Japan (Subías 2004).

Characters (from Balogh & Balogh 1992): Lamellae synlamellate; pteromorphae immovable; octotaxic system sacculonotic, four pairs of sacculi; ten pairs of notograstral setae present, setae lm and lp originating almost in same longitudinal row as setae c2, h1 and h3; six pairs of genital setae present, one pair of aggenital setae, two pairs of anal setae; three pairs of adanal setae; legs tridactyle.

Austrachipteria Balogh & Mahunka 1966 [incl. Parahypozetes Hammer 1967] S04
    |--*A. lamellata Balogh & Mahunka 1966 S04
    |--A. bidactyla (Balogh & Balogh 1983) [=Parahypozetes bidactylus] S04
    |--A. bidentata (Hammer 1967) [=Parahypozetes bidentatus] S04
    |--A. breviseta (Balogh & Balogh 1983) [=Parahypozetes brevisetus] S04
    |--A. furcata (Hammer 1967) [=Parahypozetes furcatus] S04
    |--A. gigantea (Hammer 1967) [=Parahypozetes giganteus] S04
    |--A. grandis (Hammer 1967) S04 [=*Parahypozetes grandis CH98]
    |--A. hammerae Balogh & Balogh 1992 (see below for synonymy) S04
    |--A. lobata (Hammer 1967) [=Parahypozetes lobatus] S04
    |--A. macrodentata (Hammer 1967) [=Parahypozetes macrodentatus] S04
    |--A. maxima (Hammer 1967) [=Parahypozetes maximus] S04
    |--A. pulla Aoki & Honda 1985 S04
    `--A. quadridentata (Hammer 1967) [=Parahypozetes quadridentatus] S04

Austrachipteria hammerae Balogh & Balogh 1992 [incl. Parahypozetes hammerae Mahunka 1997 non A. hammerae Balogh & Balogh 1992, P. lobatus Balogh & Balogh 1983 non Hammer 1967, Austrachipteria marieae Colloff & Halliday 1998] S04

*Type species of generic name indicated

REFERENCES

Balogh, J., & P. Balogh. 1992. The Oribatid Mites Genera of the World vol. 1. Hungarian Natural History Museum: Budapest.

[CH98] Colloff, M. J., & R. B. Halliday. 1998. Oribatid Mites: A catalogue of Australian genera and species. CSIRO Publishing: Collingwood.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

Ameronothroidea

Tegeocranellus alas, copyright B. Eamer.

Belongs within: Poronoticae.
Contains: Halozetes.

The Ameronothroidea are a group of mostly semi-aquatic oribatid mites, inhabiting locations such as ephemeral rock pools or intertidal littoral zones. Gas exchange may be facilitated during immersion by a cerotegumental plastron and/or taenidia and tecta in the pleural region (Norton & Behan-Pelletier 2009). Some species of Ameronothridae are noteworthy for being found in extreme environments in the Arctic or Antarctic (Norton & Behan-Pelletier 2009). Members of the genera Antarcticola and Pseudantarcticola have fewer setae on the notogaster than other genera of Ameronothridae, with only ten or eleven pairs versus the twelve or more pairs of the latter (Balogh & Balogh 1992).

Characters (from Norton & Behan-Pelletier 2009): Adults with integument often weakly sclerotised and easily distorted. Prodorsum with or without lamellae, adjacent medially or separated, and lamellar cusps. Tutorium and translamella present or absent. Gena1 notch absent. Bothridial seta normally developed or extremely small, or absent. Chelicerae rhelate-dentate; palpal eupathidium acm separate from solenidion; subcapitular mentum without tectum. Axillary saccule of subcapitulum absent. Dorsophragmata and pleurophragmata present or absent. Discidium present or absent, custodium absent. Pedotectum I large, extending to base of bothridium, subdivided, or absent. Pedotectum II present or absent. With or without tecta laterally to maintain plastron around sejugal region and acetabulum III. Coxisternal setation various (3-1-3-2, 3-1-2-2, 3-1-3-2, 2-1-2-3, or 1-1-1-1). Genital setation 3, 5, or 6 pairs. Ovipositor and spermatopositor with 11-12 setae. Notogaster overhanging ventral plate. Lenticulus present or absent. Immatures apheredermous, plicate, opisthonotum often smooth and convex centrodorsally; unideficient or bideficient. Prodorsal, adanal, and latero-hysterosomal porose regions (or their saccule homologues) present or absent. Seta d variably present on tibiae and genua when respective solenidion is present.

<==Ameronothroidea S04
    |--Ameronothridae SK10
    |    |--Aquanothrus Engelbrecht 1975 [Aquanothridae] S04
    |    |    `--*A. montanus Engelbrecht 1975 S04
    |    |--Capillibates Hammer 1966 S04
    |    |    `--*C. stagaardi Hammer 1966 S04
    |    |--Chudalupia Wallwork 1981 S04
    |    |    `--*C. meridionalis Wallwork 1981 S04
    |    |--Pseudantarcticola Balogh 1970 S04
    |    |    `--*P. tropica Balogh 1970 S04
    |    |--Antarcticola Wallwork 1967 [incl. Petrozetes Sitnikova 1968] S04
    |    |    |--*A. meyeri Wallwork 1967 [incl. Petrozetes oblongus Sitnikova 1968] S04
    |    |    `--A. georgiae Wallwork 1970 S04
    |    |--Alaskozetes Hammer 1955 S04
    |    |    |--*A. coriaceus Hammer 1955 S04
    |    |    |--A. antarcticus (Michael 1903) S04 [=Notaspis antarctica S04, Halozetes antarctica CH98]
    |    |    |    |--A. a. antarcticus S04
    |    |    |    |--A. a. grandjeani (Dalenius 1958) S04 [=Halozetes antarctica grandjeani CH98]
    |    |    |    `--A. a. intermedius Wallork 1967 S04
    |    |    `--A. bouvetoyaensis Pletzen & Kok 1971 S04
    |    |--Ameronothrus Berlese 1896 [incl. Hygroribates Jacot 1934] S04
    |    |    |--*A. lineatus (Thorell 1871) (see below for synonymy) S04
    |    |    |--A. bilineatus (Michael 1888) [=Scutovertex bilineatus] S04
    |    |    |--A. dubinini Sitnikova 1975 S04
    |    |    |--A. harioti (Michael 1891) (n. d.) [=Scutovertex harioti] S04
    |    |    |--A. lapponicus Dalenius 1963 S04
    |    |    |--A. maculatus (Michael 1882) (see below for synonymy) S04
    |    |    |--A. marinus (Banks 1896) S04 (see below for synonymy)
    |    |    |--A. nidicola Sitnikova 1975 S04
    |    |    |--A. nigrofemoratus (Koch 1879) [=Nothrus nigrofemoratus; incl. A. lineatus brevipes Willmann 1937] S04
    |    |    |--A. oblongus Sitnikova 1975 S04
    |    |    |--A. schneideri (Oudemans 1905) [=Scutovertex schneideri] S04
    |    |    |--A. schubarti Weigmann & Schulte 1975 S04
    |    |    `--A. schusteri Schubart 1970 S04
    |    `--Halozetes S04
    |--Fortuyniidae S04
    |    |--Circellobates Luxton 1992 S04
    |    |    `--*C. venustus Luxton 1992 S04
    |    |--Alismobates Luxton 1992 S04
    |    |    |--*A. reticulatus Luxton 1992 S04
    |    |    `--A. rotundus Luxton 1992 S04
    |    `--Fortuynia Hammen 1960 S04
    |         |--*F. marina Hammen 1960 S04
    |         |--F. elamellata Luxton 1967 S04
    |         |    |--F. e. elamellata S04
    |         |    |--F. e. micromorpha Marshall & Pugh 2002 S04
    |         |    `--F. e. shibai Aoki 1974 S04
    |         |--F. inhambanensis Marshall & Pugh 2002 S04
    |         |--F. maculata Luxton 1986 S04
    |         |--F. rotunda Marshall & Pugh 2002 S04
    |         |--F. sinensis Luxton 1992 S04
    |         `--F. yunkeri Hammen 1963 S04
    |--Selenoribatidae S04
    |    |--Thalassozetes Schuster 1963 S04
    |    |    `--*T. riparius Schuster 1963 S04
    |    |--Psednobates Luxton 1992 S04
    |    |    `--*P. uncunguis Luxton 1992 S04
    |    |--Arotrobates Luxton 1992 S04
    |    |    |--*A. lanceolatus Luxton 1992 S04
    |    |    `--A. granulatus Luxton 1992 S04
    |    |--Selenoribates Strenzke 1961 S04
    |    |    |--*S. foveiventris Strenzke 1961 S04
    |    |    |--S. ghardaqensis Abdel-Hamid 1973 S04
    |    |    `--S. mediterraneus Grandjean 1966 S04
    |    `--Schusteria Grandjean 1968 S04
    |         |--*S. littorea Grandjean 1968 S04
    |         |--S. melanomerus Marshall & Pugh 2000 S04
    |         `--S. ugraseni Marshall & Pugh 2000 S04
    `--Tegeocranellus Berlese 1913 NB-P09, H98 [=Tegeogranellus H98; Tegeocranellidae]
         |--*T. laevis (Berlese 1905) [=Tegeocranus laevis] S04
         |--T. alas Behan-Pelletier 1997 S04
         |--T. barbarae Behan-Pelletier 1997 S04
         |--T. bolivianus Balogh & Mahunka 1969 S04
         |--T. bosniae (Frank 1961) [=Carabodes bosniae] S04
         |--T. concavus Balogh & Balogh 1983 S04
         |--T. convexus Balogh & Balogh 1983 S04
         |--T. kethleyi Behan-Pelletier 1997 S04
         |--T. knysnaensis Kok 1968 [incl. T. africanus Mahunka 1985] S04
         |--T. mediolamellatus Behan-Pelletier 1997 S04
         |--T. mississippii Behan-Pelletier 1997 S04
         |--T. muscorum Behan-Pelletier 1997 S04
         |--T. opcus Tseng 1982 S04
         `--T. sacchareus Kok 1968 S04

*Ameronothrus lineatus (Thorell 1871) [=Eremaeus lineatus; incl. Scutovertex corrugatus Michael 1888, S. occidentalis Hull 1918] S04

Ameronothrus maculatus (Michael 1882) [=Scutovertex maculatus; incl. S. maculatus groenlandicus Trägardh 1904, S. pseudomaculatus insularis Hull 1914, S. pseudomaculatus Hull 1914] S04

Ameronothrus marinus (Banks 1896) S04 [=Nothrus marinus S04, *Hygroribates marinus B65; incl. A. fucicolum Brady 1875 S04, Scutovertex spoofi Oudemans 1900 S04, A. spoofi G32]

*Type species of generic name indicated

REFERENCES

[B65] Balogh, J. 1965. A synopsis of the world oribatid (Acari) genera. Acta Zoologica Academiae Scientiarum Hungaricae 11 (1–2): 5–99.

