Belongs within: Neohydrachnidia.
The Anisitsiellidae are a diverse group of freshwater mites of questionable monophyly, larvae of which are (where known) parasites of nematocerous Diptera (Walter et al. 2009).
Characters (from Walter et al. 2009): Adults with claws either present or absent on legs IV; when claws present, then lateral eyes usually located on platelets associated with ventral shield; venter with suture lines between coxal plates II-III complete; genital field with movable genital flaps flanking gonopore; genital acetabula conspicuous, lying free in gonopore.
<==Anisitsiellidae
| i. s.: Fuenticola Goldschmidt 2004 G04
| `--*F. sancheziana Goldschmidt 2004 G04
| Mamersopsides Viets 1916 G04
| |--*M. sigthori Viets 1916 G04
| |--M. neotropica Goldschmidt 2004 G04
| `--M. vietsi Cook 1966 G04
| Ranautonia dentipalpis Viets 1935 G04
|--Nilotoniinae G04
| |--Mamersonia Viets 1954 G04
| | |--*M. amazonica Viets 1954 G04 [=Nilotonia (*Mamersonia) amazonica RFF05]
| | |--M. biscutata Bader 1995 G04
| | |--M. cascada Goldschmidt 2004 G04
| | |--M. monoscutata Bader 1995 G04
| | |--M. scutata Cook 1966 G04
| | |--M. similis Bader 1995 G04
| | `--M. testudinata Cook 1966 G04
| `--Nilotonia Thor 1905 G04
| | i. s.: N. fundamentalis Bader 1995 G04
| | N. gallica Motas & Angelier 1967 G04
| | N. longipora WP99
| | N. ortali Bader 1995 G04
| | N. parva Walter 1931 G04
| | N. petri Cook 1979 G04
| | N. pontifica Gerecke 1992 G04
| |--*N. (Nilotonia) loricata (Nordenskiöld 1905) [=Teutonia loricata] G04
| |--N. (Bolivartonia Orghidan & Gruia 1983) G04
| | |--N. (*B.) simoni Orgjhidan & Gruia 1983 G04
| | |--N. (B.) conjuncta Orghidan & Gruia 1983 S86
| | |--N. (B.) curtipalpis Goldschmidt 2004 G04
| | `--N. (B.) panzuda Goldschmidt 2004 G04
| |--N. (Dartia Soar 1917) G04
| |--N. (Guanacastonia Goldschmidt 2004) G04
| | `--N. (*G.) pablito Goldschmidt 2004 G04
| `--N. (Manotonia Viets 1935) G04
`--Anisitsiellinae [Mamersopsidae, Mamersopsinae] G04
|--Mamersopsis Nordenskiöld 1905 G04
|--Platymamersopsis Viets 1914 G04
|--Anisitsiella Daday 1905 G04
| `--A. aculeata Daday 1905 RFF05
|--Sigthoriella Besch 1964 G04
| `--S. hygropetrica Besch 1964 RFF05
|--Psammotorrenticola Angelier 1951 [Psammotorrenticolinae] S86
| `--P. gracilis Angelier 1951 S86
|--Utaxatax Habeeb 1964 S86
| `--U. ovalis Cook 1974 S86
|--Mamersella Viets 1929 G04
| `--M. newelli Habeeb 1959 (n. d.) C91
|--Stygomamersopsis Cook 1974 S86
| |--S. anisitisipalpis Cook 1974 S86
| `--S. viedmai Cook 1974 S86
|--Mamersellides Lundblad 1937 G04
| |--*M. ventriperforatus Lundblad 1937 G04
| `--M. costenius Cramer & Smith 1993 G04
|--Anisitsiellides Lundblad 1941 G04
| |--A. australis Smit 2002 RFF05
| |--A. chilensis Cook 1988 RFF05
| |--A. lundbladi Cook 1980 RFF05
| `--A. monticolus Lundblad 1941 RFF05
|--Rutacarus Lundblad 1937 G04
| |--R. (Rutacarus) S86
| | |--R. (R.) feradasae Cook 1980 S86
| | `--R. (R.) pyriformis Lundblad 1937 RFF05
| `--R. (Neorutacarus Orghidan & Gruia 1983) S86
| `--R. (N.) angelieri Orghidan & Gruia 1983 RFF05
|--Bandakiopsis fonticola WL09
|--Cookacarus WL09
|--Oregonacarus WL09
|--Australiotonia WL09
|--Rubicundula placibilis WL09
`--Bandakia Thor 1913 G04
|--*B. concreta Thor 1913 G04
|--B. borealis Smith 1979 S86
|--B. corsica Angelier 1951 G04
|--B. elhacha Goldschmidt 2004 G04
|--B. elongata Cook 1963 [incl. B. longipalpis Cook 1974] S86
|--B. fragilis WL09
|--B. hoffmannae Cramer & Smith 1991 G04
|--B. mexicana Cramer & Smith 1991 G04
|--B. oregonensis Smith 1979 S86
|--B. phreatica Cook 1974 S86
`--B. speciosa Viets 1953 G04
*Type species of generic name indicated
REFERENCES
[C91] Conroy, J. C. 1991. Acarology—a modern science. In Dusbábek, F. & V. Bukva (eds) Modern Acarology: Proceedings of the VIII International Congress of Acarology, held in České Budĕjovice, Czechoslovakia, 6–11 August 1990 vol. 1 pp. 5-16. SPB Academic Publishing: The Hague.
[G04] Goldschmidt, T. 2004. Untersuchungen über Wassermilben der Familie Anisitsiellidae Koenike 1910 aus der Neotropis (Arachnida, Acari, Actinedida, Hydrachnidia). Senckenbergiana Biologica 83 (2): 113-150.
[RFF05] Rosso de Ferradás, B. & H. R. Fernández. 2005. Elenco y biogeografía de los ácaros acuáticos (Acari, Parasitengona, Hydrachnidia) de Sudamérica. Graellsia 61 (2): 181-224.
[S86] Schwoerbel, J. 1986. Acari: “Hydrachnellae”. In Stygofauna Mundi: A Faunistic, Distributional, and Ecological Synthesis of the World Fauna inhabiting Subterranean Waters (including the Marine Interstitial) (L. Botosaneanu, ed.) pp. 652-696. E. J. Brill / Dr. W. Backhuys: Leiden.
[WL09] Walter, D. E., E. E. Lindquist, I. M. Smith, D. R. Cook & G. W. Krantz. 2009. Order Trombidiformes. In: Krantz, G. W. & D. E. Walter (eds) A Manual of Acarology, pp. 233-420. Texas Tech University Press.
[WP99] Walter, D. E. & H. C. Proctor. 1999. Mites: Ecology, Evolution and Behaviour. CABI Publishing: Wallingford (UK).
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