Umbilicaria

Umbilicaria leiocarpa, copyright Jason Hollinger.

Belongs within: Lecanoromycetes.
Contains: Umbilicaria section Glabrae, Umbilicaria section Polymorphae, Umbilicaria secton Velleae.

Umbilicaria, rock tripe, is a cosmopolitan genus of saxicolous lichens that derive their name from their attachment to the substrate by a distinct 'navel'.

Characters (from Frey 1933): Thallus foliaceous, navel simple, thin or often divided into several strands, with the base very firmly attached, upper surface smooth or cracked-areolate, often with scaly or sheet-like attachments or protruding ribs. Apothecia sessile on upper surface, occasionally slightly sunk or pressed, often top-shaped, short-stalked, black. Underside of the thallus smooth and hairless or cracked-areolate, with or without rhizines. Rhizines simple or branched or warty, also often reticulate or pitted and with anastomosing discontinuous ribs. More or less radially arranged bars often extending outward from navel. Thallus layered, upper crust usually solid (exceptionally thin), forming a palisade plectenchyma, paraplectenchyma or an almost amorphous-appearing tissue. Gonidia Cytococcus, usually in a continuous or more or less regularly interrupted zone. Hyphae of gonidial zone loose, rarely fused like the rind. Central mass usually loose, rarely dense; lower crust usually similar to top layer, rhizines mostly similarly textured to underside of thallus. Apothecia always without gonidia, often quite black inside or at least partly dark, hypothecium usually at least brownish, hymenium often with a tendency to split. Spores unicellular and hyaline, or 2- to multicellular (forming a brick-like arrangement), hyaline or partially brown. Pycnidia sunk in thallus, but alwayswith a clear, point-like, black crown, pitcher-shaped or pear-shaped, often quite strongly broadened at the base.

Umbilicaria Hoffmann ex Linnaeus 1891 (see below for synonymy) F33
    |--U. subg. Gyrophora Endlich 1836 F33
    |    |--U. sect. Agyrophora Nylander 1878 (see below for synonymy) F33
    |    |    |--U. cinerascens (Arnold) Frey 1931 (see below for synonymy) F33
    |    |    |    |--U. c. f. cinerascens F33
    |    |    |    `--U. c. f. laciniata Frey 1933 non Lichen corneus var. laciniatus Retzius 1779 F33
    |    |    |--U. laevis (Schaerer) Frey 1931 (see below for synonymy) F33
    |    |    |--U. leiocarpa (de Candolle in de Lamarck & de Candolle 1805) Frey 1931 (see below for synonymy) F33
    |    |    |--U. microphylla (Laurer) Massal. 1852 (see below for synonymy) F33
    |    |    |--U. reticulata F33 (see below for synonymy)
    |    |    |    |--U. r. f. reticulata F33
    |    |    |    |--U. r. f. maxima F33
    |    |    |    |--U. r. f. perforata F33
    |    |    |    `--U. r. f. typica F33
    |    |    |--U. rigida (Du Rietz) Frey 1931 [=Gyrophora rigida Du Rietz 1925] F33
    |    |    |    |--U. r. var. rigida F33
    |    |    |    `--U. r. var. discissa F33
    |    |    `--U. subglabra  (Nylander) Harm. 1910 [=Gyrophora subglabra Nylander 1896] F33
    |    |         |--U. s. var. subglabra F33
    |    |         |--U. s. var. pallens (Nylander) Frey 1933 (see below for synonymy) F33
    |    |         `--U. s. var. schmidtii Frey 1933 F33
    |    |--U. sect. Glabrae F33
    |    |--U. sect. Polymorphae F33
    |    `--U. sect. Velleae F33
    `--U. subg. Gyrophoropsis (Elenkin & Sav.) Zahlbr. 1927 [=Gyrophoropsis Elenkin & Sav. 1911] F33
         |--U. cinereorufescens (Schaerer) Frey 1931 (see below for synonymy) F33
         |--U. crustulosa (Acharius) Frey 1931 (see below for synonymy) F33
         |    |--U. c. var. crustulosa F33
         |    `--U. c. var. badiofusca Frey 1933 F33
         |--U. depressa (Acharius) Duby 1830 [=Gyrophora crustulosa var. depressa Acharius 1810] F33
         |--U. ruebeliana (DR. & Frey) Frey 1931 [=Gyrophora ruebeliana DR. & Frey 1929] F33
         |    |--U. r. f. ruebeliana F33
         |    `--U. r. f. rhizinosa Frey 1933 F33
         `--U. spodochroa (Acharius) Frey 1931 (see below for synonymy) F33
              |--U. s. var. spodochroa F33
              `--U. s. var. lapponica Frey 1933 F33