[CH98] Colloff, M. J., & R. B. Halliday. 1998. Oribatid Mites: A catalogue of Australian genera and species. CSIRO Publishing: Collingwood.

[G32[ Grandjean, F. 1932. Observations sur les oribates (3^e série). Bulletin du Muséum, 2e série, 4 (3): 292–306.

[H98] Halliday, R. B. 1998. Mites of Australia: A checklist and bibliography. CSIRO Publishing: Collingwood.

[NB-P09] Norton, R. A., & V. M. Behan-Pelletier. 2009. Suborder Oribatida. In: Krantz, G. W., & D. E. Walter (eds) A Manual of Acarology 3^rd ed. pp. 430–564. Texas Tech University Press.

[SK10] Schäffer, S., S. Koblmüller, T. Pfingstl, C. Sturmbauer & G. Krisper. 2010. Ancestral state reconstruction reveals multiple independent evolution of diagnostic morphological characters in the "Higher Oribatida" (Acari), conflicting with current classification. BMC Evolutionary Biology 10: 246.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

Scheloribates (Scheloribates)

Dorsum of Scheloribates eusetosus, from Lee & Pajak (1990).

Belongs within: Scheloribatidae.

The type subgenus of the genus Scheloribates is a hyperdiverse cosmopolitan group of oribatid mites with well over 200 currently recognised species (Subías 2004). Lee & Pajak (1990) recognised three species groups primarily on the basis of a revision of Australian species. In the S. heterosetosus group, some of the hysteronotal setae are reduced to microsetae, seemingly represented by their alveoli only. In the other two species groups, all hysteronotal setae are small but definitely present. Scheloribates crassipodus has tarsi on the first three legs that are subrectangular, almost parallel-sided for over two-thirds of their length, in contrast to members of the S. eusetosus group whose tarsi taper distally for over half their length.

See also: Scheloribates.

Characters (from Balogh & Balogh 1992): Prodorsum with lamellae present, translamella absent; tutorium absent. Dorsosejugal suture arched. Notogaster with ten pairs of setae and/or setal alveoli; four pairs of sacculi; pteromorphae present, immovable. Four pairs of genital setae; one pair of aggenital setae; two pairs of anal setae; three pairs of adanal setae. Legs tridactyle.

<==Scheloribates Berlese 1908 (Scheloribates) S04 (see below for synonymy)
    |--S. (S.) crassipodus Lee & Pajak 1990 S04, LP90
    |--S. eusetosus complex LP90
    |    |--S. (S.) brevipodus Lee & Pajak 1990 LP90
    |    |--S. (S.) diforamenatus Lee & Pajak 1990 LP90
    |    |--S. (S.) eusetosus Lee & Pajak 1990 S04
    |    |--S. (S.) heterotrichus Mahunka 1984 LP90
    |    |--S. (S.) nanus Lee & Pajak 1990 LP90
    |    |--S. (S.) parabrevipodus Lee & Pajak 1990 LP90
    |    |--S. (S.) parananus Lee & Pajak 1990 LP90
    |    |--S. (S.) praeincisus (Berlese 1910) LP90 (see below for synonymy)
    |    |    |--S. p. praeincisus S04
    |    |    |--S. p. acuticlava Pérez-Íñigo & Baggio 1986 S04
    |    |    |--S. p. atlanticus Pérez-Íñigo 1982 S04
    |    |    |--S. p. cubanus Scull, Jeleva & Cruz 1984 S04
    |    |    |--S. p. interruptus (Berlese 1916) [=Protoribates (Scheloribates) praeincisus var. interruptus] S04
    |    |    |--S. p. sandvicensis (Jacot 1934) S04 (see below for synonymy)
    |    |    `--S. p. tenuiseta Hammer 1971 S04
    |    `--S. (S.) repetitivus Subías 2004 S04 (see below for synonymy)
    `--S. heterosetosus complex LP90
         |--S. (S.) heterosetosus Lee & Pajak 1990 S04
         `--S. (S.) unisetosus Lee & Pajak 1990 LP90

Scheloribates incertae sedis:
  S. (S.) pallidulus (Koch 1841) S04 (see below for synonymy)
  S. (S.) abraensis Corpuz-Raros 1980 S04
  S. (S.) acutirostris Calugar & Vasiliu 1983 S04
  S. (S.) aequalis Hammer 1967 S04
  S. (S.) aethiopicus Mahunka 1982 S04
  S. (S.) agilis (Nicolet 1855) (n. d.) [=Oribata agilis] S04
  S. (S.) agricolus Corpuz-Raros 1980 S04
  S. (S.) alexandrinae Vasiliu & Ivan 1995 S04
  S. (S.) angulatus Willmann 1931 S04
  S. (S.) artigasi Pérez-Íñigo & Baggio 1980 S04
  S. (S.) ascendens Weigmann & Wunderle 1990 S04
  S. (S.) atahualpensis Hammer 1961 S04
  S. (S.) azovicus Subbotina 1989 S04
  S. (S.) badius (Ewing 1908) [=Oribata badia] S04
  S. (S.) baloghi Kardar 1976 S04
  S. (S.) barbatulus Mihelčič 1956 [incl. S. subsimilis Mihelčič 1956] S04
  S. (S.) bhadurii Sanyal 1992 S04
  S. (S.) bhutanensis Sanyal 1988 S04
  S. (S.) bicarcualis Hammer 1973 S04
  S. (S.) bicornis Mahunka 1991 S04
  S. (S.) bicuspidatus Kardar 1976 S04
  S. (S.) bidactylus Hammer 1961 S04
  S. (S.) bilineatus Balogh 1958 (n. d.) S04
  S. (S.) biunguis (Berlese 1920) [=Protoribates (S.) biunguis] S04
  S. (S.) brachypterus Wallwork 1977 S04
  S. (S.) brasilocompressus Badejo, Woas & Beck 2002 S04
  S. (S.) brasilosphericus Badejo, Woas & Beck 2002 S04
  S. (S.) callipus (Berlese 1908) (see below for synonymy) S04
  S. (S.) camtschaticus Subbotina 1989 S04
  S. (S.) caprai Bernini 1973 (see below for synonymy) S04
  S. (S.) carneus (Hull 1916) (n. d.) [=Oribatula carnea] S04
  S. (S.) castlei (Jacot 1934) [=Protoschelobates castlei] S04
  S. (S.) chauhani Baker 1945 S04
  ‘*Semischeloribates’ clavatus Hammer 1973 CH98
  S. (S.) clavilanceolatus (Ewing 1907) [=Oribata clavilanceolata] S04
  S. (S.) clavipectinatus (Ewing 1907) [=Oribata clavipectinata] S04
  S. (S.) concentricus Balogh 1962 S04
  S. (S.) confusia Coetzer 1968 S04
  S. (S.) conirostris Corpuz-Raros 1980 S04
  S. (S.) corpusculum Bayartogtokh 2000 S04
  S. (S.) crassus Hammer 1967 S04
  S. (S.) crimicus Subbotina 1989 S04
  S. (S.) cruciseta Jeleva & Vu 1987 S04
  S. (S.) curvialatus Hammer 1961 S04
  S. (S.) cuyi Corpuz-Raros 1980 S04
  S. (S.) dalawaeus Corpuz-Raros 1980 S04
  S. (S.) danaus Subbotina 1987 S04
  S. (S.) decarinatus Aoki 1984 S04
  S. (S.) dentatus Katsumata 1988 non Oribata dentata Berlese 1883 (not preoc. if in dif. gen.) S04
  S. (S.) dimentmani (Vasiliu & Ivan 1995) [=Hemileius dimentmani] S04
  S. (S.) discifer Balogh 1959 S04
  S. (S.) distinctus Mihelčič 1964 S04
  S. (S.) diversidactylus (Hammer 1961) [=*Andeszetes diversidactylus] EA14
  S. (S.) elegans Hammer 1958 S04
  S. (S.) elegantulus Hammer 1961 S04
  S. (S.) elongatus (Warburton 1912) [=Oribata elongata] S04
  S. (S.) elsi Pletzen 1965 S04
  S. (S.) exiguus Mahunka 1992 S04
  S. (S.) feideri Calugar & Vasiliu 1983 S04
  S. (S.) femoralis (Nicolet 1855) (n. d.) [=Oribata femoralis] S04
  S. (S.) femoratus Willmann 1931 S04
  S. (S.) femoroserratus Pérez-Íñigo & Baggio 1980 S04
  S. (S.) fijiensis Hammer 1971 [=S. praeincisus fijiensis] S04
  S. (S.) filipinus (Corpuz-Raros 1980) S04 [=*Philoribates filipinus BB92]
  S. (S.) fimbriatoides Hammer 1977 S04
  S. (S.) fimbriatus Thor 1930 S04
    |--S. f. fimbriatus (see below for synonymy) S04
    |--S. f. africanus Wallwork 1964 S04
    |--S. f. calcaratus Jacot 1934 S04
    |--S. f. javensis Willmann 1931 S04
    `--S. f. whitteni Jacot 1934 S04
  S. (S.) flagellatus Wallwork 1966 S04
  S. (S.) flagellisetosus Ermilov & Anichkin 2014 EA14
  S. (S.) fuscosensillus Corpuz-Raros 1980 S04
  S. (S.) giganteus Hammer 1961 EA14 [=Megascheloribates giganteus LP90]
  S. (S.) gilvulus (Koch 1841) (n. d.) [=Zetes gilvulus] S04
  S. (S.) grandiporosus (Hammer 1973) EA14 (see below for synonymy)
  S. (S.) grandis Subbotina 1979 S04
  S. (S.) guhitanus Corpuz-Raros 1980 S04
  S. (S.) gunini Bayartogtokh 2000 S04
  S. (S.) helenensis Wallwork 1977 S04
  S. (S.) heterodactylus (Mahunka 1988) (see below for synonymy) S04
  S. (S.) huancayensis Hammer 1961 S04
  S. (S.) humeratus (Hull 1916) (n. d.) [=Oribatula humerata] S04
  S. (S.) imperfectus Hammer 1972 S04 [=Semischeloribates imperfectus BB92]
  S. (S.) indicus (Oudemans 1915) [=Murcia indica] S04
    |--S. i. indicus S04
    `--S. i. marquesalis Jacot 1934 S04
  S. (S.) jucundior (Berlese 1923) [=Protoribates (S.) parvulus jucundior] S04
  S. (S.) kraepelini (Berlese 1908) [=Protoribates (S.) kraepelini] S04
  S. (S.) labyrinthicus Jeleva 1962 S04
    |--S. l. labyrinthicus S04
    `--S. l. oscensis Pérez-Íñigo, Herrero & Pérez-Íñigo 1987 S04
  S. (S.) laevigatus (Koch 1835) (see below for synonymy) S04
  S. (S.) laminatus (Ewing 1909) [=Oribata laminata] S04
  S. (S.) laticlava Hammer 1961 EA14 [=Grandjeanobates laticlava S04]
  S. (S.) latoincisus Hammer 1973 S04
  S. (S.) leleupi Balogh 1959 S04
  S. (S.) lencoranicus Subbotina 1981 S04
  S. (S.) longiporosus Kulijev 1968 S04
  S. (S.) longisetosus Feider, Vasiliu & Calugar 1973 S04
  S. (S.) longus Kulijev 1968 S04
  S. (S.) louwi Pletzen 1965 S04
    |--S. l. louwi EA14
    `--S. l. nimirumi Coetzer 1968 EA14
  S. (S.) lucasiformis (Trägardh 1905) (n. d.) [=Oribata lucasiformis] S04
  S. (S.) luchili Wharton 1938 S04
  S. (S.) luciensis Mahunka 1985 S04
  S. (S.) luteomarginatus Hammer 1958 S04
  S. (S.) maculatus Wallwork 1977 S04
  S. (S.) magnus Abdel-Hamid, Al-Assiuty & Trrad 1983 S04
  S. (S.) manoai Jacot 1934 S04
  S. (S.) maoriensis Hammer 1968 S04
  S. (S.) marginedentatus (Trägårdh 1931) [=Chamobates marginedentata] S04
  S. (S.) matulisus Corpuz-Raros 1980 S04
  S. (S.) megalonyx (Berlese 1916) [=Protoribates (S.) megalonyx] S04
  S. (S.) milleri Jaoct 1936 S04
  S. (S.) minifimbriatus Mínguez, Subías & Ruiz 1986 (see below for synonymy) S04
  S. (S.) mochlosimilaris Badejo, Woas & Beck 2002 S04
  S. (S.) moestus (Banks 1895) [=Oribata moesta] S04
  S. (S.) muiri Jacot 1934 S04
  S. (S.) muiricius Jacot 1937 S04
  S. (S.) multiiteratus Subías 2004 [=Oribata simplex Storkan 1925 nec Gervais 1849 nec Haller 1884] S04
  S. (S.) multirepetitus Subías 2004 [=Incabates striatus Corpuz-Raros 1980 non S. striatus Hammer 1958] S04
  S. (S.) mumfordi Jacot 1934 S04 [=S. (*Paracheloribates) mumfordi CH98]
    |--S. m. mumfordi S04
    `--S. m. uapoui Jacot 1934 [=S. (Paraschelobates) mumfordi uapoui] S04
  S. (S.) neonominatus Subías 2004 [=S. baloghi Calugar & Vasiliu 1983 non Kardar 1976] S04
  S. (S.) nigeriocompressus Badejo, Woas & Beck 2002 S04
  S. (S.) nigeriosphericus Badejo, Woas & Beck 2002 S04
  S. (S.) nudus Woodring 1965 S04
  S. (S.) oahuensis Jacot 1934 S04
  S. (S.) obesus Hammer 1958 S04
  S. (S.) obsessus Subías 2004 [=S. indicus Sanyal 1992 non Murcia indica Oudemans 1915] S04
  S. (S.) obtusus Pletzen 1963 S04
  S. (S.) orixaensis Badejo, Woas & Beck 2002 S04
  S. (S.) ornatus Balogh 1959 S04
  S. (S.) oryzae Lan, Xin & Aoki 1986 EA14
  S. (S.) pacificus (Sellnick 1959) [=Styloribates pacificus] S04
  S. (S.) pahabaeus (Corpuz-Raros 1980) [=Incabates pahabaeus] S04
  S. (S.) palawanus Corpuz-Raros 1980 S04
  S. (S.) papillaris Tseng 1984 S04
  S. (S.) parabilis Woodring 1965 S04
  S. (S.) parvulus (Berlese 1916) [=Protoribates (S.) parvulus] S04
  S. (S.) parvus Pletzen 1963 S04
    |--S. p. parvus S04
    `--S. p. conglobatus Wallwork 1964 S04
  S. (S.) pauliensis Pérez-Íñigo & Baggio 1980 S04
  S. (S.) peracutus Balogh 1962 S04
  S. (S.) perforatus Wallwork 1964 S04
  S. (S.) perisi Pérez-Íñigo 1982 S04
  S. (S.) philippinensis Corpuz-Raros 1980 S04
  S. (S.) pilosus Sellnick 1943 (n. d.) S04
  S. (S.) polygonatus Balogh & Mahunka 1974 S04
  S. (S.) ponticuliger (Berlese 1916) [=Protoribates (S.) ponticuliger] S04
  S. (S.) ponticus Subbotina 1989 S04
  S. (S.) praelineatus Hammer 1977 S04
  S. (S.) praeoccupatissimus Subías 2004 [=S. minutus Mahunka 1984 non Pletzen 1965] S04
  S. (S.) praestantissimus (Berlese 1916) [=Protoribates (S.) praestantissimus] S04
    |--S. p. praestantissimus S04
    `--S. p. major (Berlese 1916) [=Protoribates (Scheloribates) praestantissimus major] S04
  S. (S.) primoricus (Subbotina 1987) [=Semischeloribates primoricus] S04
  S. (S.) pseudomochlosimilaris Badejo, Woas & Beck 2002 S04
  S. (S.) pseudoprincipalis Balogh 1962 S04
  S. (S.) pubescens Mahunka 1984 S04
  S. (S.) quintus Wunderle, Beck & Woas 1990 S04
  S. (S.) rakhali Sanyal 1992 S04
  S. (S.) rectus Hammer 1958 S04
  S. (S.) repetitus Subías 2004 [=S. crassus Kulijev 1979 non Hammer 1967] S04
  S. (S.) rigidisetosus Willmann 1951 S04
  S. (S.) robustus Pletzen 1963 S04
  S. (S.) rostropilosus Sanyal 1988 S04
  S. (S.) rotundatus Hammer 1955 S04
  S. (S.) rufafulvus Kardar 1976 S04
  S. (S.) rugiceps Balogh 1962 S04
  S. (S.) rugosus Hammer 1958 S04
  S. (S.) sacsahuamanensis Hammer 1961 S04
  S. (S.) saswatii Dhali & Bhaduri 1981 S04
  S. (S.) saudicus Bayoumi & Al-Khalifa 1985 S04
  S. (S.) saxicola (Halbert 1920) [=Oribatula saxicola] S04
  S. (S.) schauenbergi Mahunka 1988 S04
  S. (S.) semidesertus Bulanova-Zachvatkina & Mahmudova 1991 S04
  S. (S.) sergienkoae Subbotina 1987 S04
  S. (S.) sikkimensis Dhali & Bhaduri 1981 S04
  S. (S.) simplex (Gervais 1849) (n. d.) S04 [=Oribata simplex S04, *Storkania simplex CH98]
  S. (S.) sine Kulijev 1979 S04
  S. (S.) sphaeroides Hammer 1973 S04
  S. (S.) spirulatus Vasiliu & Calugar 1973 S04
  S. (S.) striatus Hammer 1958 S04
  S. (S.) striolatus Balogh 1961 S04
  S. (S.) teldanicus Vasiliu & Ivan 1995 S04
  S. (S.) thermophilus Hammer 1961 S04
    |--S. t. thermophilus S04
    `--S. t. corolevuensis Hammer 1971 S04
  S. (S.) translamellaris Kardar 1976 S04
  S. (S.) transplicatus Abdel-Hamid, Al-Assiuty & Trrad 1983 S04
  S. (S.) tricarinatus Coetzer 1968 S04
  S. (S.) tuberculatus Pérez-Íñigo, Herrero & Pérez-Íñigo 1987 S04
  S. (S.) tubuaiensis Sellnick 1959 S04
  S. (S.) turcmensis Subbotina 1987 S04
  S. (S.) uluguruensis Mahunka 1983 S04
  S. (S.) urenicus Subbotina 1978 S04
  S. (S.) vanzwaluwenburgi (Jacot 1934) [=Protoschelobates vanzwaluwenburgi] S04
  S. (S.) variabilis Pletzen 1965 S04
  S. (S.) viguerasis Scull, Jeleva & Cruz 1984 S04
  S. (S.) volcanensis Hammer 1962 S04
  S. (S.) vulgaris Hammer 1961 S04
  S. (S.) willmanni Hammer 1961 [incl. S. angulatus Hammer 1958 non Willmann 1931] S04
  S. (S.) xylobatoides Mahunka 1977 S04
  S. (S.) yezoensis Fujita & Fujikawa 1987 S04
  S. (S.) yorubaensis Badejo, Woas & Beck 2002 S04
  S. (S.) zealandicus Hammer 1967 S04