Umbilicaria incertae sedis:
  U. angulata F33
  U. calvescens F33
  U. dichroa F33
  U. esculenta F33
  U. haplocarpa F33
  U. krempelhuberi F33
  U. mammulata JK06
  U. membranacea F33
  U. muehlenbergii F33
  U. nanella Frey & Poelt 1977 S03
  U. pennsylvanica F33
  U. phaea F33
  U. semitensis F33
  U. yunnana F33

Umbilicaria Hoffmann ex Linnaeus 1891 [incl. Gyrophora Acharius 1803; Gyrophoraceae, Umbilicariaceae, Umbilicariales] F33

Umbilicaria sect. Agyrophora Nylander 1878 [=Gyrophora sect. Agyrophora (Nylander) Zahlbr. 1906; incl. U. sect. Anthracinae Frey 1931] F33

Umbilicaria cinerascens (Arnold) Frey 1931 [=Gyrophora cinerascens Arnold 1875, U. polyphylla var. cinerascens (Arnold) Wain. 1899] F33

Umbilicaria laevis (Schaerer) Frey 1931 [=Gyrophora atropruniosa var. laevis Schaerer 1848, G. laevis (Schaerer) Du Rietz 1925, Umbilicaria anthracina var. laevis (Schaerer) Schaerer 1850; incl. U. anthracina var. laevis f. nuda Schaere 1850] F33

Umbilicaria leiocarpa (de Candolle in de Lamarck & de Candolle 1805) Frey 1931 [=Gyrophora leiocarpa (de Candolle) Du Rietz 1925; incl. G. anthracina Körb. 1855, Umbilicaria atropruinosa Schaerer 1823, G. atropruinosa var. tessellata Schaerer 1818, U. anthracina var. tessellata (Schaerer) Schaerer 1850, U. atropruinosa var. tessellata (Schaerer) Fries 1831] F33

Umbilicaria microphylla (Laurer) Massal. 1852 [=U. atropruinosa var. microphylla Laurer 1832, Gyrophora anthracina var. microphylla (Laurer) Jatta 1911, G. microphylla (Laurer) Arnold 1878, Umbilicaria anthracina var. microphylla (Laurer) Schaerer 1850, U. cinerascens var. microphylla (Laurer) Stizenb. 1882; incl. Gyrophora microphylloides Zahlbr. 1927] F33

Umbilicaria reticulata F33 [=Gyrophora polymorpha var. reticulata Schaerer 1818 F33, G. anthracina var. reticulata (Schaerer) Rabenhorst 1859 F33, G. reticulata (Schaerer) Fries 1871 F33, Lecidea atropruinosa var. reticulata (Schaerer) Schaerer 1828 F33, Umbilicaria anthracina var. reticulata (Schaerer) Tuck. 1848 F33, U. atropruinosa var. reticulata (Schaerer) Fries 1831 F33, U. tessellata var. reticulata (Schaerer) Duby 1830 F33; incl. Lichen decussatus Vill. 1789 F33, Gyrophora decussata (Vill.) Zahlbr. 1927 F33, Umbilicaria decussata C74, G. discolor Fries 1867 F33, U. ptychophora Nylander 1869 F33]

Umbilicaria subglabra var. pallens (Nylander) Frey 1933 [=U. atropruinosa var. pallens Nylander 1864, U. cinerascens var. pallens (Nylander) Nylander 1869] F33

*Type species of generic name indicated

REFERENCES

[C74] Coineau, Y. 1974. Éléments pour une monographie morphologique, écologique et biologique des Caeculidae (Acariens). Mémoires du Muséum National d'Histoire Naturelle, nouvelle série, Série A, Zoologie 81: 1–299, 224 pls.