Scheloribates Berlese 1908 (Scheloribates) S04 [incl. Andeszetes Hammer 1961 EA14, Bischeloribates Mahunka 1988 S04, Neoscheloribates Hammer 1973 EA14, Parascheloribates Jacot 1934 S04, Philoribates Corpuz-Raros 1980 S04, Protoscheloribates Jacot 1934 S04, Semischeloribates Hammer 1973 S04, Storkania Jacot 1929 S04]

Scheloribates (Scheloribates) callipus (Berlese 1908) [=Protoribates (S.) callipus; incl. Protoschelobates seghetti Runkel & Kates 1947] S04

Scheloribates (Scheloribates) caprai Bernini 1973 [incl. S. baloghi Pérez-Íñigo 1982 non Kardar 1976, S. subsimilis Balogh 1962 non Mihelčič 1956] S04

Scheloribates (Scheloribates) fimbriatus fimbriatus Thor 1930 [incl. Protoribates (S.) artifex Berlese 1916 (n. d.), Oribata helvina Ewing 1909 (n. d.), P. (S.) lanceoliger Berlese 1908 (n. d.), Scheloribates penicillatus Mihelčič 1957, Galumna sylvicola Banks 1909 (n. d.)] S04

Scheloribates (Scheloribates) grandiporosus (Hammer 1973) [=*Neoscheloribates grandiporosus, S. (Andeszetes) grandiporosus] EA14

Scheloribates (Scheloribates) heterodactylus (Mahunka 1988) non Pletzen 1963 (not preoc. if in dif. gen.) [=Bischeloribates heterodactylus] S04

Scheloribates (Scheloribates) laevigatus (Koch 1835) [=Zetes laevigatus; incl. S. angustirostris Mihelčič 1957, Oribata lucasi Nicolet 1855, O. michaeli Hull 1914, Scheloribates robustus Mihelčič 1969 non Pletzen 1963] S04

Scheloribates (Scheloribates) minifimbriatus Mínguez, Subías & Ruiz 1986 [incl. Protoribates (S.) exilior Berlese 1916 (n. d.)] S04

Scheloribates (Scheloribates) pallidulus (Koch 1841) S04 [=Zetes pallidulus S04; incl. S. campestris Mihelčič 1966 S04, Protoribates (Scheloribates) frigidus Berlese 1908 S04, Zetes fuscomaculatus Koch 1841 S04, Z. latipes Koch 1844 S04, Oribates latipes CF77, Protoribates (*Scheloribates) latipes CH98]

Scheloribates (Scheloribates) praeincisus (Berlese 1910) LP90 [=Protoribates (S.) praeincisus S04; incl. Murcia insularis Oudemans 1915 S04, *Protoscheloribates insularis CH98, Scheloribates insularis SN98, S. zaherii Abdel-Hamid, Al-Assiuty & Trrad 1983 S04]

Scheloribates (Scheloribates) praeincisus sandvicensis (Jacot 1934) S04 [=Protoschelobates insularis sandvicensis S04, S. insularis sandwichensis SN98; incl. S. praeincisus rotundiclava Pérez-Íñigo & Baggio 1986 S04]

Scheloribates (Scheloribates) repetitivus Subías 2004 S04 [=S. pacificus Hammer 1967 non Styloribates pacificus Sellnick 1959 LP90]

*Type species of generic name indicated

REFERENCES

[BB92] Balogh, J., & P. Balogh. 1992. The Oribatid Mites Genera of the World vol. 1. Hungarian Natural History Museum: Budapest.

[CF77] Canestrini, G., & F. Fanzago. 1877. Intorno agli Acari Italiani. Atti del Reale Istituto Veneto di Scienze, Lettere ed Arti, Serie 5, 4: 69–208, pls 2–7.

[CH98] Colloff, M. J., & R. B. Halliday. 1998. Oribatid Mites: A catalogue of Australian genera and species. CSIRO Publishing: Collingwood.

[EA14] Ermilov, S. G., & A. E. Anichkin. 2014. A new species of Scheloribates (Scheloribates) from Vietnam, with notes on taxonomic status of some taxa in Scheloribatidae (Acari, Oribatida). International Journal of Acarology 40 (1): 109–116.

[LP90] Lee, D. C., & G. A. Pajak. 1990. Scheloribates Berlese and Megascheloribates gen. nov. from southeastern Australia, with comments on Scheloribatidae (Acarida: Cryptostigmata: Oriopodoidea). Invertebrate Taxonomy 4: 205–246.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

[SN98] Swift, S. F., & R. A. Norton. 1998. Preliminary report on oribatid mite (Acari: Oribatida) diversity in the Hawaiian Islands. Bishop Museum Occasional Papers 57: 1–44.

Last updated: 19 December 2018.

Lauritzenia (Incabates)

Dorsal view of Lauritzenia major, from Balogh & Balogh (1992 vol. 2).

Belongs within: Haplozetidae.

The subgenus Incabates of the genus Lauritzenia is a tropical and subtropical group of oribatid mites. They are distinguished from other subgenera in Lauritzenia by their tridactyle legs and terminally dilated or capitate sensillus.

Characters (from Balogh & Balogh 1992): Prodorsum with lamellae present, translamella absent; sensillus with short and dilated or capitate head. Notogaster with ten pairs of setae; four pairs of sacculi; pteromorphae movable. Four pairs of genital setae; one pair of aggenital setae; two pairs of anal setae; three pairs of adanal setae; fissures iad adanal. Legs tridactyle.

<==Lauritzenia (Incabates Hammer 1961) [incl. Canaribates Pérez-Íñigo & Peña 1994] S04
    |--L. (*I.) nuda (Hammer 1961) [=*Incabates nudus] S04
    |--L. (I.) angusta (Hammer 1971) [=Incabates angustus; incl. I. medius Hammer 1971] S04
    |--L. (I.) aokii (Choi 1985) [=Incabates aokii] S04
    |--L. (I.) atlantica Pérez-Íñigo & Peña 1996 S04
    |--L. (I.) barbata Choi & Kim 2002 S04
    |--L. (I.) canariensis Pérez-Íñigo & Peña 1997 S04
    |--L. (I.) depilis Pérez-Íñigo & Peña 1996 S04
    |--L. (I.) elegans (Kunst 1977) [=Haplozetes elegans] S04
    |--L. (I.) elongata (Lee & Shepherd 1990) [=Magnobates elongatus] S04
    |--L. (I.) globula (Lee & Shepherd 1990) [=Magnobates globulus] S04
    |--L. (I.) hispanica Morell 1991 S04
    |--‘Incabates’ macronudus Lee 1993 CH98
    |--L. (I.) major (Aoki 1970) [=Incabates major] S04
    |--L. (I.) mucronata (Lee 1993) [=Incabates mucronatus] S04
    |--L. (I.) pallida (Mihelčič 1956) [=Peloribates pallidus; incl. Protoribates meridionalis Mihelčič 1965] S04
    |--L. (I.) punctata (Lee 1993) S04 [=Incabates punctatus CH98, Indoribates punctatus S04]
    |--L. (I.) similis Subías & Gil-Martín 1995 S04
    |--L. (I.) sinuata (Pérez-Íñigo 1990) [=Haplozetes sinuatus] S04
    `--L. (I.) tenuifusus (Berlese 1916) (see below for synonymy) S04

Lauritzenia (Incabates) tenuifusus (Berlese 1916) [=Protoribates (Scheloribates) tenuifusus; incl. Canaribates chamobatoides Pérez-Íñigo & Peña 1994] S04

*Type species of generic name indicated

REFERENCES

Balogh, J., & P. Balogh. 1992. The Oribatid Mites Genera of the World vol. 1. Hungarian Natural History Museum: Budapest.

[CH98] Colloff, M. J., & R. B. Halliday. 1998. Oribatid Mites: A catalogue of Australian genera and species. CSIRO Publishing: Collingwood.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

Last updated: 19 December 2018.

Edwardzetes

Dorsal (left) and ventral views of Edwardzetes elongatus, from Balogh & Balogh (1992 vol. 2).

Belongs within: Ceratozetoidea.

Edwardzetes is a genus of oribatid mites found in cooler regions of the world. They are divided between two subgenera, with six pairs of genital setae present in the type subgenus but only five pairs present in E. (Gamerozetes) brevitutorium (Balogh & Balogh 1992).

Characters (from Balogh & Balogh 1992): Prodorsum with well separated lamellae, without cuspis; translamella medially interrupted or absent. Notogaster with ten pairs of setae; four pairs of areae porosae; pteromorphae immovable. Six pairs of genital setae; one pair of aggenital setae; two pairs of anal setae; three pairs of adanal setae. Legs tridactyle.

<==Edwardzetes Berlese 1914 S04
    |--E. (Edwardzetes) S04
    |    |--*E. (E.) edwardsi (Nicolet 1855) [=Oribata edwardsi; incl. Notaspis edwards lapponicus Trägardh 1902] S04
    |    |--E. (E.) andicola Hammer 1958 S04
    |    |--E. (E.) armatus (Hammer 1958) [=Jugatala armata] S04
    |    |--E. (E.) australis Starý & Block 1995 S04
    |    |--E. (E.) dentifer Hammer 1962 S04
    |    |--E. (E.) elongatus Wallwork 1966 S04
    |    |--E. (E.) novazealandicus Hammer 1967 S04
    |    |--E. (E.) trilobus Mihelčič 1957 S04
    |    `--E. (E.) ubali Iturrondobeitia & Caballero 2000 S04
    `--E. (Gamerozetes Balogh & Balogh 1990) S04
         `--E. (*G.) brevitutorium (Covarrubias 1967) [=Magellozetes brevitutorium] S04

*Type species of generic name indicated

REFERENCES

Balogh, J., & P. Balogh. 1992. The Oribatid Mites Genera of the World vol. 1. Hungarian Natural History Museum: Budapest.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.


Last updated: 15 December 2018.

Punctoribates

Punctoribates punctum, copyright R. Penttinen.

Belongs within: Ceratozetoidea.

Punctoribates is a primarily Holarctic genus of oribatid mites, with the species P. punctum also being known from New Zealand (Subías 2004). Species of this genus are found in damp habitats such as among litter, moss or sphagnum bogs.

Characters (from Balogh & Balogh 1992): Prodorsum with lamellae almost parallel, with short translamella and long cuspis, lamellar arrangement almost H-shaped; interlamellar region with sclerotised bridge bearing interlamellar setae; anterior tectum of notogaster overhanging bothridium and origin of interlamellar setae. Notogaster with ten pairs of setal alveoli only; four pairs of areae porosae; pteromorphae movable. Six pairs of genital setae; one pair of aggenital setae; two pairs of anal setae; three pairs of adanal setae. Legs tridactyle.

<==Punctoribates Berlese 1908 S04
    |--*P. punctum (Koch 1839) S04 [=Oribata punctum S04, Oribates punctum CH98]
    |--P. angulatus Bayartogtokh, Grobler & Cobanoglu 2000 S04
    |--P. armipes (Banks 1906) [=Galumna armipes] S04
    |--P. brevicuspidatus Bayartogtokh & Aoki 1998 S04
    |--P. eoeryi Mahunka 1972 S04
    |--P. ghilarvi Shaldybina 1969 S04
    |--P. grishinae Shaldybina 1977 S04
    |--P. latilobatus Kunst 1958 S04
    |--P. liber Paulitchenko 1991 S04
    |--P. meridianus Shaldybina 1973 S04
    |--P. minimus Shaldybina 1969 S04
    |--P. moldavicus Vasiliu & Calugar 1976 S04
    |--P. mundus Shaldybina 1973 S04
    |--P. palustris (Banks 1895) [=Oribata palustris] S04
    |--P. pannonicus Mahunka 1987 S04
    |--P. pragensis Winkler 1957 S04
    |--P. quadrivertex (Halbert 1920) AN96
    |--P. selgae (Pérez-Íñigo 1976) [=Minunthozetes selgae] S04
    |--P. sellnicki Willmann 1991 S04
    |    |--P. s. sellnicki S04
    |    `--P. s. crassirostris Strenzke 1952 S04
    |--P. sibiricus Grishina 1987 S04
    `--P. sphaericus Shaldybina 1987 S04

*Type species of generic name indicated

REFERENCES

[AN96] Alberti, G., R. A. Norton, J. Adis, N. A. Fernandez, E. Franklin, M. Kratzmann, A. I. Moreno, E. F. Ribeiro, G. Weigmann & S. Woas. 1996. Porose areas and related organs in oribatid mites (Oribatida). In: Mitchell, R., D. J. Horn, G. R. Needham & W. C. Welbourn (eds) Acarology IX vol. 1. Proceedings pp. 277–283. Ohio Biological Survey: Columbus (Ohio).

Balogh, J., & P. Balogh. 1992. The Oribatid Mites Genera of the World vol. 1. Hungarian Natural History Museum: Budapest.

[CH98] Colloff, M. J., & R. B. Halliday. 1998. Oribatid Mites: A catalogue of Australian genera and species. CSIRO Publishing: Collingwood.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

Last updated: 15 December 2018.

Humerobates

Aggregation of Humerobates rostrolamellatus, copyright Theodoor Heijerman.

Belongs within: Ceratozetoidea.

Humerobates is a cosmopolitan genus of oribatid mites in which the pteromorphs may be movable or immovable, and the notogastral setae are represented by alveoli only (Balogh & Balogh 1992). The subgenus Cordylobates is represented by a single species H. (C.) fragilis from Ascension Island in the South Atlantic.

Characters (from Balogh & Balogh 1992): Prodorsum with lamellae present, translamella absent. Notogaster with ten pairs of setal alveoli only; four pairs of areae porosae; pteromorphs well-developed, movable or immovable. Six pairs of genital setae; one pair of aggenital setae; two pairs of anal setae; three pairs of adanal setae. Legs tridactyle.

<==Humerobates Sellnick 1928 S04
    |--H. (Humerobates) (see below for synonymy) S04
    |    |--H. (H.) rostrolamellatus Grandjean 1936 S04 (see below for synonymy)
    |    |    |--H. r. rostrolamellatus S04
    |    |    |--H. r. giganteus Willmann 1939 [incl. H. fungorum giganteus] S04
    |    |    |--H. r. guadarramicus Pérez-Íñigo 1972 [incl. Chamobates magnus Mihelčič 1957 (n. d.)] S04
    |    |    `--H. r. mauritius Jacot 1936 [=H. humeralis mauritius] S04
    |    |--H. (H.) africanus (Mahunka 1984) [=Baloghobates africanus] S04
    |    |--H. (H.) cornutus (Sellnick 1959) S04 [=*Trihumerozetes cornutus BB92]
    |    |--H. (H.) crozetensis (Richters 1908) S04 (see below for synonymy)
    |    |--H. (H.) flechtmanni Pérez-Íñigo & Pérez-Íñigo 1993 S04
    |    |--H. (H.) fungorum (Linnaeus 1758) (n. d.) [=Acarus fungorum] S04
    |    |--H. (H.) nudus (Hammer 1967) S04 [=*Baloghobates nudus CH98]
    |    |--H. (H.) papuanus Balogh 1970 S04
    |    |--H. (H.) parvoglobosus (Hammer 1967) [=Baloghobates parvoglobosus] S04
    |    |--H. (H.) perkinsi Jacot 1934 S04 [=H. humeralis perkinsi S04, H. rostrolamellatus perkinsi SN98]
    |    |--H. (H.) pomboi Pérez-Íñigo 1992 S04
    |    |--H. (H.) setosus Behan-Pelletier & Mahunka 1993 S04
    |    |--H. (H.) taiwanensis Tseng 1984 S04
    |    |--H. (H.) varius Ohkubo 1982 S04
    |    `--H. (H.) zumpti Sellnick 1957 S04
    `--H. (Cordylobates Luxton 1995) S04
         `--H. (*C.) fragilis (Luxton 1995) [=*Cordylobates fragilis] S04

Humerobates Sellnick 1928 (Humerobates) [incl. Antarctozetes Balogh 1961, Baloghobates Hammer 1967, Banksinus Jacot 1938, Jeannelia Dalenius 1958 non Raffray 1913, Trihumerozetes Sellnick 1959] S04

Humerobates (Humerobates) crozetensis (Richters 1908) S04 [=Oribata crozetensis S04, *Antarctozetes crozetensis CH98, *Jeannelia crozetensis CH98]

Humerobates (Humerobates) rostrolamellatus Grandjean 1936 S04 [=Banksinus arboreus rostrolamellatus J38; incl. Oribata arborea Banks 1895 (n. d.) S04, *Banksinus arboreus J38, Humerobates humeralis arborea S59, Sphaerobates dentatus Mihelčič 1957 S04, Notaspis humeralis Sellnick 1928 non Hermann 1804 S04, *Humerobates humeralis S04, Murcia rubra Koch 1841 (n. d.) S04]

*Type species of generic name indicated

REFERENCES

[BB92] Balogh, J., & P. Balogh. 1992. The Oribatid Mites Genera of the World vol. 1. Hungarian Natural History Museum: Budapest.

[CH98] Colloff, M. J., & R. B. Halliday. 1998. Oribatid Mites: A catalogue of Australian genera and species. CSIRO Publishing: Collingwood.

[J38] Jacot, A. P. 1938. Some new western North Carolina moss-mites. Proceedings of the Entomological Society of Washington 40 (1): 10–15.

[S59] Sellnick, M. 1959. Acarina from southeastern Polynesia—II (Oribatidae). Occasional Papers of the Bernice P. Bishop Museum 22 (9): 109–152.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

[SN98] Swift, S. F., & R. A. Norton. 1998. Preliminary report on oribatid mite (Acari: Oribatida) diversity in the Hawaiian Islands. Bishop Museum Occasional Papers 57: 1–44.

Last updated: 15 December 2018.

Protoribates (Protoribates)

Protoribates lophothrichus, copyright R. Penttinen.

Belongs within: Haplozetidae.

The type subgenus of the genus Protoribates is a cosmopolitan group of oribatid mites (Subías 2004), albeit one of uncertain composition owing to confusion over the identity of the type species. True Protoribates have movable pteromorphs and had previously been referred to by the junior name Xylobates (Colloff & Halliday 1998).

Characters (from Balogh & Balogh 1992a, as Protoribates and Xylobates): Prodorsum with lamellae present, translamella absent; interlamellar setae long. Dorsosejugal suture present. Notogaster smooth; ten pairs of notogastral setae short but present; four pairs of areae porosae; areae porosae Aa, A2 and A3 round; pteromorphs well developed, movable. Four to six pairs of genital setae; one pair of aggenital setae; two pairs of anal setae; three pairs of adanal setae. Legs monodactyle.

<==Protoribates Berlese 1908 (Protoribates) (see below for synonymy) S04
    |--*P. (P.) dentatus (Berlese 1883) S04 (see below for synonymy) S04
    |--P. (P.) albidus (Ewing 1908) S04 [=Oribata albida S04, Oribatula albida CH98, *Propescheloribates albidus CH98]
    |--P. (P.) angulatus Choi 1986 S04
    |--P. (P.) antillensis (Mahunka 1985) [=Xylobates antillensis] S04
    |--P. (P.) bayanicus Bayartogtokh 2000 S04
    |--P. (P.) boraboraensis Sellnick 1959 S04
    |--P. (P.) brevisetosus (Fujita 1989) [=Xylobates brevisetosus] S04
    |--P. (P.) capucinus Berlese 1908 S04 [=Xylobates capucinus BB92]
    |    |--P. c. capucinus S04 (see below for synonymy)
    |    |--P. c. bonairensis Willmann 1936 S04
    |    `--P. c. tentaculus Mihelčič 1958 S04
    |--P. (P.) concretus (Kramer 1898) (n. d.) [=Oribata concreta] S04
    |--P. (P.) crassisetiger Choi 1986 S04
    |    |--P. c. crassisetiger S04
    |    `--P. c. nipponicus Fujita 1989 S04
    |--P. (P.) diani (Mahunka 1986) [=Xylobates diani] S04
    |--P. (P.) geonjiensis Choi 1994 S04
    |--P. (P.) gracilis (Aoki 1982) [=Xylobates gracilis] S04
    |--P. (P.) hakonensis Aoki 1994 S04
    |--P. (P.) halleri (Michael 1898) (n. d.) [=Oribata halleri; incl. O. simplex Haller 1884 non Gervais 1849] S04
    |--P. (P.) imperfectus (Banks 1906) [=Galumna imperfecta] S04
    |--P. (P.) iracemae Pérez-Íñigo & Baggio 1994 S04
    |--P. (P.) longiuscula (Koch 1841) (n. d.) [=Zetes longiuscula] S04
    |--P. (P.) lophothrichus (Berlese 1904) S04 (see below for synonymy)
    |--P. (P.) madagascarensis (Balogh 1961) [=Scheloribates madagascarensis] S04
    |--P. (P.) magnus Aoki 1982 S04
    |--P. (P.) mollicoma (Hammer 1973) [=Xylobates mollicoma] S04
    |--P. (P.) myrmecophilus Aoki & Ito 1997 S04
    |--P. (P.) nagaroensis (Fujita 1989) [=Xylobates nagaroensis] S04
    |--P. (P.) natalensis (Pletzen 1963) [=Scheloribates natalensis] S04
    |--P. (P.) oblongus (Ewing 1909) [=Oribata oblonga; incl. O. longa Ewing 1909, Xylobates longisetae Jacot 1937] S04
    |--P. (P.) paracapucinus (Mahunka 1988) [=Xylobates paracapucinus] S04
    |--P. (P.) ‘parvulus’ Willmann 1931 non Berlese 1916 S04
    |--P. (P.) pembertoni (Jacot 1934) [=Xylobates pembertoni] S04
    |--‘Xylobates’ punctulatus (Sellnick 1925) C-R79
    |--P. (P.) robustior (Jacot 1937) (see below for synonymy) S04
    |--P. (P.) rodriguezi (Mahunka 1988) [=Xylobates rodriguezi] S04
    |--P. (P.) rotundus (Aoki 2002) [=Xylobates rotundus] S04
    |--P. (P.) souchnaiensis Abdel-Hamid 1964 S04
    |--P. (P.) tenuis (Wen, Aoki & Wang 1984) [=Xylobates tenuis] S04
    |--P. (P.) triangularis (Hammer 1971) [=Xylobates triangularis] S04
    |--P. (P.) varisetiger (Wen, Aoki & Wang 1984) [=Xylobates varisetiger] S04
    `--P. (P.) yezoensis (Fujikawa 1983) [=Xylobates yezoensis] S04

Protoribates Berlese 1908 (Protoribates) [incl. Alloribates Banks 1947, Propescheloribates Jacot 1936, Styloribates Jacot 1934, Xylobates Jacot 1929] S04

Protoribates (Protoribates) capucinus capucinus Berlese 1908 S04 [incl. Xylobates capucinus angustior Jacot 1937 S04, *Alloribates singularis Banks 1947 CH98, S04]

*Protoribates (Protoribates) dentatus (Berlese 1883) S04 [=Oribata dentata S04, Pr. (Scheloribates) dentatus B16; incl. Oribata monodactyla Haller 1884 S04, Oribates monodactylus S59, Protoribates monodactylus B61, Xylobates monodactylus C-R79, Peloribates vastus Mihelčič 1956 S04]

Protoribates (Protoribates) lophothrichus (Berlese 1904) S04 [=Oribata lophothricha S04, Oribates lophotrichus CH98, *Xylobates lophotrichus CH98; incl. *Styloribates pectinatus Jacot 1934 CH98, S04, Scheloribates pectinatus P63, Xylobates prionotus Woolley 1968 S04]

Protoribates (Protoribates) robustior (Jacot 1937) [=Xylobates oblongus robustior; incl. X. capucinus robustior Jacot 1937 non X. oblongus robustior Jacot 1937] S04

*Type species of generic name indicated

REFERENCES

[B61] Balogh, J. 1961. Identification keys of world oribatid (Acari) families and genera. Acta Zoologica Hungarica 1: 243–344.

[BB92] Balogh, J., & P. Balogh. 1992. The Oribatid Mites Genera of the World vol. 2. Plates. Hungarian Natural History Museum: Budapest.

[B16] Berlese, A. 1916. Centuria terza di Acari nuovi. Redia 12: 289–338.

[CH98] Colloff, M. J., & R. B. Halliday. 1998. Oribatid Mites: A catalogue of Australian genera and species. CSIRO Publishing: Collingwood.

[C-R79] Corpuz-Raros, L. A. 1979. Philippine Oribatei (Acarina). I. Preliminary list of species and descriptions of forty new species. Philippine Agriculturist 62 (1): 1–82.

[P63] Pletzen, R. van. 1963. Studies on South African Oribatei (Acarina). Acarologia 5 (4): 690–703.

[S59] Sellnick, M. 1959. Acarina from southeastern Polynesia—II (Oribatidae). Occasional Papers of the Bernice P. Bishop Museum 22 (9): 109–152.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

Last updated: 19 December 2018.

Haplozetidae

Female Rostrozetes ovulum, copyright B. Eamer.

Belongs within: Oripodoidea.
Contains: Protoribates (Protoribates), Protoribates (Triaunguis), Peloribates (Peloribates), Lauritzenia (Incabates), Indoribates (Haplozetes).

The Haplozetidae are a group of oribatid mites with well-developed, hinged pteromorphs. Members have been divided between separate families Haplozetidae and Xylobatidae or Protoribatidae based on whether the octotaxic system is composed of (respectively) sacculi or areae porosae, but this was not regarded as significant by Norton & Behan-Pelletier (2009). The Oriental species Cribrozetes multiareolatus has the usual four pairs of areae porosae disassembled into 14–18 pairs of areae porosae rougly grouped in the original position. The dorsosejugal suture is absent in the genus Perxylobates (Balogh & Balogh 1992a). Six pairs of genital setae are present in Sicaxylobates, Afroleius, Paraxylobates, Pilobates and Pilobatella; other genera have less.

Characters (from Norton & Behan-Pelletier 2009): Length 250–650µm. Subcapitulum diarthric; chelicerae chelate-dentate. Notogaster with octotaxic system of porose areas or saccules; pteromorphs well developed, with hinge; insertions of adductor muscles for pteromorphs evident on notogaster; posterior notogastral tectum absent. Podotectum II present; circumpedal carina complete. Genital and anal plates separated by more than half length of genital plate; aggenital setae present; postanal porose area absent. Femora III and IV each with one porose area; solenidion σ present on genu III.

<==Haplozetidae [Protoribatidae, Xylobatidae] NB-P09
    |--Paraxylobates Balogh & Mahunka 1969 S04
    |    `--*P. imitans Balogh & Mahunka 1969 S04
    |--Conozetes Balogh & Mahunka 1969 S04
    |    `--*C. arcualis Balogh & Mahunka 1969 S04
    |--Bolkiah Mahunka 1997 S04
    |    `--*B. hauseri Mahunka 1997 S04
    |--Borneozetes Mahunka 1997 S04
    |    `--*B. lanceolatus Mahunka 1997 S04
    |--Cantharozetes Hammer 1961 S04
    |    `--*C. lucens Hammer 1961 S04
    |--Setincabates Lee 1993 S04
    |    `--*S. hypersetosus Lee 1993 S04
    |--Cribrozetes Balogh 1970 S04
    |    `--*C. multiareolatus Balogh 1970 S04
    |--Trixylobates Balogh & Mahunka 1978 S04
    |    `--*T. bidactylus Balogh & Mahunka 1978 S04
    |--Sicaxylobates Luxton 1985 S04
    |    `--*S. sicafer (Hammer 1968) [=Xylobates sicafer] S04
    |--Baloghiella Bulanova-Zachvatkina 1960 S04
    |    |--*B. prima Bulanova-Zachvatkina 1960 S04
    |    `--B. granulata Bayartogtokh & Akrami 2000 S04
    |--Baloghia Mahunka 1994 S04
    |    |--*B. juditae Mahunka 1994 S04
    |    `--B. spinifera Mahunka 1994 S04
    |--Trachyoribates Berlese 1908 S04
    |    |--*T. ampulla (Berlese 1905) [=Oribata ampulla] S04
    |    `--T. chinensis Fukuyama & Aoki 2000 S04
    |--Transoribates Pérez-Íñigo 1992 [incl. Lagenobates Weigmann & Miko 2002] S04
    |    |--T. lagenula (Berlese 1904) (see below for synonymy) S04
    |    `--T. agricola (Nakamura & Aoki 1989) [=Protoribates agricola] S04
    |--Protoribates Berlese 1908 NB-P09, S04
    |    |  i. s.: P. longipilis Sellnick 1931 S93
    |    |--P. (Protoribates) S04
    |    `--P. (Triaunguis) S04
    |--Berlesiella Hammer 1979 S04
    |    |--*B. (Berlesiella) scutata Hammer 1979 S04
    |    `--B. (Comororibula Mahunka 1994) S04
    |         `--B. (*C.) truncata (Mahunka 1994) [=*Comororibula truncata] S04
    |--Afroleius Mahunka 1984 S04
    |    |--*A. deformis Mahunka 1984 S04
    |    |--A. minor Mahunka 1984 S04
    |    `--A. simplex Mahunka 1984 S04
    |--Tuxenia Hammer 1958 S04
    |    |--*T. complicata Hammer 1958 S04
    |    |--T. brevis Covarrubias 1967 [incl. T. brevisetosa Mahunka 1980] S04
    |    `--T. manantialis Hammer 1962 S04
    |--Acutozetes Balogh 1970 S04
    |    |--*A. rostratus Balogh 1970 S04
    |    |--A. bornemisszai Balogh & Balogh 1986 S04
    |    |--A. chibai Aoki 1976 S04
    |    |--A. javensis Hammer 1979 S04
    |    `--A. nadchatrami Balogh & Mahunka 1974 S04
    |--Pilobates Balogh 1960 S04
    |    |--P. (Pilobates) S04
    |    |    |--*P. (P.) pilosellus (Balogh 1958) [=Protoribates pilosellus] S04
    |    |    `--P. (P.) carpentanus Pérez-Íñigo 1969 (see below for synonymy) S04
    |    `--P. (Italobates Mahunka 1994) S04
    |         `--P. (*I.) incisura (Berlese 1916) [=Protoribates (Scheloribates) incisura] S04
    |--Peloribates Berlese 1908 S04
    |    |  i. s.: P. antillensis (Mahunka 1985) S07
    |    |--P. (Peloribates) S04
    |    |--P. (Aokibates Mahunka 1988) S04
    |    |    `--P. (*A.) yoshii (Mahunka 1988) [=*Aokibates yoshii] S04
    |    `--P. (Tentaculozetes Balogh 1970) S04
    |         `--P. (*T.) loksai (Balogh 1970) [=*Tentaculozetes loksai] S04
    |--Pilobatella Balogh & Mahunka 1967 S04
    |    |--*P. punctulata Balogh & Mahunka 1967 S04
    |    |--P. berlesei Bhattacharya & Banerjee 1979 S04
    |    |--P. maurensis Scull 1985 S04
    |    |--P. monstruosa Mahunka 1986 S04
    |    |--P. pseudovermiseta Corpuz-Raros 1979 S04
    |    |--P. schauenbergi Mahunka 1977 S04
    |    `--P. xena Mahunka 1977 S04
    |--Vilhenabates Balogh 1963 S04
    |    |--V. (Vilhenabates) S04
    |    |    |--*V. (V.) minutus (Balogh 1958) [=Peloribates minutus] S04
    |    |    |--V. (V.) hyalinus Tseng 1984 S04
    |    |    `--V. (V.) simplex Balogh 1970 S04
    |    `--V. (Phalacrozetes Aoki 1965) S04
    |         |--V. (*P.) sinatus (Aoki 1965) [=*Phalacrozetes sinatus] S04
    |         `--V. (P.) similis (Mahunka 1988) [=Phalacrozetes similis] S04
    |--Setoxylobates Balogh & Mahunka 1967 S04
    |    |--*S. (Setoxylobates) foveolatus Balogh & Mahunka 1967 S04
    |    |--S. (Plenoxylobates Hammer 1979) S04
    |    |    |--S. (*P.) ramosus (Hammer 1979) [=*Plenoxylobates ramosus] S04
    |    |    |--S. (P.) curtiseta (Hammer 1979) [=Plenoxylobates curtiseta] S04
    |    |    `--S. (P.) mayuloeus (Corpuz-Raros 1979) [=Plenoxylobates mayuloeus] S04
    |    `--S. (Polyxylobates Hammer 1973) S04
    |         |--S. (*P.) diversiporosus (Hammer 1973) [=*Polyxylobates diversiporosus] S04
    |         `--S. (P.) principalis (Berlese 1916) [=Protoribates principalis] S04
    |--Lauritzenia Hammer 1958 S04
    |    |--L. (Lauritzenia) S04
    |    |    |--*L. (L.) longipluma Hammer 1958 S04
    |    |    |--L. (L.) furtadoi (Balogh & Mahunka 1974) [=Haplozetes furtadoi] S04
    |    |    |--L. (L.) glabra Pérez-Íñigo & Baggio 1996 S04
    |    |    |--L. (L.) insignis (Balogh & Mahunka 1966) [=Haplozetes insignis] S04
    |    |    |--L. (L.) loongchiensis (Tseng 1984) [=Haplozetes loongchiensis] S04
    |    |    `--L. (L.) minimicoma (Beck 1964) [=Haplozetes minimicoma] S04
    |    |--L. (Incabates) S04
    |    `--L. (Magnobates Hammer 1967) [incl. Gymnozetes Calugar & Vasiliu 1983] S04
    |         |--L. (*M.) flagellifer (Hammer 1967) [=*Magnobates flagellifer] S04
    |         `--L. (M.) orghidani (Calugar & Vasiliu 1983) S04 [=*Gymnozetes orghidani BB92a]
    |--Perxylobates Hammer 1972 [incl. Octodurozetes Mahunka 1993] S04
    |    |--*P. vermiseta (Balogh & Mahunka 1968) [=Xylobates vermiseta] S04
    |    |--P. barbatus Hammer 1972 S04
    |    |--P. berndhauseri (Mahunka 1993) [=Octodurozetes berndhauseri] S04
    |    |--P. brevisetus Mahunka 1988 S04
    |    |--P. coreanus Coi & Aoki 1993 S04
    |    |--P. guehoi Mahunka 1978 S04
    |    |--P. longissimus (Warburton 1912) [=Oribata longissima] S04
    |    |--P. mahunkai Bayoumi 1980 S04
    |    |--P. mayuloeus Corpuz-Raros 1979 C-R79
    |    |--P. obesus Bayoumi 1977 S04
    |    |--P. paravermiseta Mahunka 1976 S04
    |    |--P. sinensis Wen, Aoki & Wang 1984 S04
    |    |--P. sinlimes (Hammer 1971) C-R79
    |    |--P. taidinchani Mahunka 1976 S04
    |    `--P. vietnamensis (Jeleva & Vu 1987) [=Xylobates vietnamensis] S04
    |--Magyaria Balogh 1963 S04
    |    |--*M. reticulata (Balogh 1958) [=Scheloribates reticulatus] S04
    |    |--M. agusana Corpuz-Raros 1991 S04
    |    |--M. annobonica Pérez-Íñigo 1982 S04
    |    |--M. atlantica Pérez-Íñigo 1983 S04
    |    |--M. breviseta Mahunka 1978 BB92b
    |    |--M. cancellata (Beck 1964) [=Haplozetes cancellatus] S04
    |    |--M. fenestrata Balogh & Mahunka 1974 S04
    |    |--M. filipina Corpuz-Raros 1979 S04
    |    |--M. florida Mahunka 1985 S04
    |    |--M. incisa Balogh & Mahunka 1974 S04
    |    |--M. javensis Hammer 1979 S04
    |    |--M. luzonica Corpuz-Raros 1991 S04
    |    |--M. mindanensis Corpuz-Raros 1979 S04
    |    |--M. ornata Balogh 1963 S04
    |    |--M. pulcherrima Balogh 1970 S04
    |    |--M. strinovichi Balogh 1970 S04
    |    `--M. triungulata Mahunka 1988 S04
    |--Indoribates Jacot 1929 S04
    |    |--I. (Indoribates) [incl. Nixozetes Mahunka 1977, Sundazetes Hammer 1979] S04
    |    |    |--*I. (I.) punctulatus (Sellnick 1925) S04 (see below for synonymy)
    |    |    |--I. (I.) crassisetiger (Fukuyama & Aoki 2000) [=Nixozetes crassisetiger] S04
    |    |    |--I. (I.) gregoryi (Balogh 1970) S04 [=Cosmobates gregoryi S04, Nixozetes gregoryi BB92b]
    |    |    |--I. (I.) japonicus (Aoki 1988) [=Sundazetes japonicus] S04
    |    |    |--I. (I.) javanus (Mahunka 1977) S04 [=*Nixozetes javanus BB92a]
    |    |    |--I. (I.) multisetus (Wen & Zhao 1994) [=Sundazetes multisetus] S04
    |    |    |--I. (I.) nobilis (Golosova 1984) [=Cosmobates nobilis] S04
    |    |    `--I. (I.) panabokkei (Balogh 1970) (see below for synonymy) S04
    |    |--I. (Haplozetes) S04
    |    |--I. (Mancoribates Hammer 1961) [incl. Minasbates Pérez-Íñigo & Baggio 1996] S04
    |    |    |--I. (*M.) rostropilosus (Hammer 1961) [=*Mancoribates rostropilosus] S04
    |    |    |--I. (M.) acutirostrum (Hammer 1968) [=Lauritzenia acutirostrum] S04
    |    |    |--I. (M.) rotundirostrum (Hammer 1968) [=Lauritzenia rotundirostrum] S04
    |    |    `--I. (M.) sphaeroclava (Pérez-Íñigo & Baggio 1996) [=Minasbates sphaeroclava] S04
    |    |--I. (Philippizetes Corpuz-Raros 1979) S04
    |    |    |--I. (*P.) philippinensis (Corpuz-Raros 1979) [=Nixozetes (*P.) philippinensis] S04
    |    |    `--I. (P.) corpusrarosae (Mahunka 1987) [=Nixozetes (P.) corpusrarosae] S04
    |    `--I. (Triungulozetes Subías 2001) [incl. Mixobates Gil-Martín & Subías 1993 non Thor 1905] S04
    |         |--I. (*T.) triangulatus (Beck 1964) [=Haplozetes triangulatus] S04
    |         `--I. (T.) fusifer (Berlese 1908) (see below for synonymy) S04
    `--Rostrozetes Sellnick 1925 NB-P09, S04 [incl. Carabozetes Mihelčič 1957 S04, Zaherizetes Yousef & Nasr 1976 S04]
         |--R. ovulum (Berlese 1908) [=Trachyoribates (Rostrozetes) ovulum) S04
         |    |--R. o. ovulum S04 (see below for synonymy)
         |    |--R. o. appalachicola Jacot 1938 (see below for synonymy) S04
         |    `--R. o. poensis (Mihelčič 1957) [=Carabozetes poensis, Trachyoribates (Rostrozetes) ovulum poensis] S04
         |--R. aegypticus (Yousef & Nasr 1976) [=Zaherizetes aegypticus, Trachyoribates (Rostrozetes) aegypticus] S04
         |--R. africanus (Yousef & Nasr 1976) [=Zaherizetes africanus, Trachyoribates (Rostrozetes) africanus] S04
         |--R. angulifer Balogh & Mahunka 1979 [=Trachyoribates (Rostrozetes) angulifer] S04
         |--R. bothulifer Balogh & Mahunka 1979 [=Trachyoribates (Rostrozetes) bothulifer] S04
         |--R. carinatus Beck 1965 [=Trachyoribates (Rostrozetes) carinatus] S04
         |--R. cristatus Balogh & Mahunka 1969 [=Trachyoribates (Rostrozetes) cristatus] S04
         |--R. cubanus Balogh & Mahunka 1974 [=Trachyoribates (Rostrozetes) cubanus] S04
         |--R. dimorphichaites Higgins 1996 [=Trachyoribates (Rostrozetes) dimorphichaites] S04
         |--R. dispar (Balogh 1958) [=Peloribates dispar, Trachyoribates (Rostrozetes) dispar] S04
         |--R. florens Balogh 1970 [=Trachyoribates (Rostrozetes) florens] S04
         |--R. geminosetosus Balogh & Mahunka 1978 [=Trachyoribates (Rostrozetes) geminosetosus] S04
         |--R. glaber Beck 1965 [=Trachyoribates (Rostrozetes) glaber] S04
         |--R. heterotrichus Pérez-Íñigo & Baggio 1991 [=Trachyoribates (Rostrozetes) heterotrichus] S04
         |--R. inornatus Pérez-Íñigo & Baggio 1991 [=Trachyoribates (Rostrozetes) inornatus] S04
         |--R. irregularis Balogh & Mahunka 1969 [=Trachyoribates (Rostrozetes) irregularis] S04
         |--R. komodensis Mahunka 1977 [=Trachyoribates (Rostrozetes) komodensis] S04
         |--R. mammillatus (Warburton 1912) [=Oribata mammillata, Trachyoribates (Rostrozetes) mamillatus] S04
         |--R. monstruosus Balogh & Mahunka 1969 [=Trachyoribates (Rostrozetes) monstruosus] S04
         |--R. nagaii Mahunka 1987 [=Trachyoribates (Rostrozetes) nagaii] S04
         |--R. nebulosus Beck 1965 PL17 (see below for synonymy)
         |--R. perezinigoi Balogh 1995 [=Trachyoribates (Rostrozetes) perezinigoi] S04
         |--R. pinguis Balogh & Mahunka 1978 [=Trachyoribates (Rostrozetes) pinguis] S04
         |--R. polygonatus Balogh & Mahunka 1969 [=Trachyoribates (Rostrozetes) polygonatus] S04
         |--R. pseudofurcatus Balogh & Mahunka 1968 [=Trachyoribates (Rostrozetes) pseudofurcatus] S04
         |--R. punctatus Karppinen 1966 [=Trachyoribates (Rostrozetes) punctatus] S04
         |--R. rimachensis Beck 1965 [=Trachyoribates (Rostrozetes) rimachensis] S04
         |--R. schalleri Beck 1965 [=Trachyoribates (Rostrozetes) schalleri] S04
         |--R. sexclavatus Sellnick 1925 [=Trachyoribates (Rostrozetes) sexclavatus] S04
         `--R. shibai Aoki 1976 [=Trachyoribates (Rostrozetes) shibai] S04

Indoribates (Indoribates) panabokkei (Balogh 1970) [=Cosmobates panabokkei; incl Nixozetes apoensis Corpuz-Raros 1979] S04

*Indoribates (Indoribates) punctulatus (Sellnick 1925) S04 [=Protoribates punctulatus S04; incl. Sundazetes crispus Hammer 1979 S04, Indoribates crispus BB92b]

Indoribates (Triungulozetes) fusifer (Berlese 1908) [=Protoribates (Scheloribates) fusifer; incl. Haplozetes (Mixobates) insularis Pérez-Íñigo & Peña 1996] S04

Pilobates (Pilobates) carpentanus Pérez-Íñigo 1969 [incl. Protoribates elongatus Mihelčič 1956 (n. d.), Pr. longior Mihelčič 1957 (n. d.) non Berlese 1908] S04

Rostrozetes nebulosus Beck 1965 PL17 [=Rostrozetes foveolatus nebulosus S04; incl. R. nebulosus tropica Beck 1963 (n. n.) B65]

Rostrozetes ovulum appalachicola Jacot 1938 [=R. foveolatus appalachicola, Trachyoribates (Rostrozetes) ovulum appalachicola] S04

Rostrozetes ovulum ovulum (Berlese 1908) S04 [incl. Peloribates areolatus Balogh 1958 S04, Trachyoribates areolatus B65, T. dorsalis Balogh 1958 S04, Rostrozetes flavus Woodring 1965 S04, *R. foveolatus Sellnick 1925 S04, R. geneuxi Mahunka 1985 S04, Trachyoribates nodosus Hammer 1958 S04, R. nodosus B65, R. pulcherrimus Balogh 1961 S04, Peloribates punctulatus Balogh 1958 S04, Trachyoribates punctulatus B65, R. punctulifer Balogh & Mahunka 1979 S04, R. trimorphus Balogh & Mahunka 1979 S04]

Transoribates lagenula (Berlese 1904) [=Oribata lagenula; incl. Protoribates latus Mihelčič 1965, *Transoribates latus, P. pupula Berlese 1910] S04

*Type species of generic name indicated

REFERENCES

[BB92a] Balogh, J., & P. Balogh. 1992a. The Oribatid Mites Genera of the World vol. 1. Hungarian Natural History Museum: Budapest.

[BB92b] Balogh, J., & P. Balogh. 1992b. The Oribatid Mites Genera of the World vol. 2. Plates. Hungarian Natural History Museum: Budapest.

[B65] Beck, L. 1965. Über Variabilität und Wertigkeit morphologischer Merkmale bei adulten Oribatiden (Arachnida, Acari) am Beispiel der Gattung Rostrozetes Sellnick 1925. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft 508: 1–64.

[C-R79] Corpuz-Raros, L. A. 1979. Philippine Oribatei (Acarina). I. Preliminary list of species and descriptions of forty new species. Philippine Agriculturist 62 (1): 1–82.

[NB-P09] Norton, R. A., & V. M. Behan-Pelletier. 2009. Suborder Oribatida. In: Krantz, G. W., & D. E. Walter (eds) A Manual of Acarology 3^rd ed. pp. 430–564. Texas Tech University Press.

[PL17] Pachl, P., A. C. Lindl, A. Krause, S. Scheu, I. Schaefer & M. Maraun. 2017. The tropics as an ancient cradle of oribatid mite diversity. Acarologia 57 (2): 309–322.

[S07] Schatz, H. 2007. Biogeography of oribatid mites (Acari, Oribatida) from the Cordillera de Talamanca, Costa Rica and Panama. In: Morales-Malacara, J. B., V. M. Behan-Pelletier, E. Ueckermann, T. M. Pérez, E. G. Estrada-Venegas & M. Badii (eds) Acarology XI: Proceedings of the International Congress pp. 151–167. Instituto de Biología and Faculdad de Ciencias, Universidad Nacional Autónoma de México, Sociedad Latinoamericana de Acarología: México.

[S93] Selden, P. A. 1993. Arthropoda (Aglaspidida, Pycnogonida and Chelicerata). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 297–320. Chapman & Hall: London.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

Last updated: 24 February 2019.