[F33] Frey, E. 1933. Cladoniaceae (unter Ausschluss der Gattung Cladonia), Umbilicariaceae. In: Die Flechten (Dr. L. Rabenhorst’s Kryptogamenflora von Deutschland, Österreich und der Schweiz vol. 9 sect. 4 pt 1. Akademische Verlagsgesellschaft M. B. H.: Leipzig (reprinted 1971. Johnson Reprint Corporation: New York).

[JK06] James, T. Y., F. Kauff, C. L. Schoch, P. B. Matheny, V. Hofstetter, C. J. Cox, G. Celio, C. Gueidan, E. Fraker, J. Miadlikowska, H. T. Lumbsch, A. Rauhut, V. Reeb, A. E. Arnold, A. Amtoft, J. E. Stajich, K. Hosaka, G.-H. Sung, D. Johnson, B. O'Rourke, M. Crockett, M. Binder, J. M. Curtis, J. C. Slot, Z. Wang, A. W. Wilson, A. Schüßler, J. E. Longcore, K. O'Donnell, S. Mozley-Standridge, D. Porter, P. M. Letcher, M. J. Powell, J. W. Taylor, M. M. White, G. W. Griffith, D. R. Davies, R. A. Humber, J. B. Morton, J. Sugiyama, A. Y. Rossman, J. D. Rogers, D. H. Pfister, D. Hewitt, K. Hansen, S. Hambleton, R. A. Shoemaker, J. Kohlmeyer, B. Volkmann-Kohlmeyer, R. A. Spotts, M. Serdani, P. W. Crous, K. W. Hughes, K. Matsuura, E. Langer, G. Langer, W. A. Untereiner, R. Lücking, B. Büdel, D. M. Geiser, A. Aptroot, P. Diederich, I. Schmitt, M. Schultz, R. Yahr, D. S. Hibbett, F. Lutzoni, D. J. McLaughlin, J. W. Spatafora & R. Vilgalys. 2006. Reconstructing the early evolution of Fungi using a six-gene phylogeny. Nature 443: 818–822.

[S03] Sinha, G. P. 2003. Additions to the lichen flora of India from Sikkim. Bulletin of the Botanical Survey of India 45 (1–4): 221–224.

Parmelia

Parmelia sulcata, copyright James Lindsey.

Belongs within: Parmeliaceae.

Parmelia is a cosmopolitan genus of foliose lichens, light-coloured above and black below with the lower surface anchored to the substrate by rhizenes, and often with squarish ends to the thallus lobes.

Parmelia
    |--P. caperata H04
    |--P. encausta F33
    |--P. glomellifera F33
    |--P. intestiniformis C74
    |--P. isidiotyla F33
    |--P. molliuscula F33
    |--P. mussooriensis BB05
    |--P. olivacea H04
    |--P. omphalodes E99
    |    |--P. o. var. omphalodes E99
    |    `--P. o. var. discordans E99
    |--P. pubescens F33
    |--P. reticulata H49
    |--P. saxatilis EB03
    |--P. sorediata F33
    |--P. stellaris H04
    |--P. stenophylla C74
    |--P. stygia C474
    |    |--P. s. var. stygia C74
    |    `--P. s. var. septentrionalis C74
    `--P. sulcata M08

*Type species of generic name indicated

REFERENCES

[BB05] Banik, D., & P. Bujarbarua. 2005. Lichens on Agapetes D. Don ex G. Don (Ericaceae) in India. Bulletin of the Botanical Survey of India 47 (1–4): 59–62.

[C74] Coineau, Y. 1974. Éléments pour une monographie morphologique, écologique et biologique des Caeculidae (Acariens). Mémoires du Muséum National d'Histoire Naturelle, nouvelle série, Série A, Zoologie 81: 1–299, 224 pls.

[E99] Eriksson, O. E. (ed.) 1999. Notes on ascomycete systematics. Nos 2440–2755. Myconet 2: 1–41.

[EB03] Eriksson, O. E., H. O. Barah, R. S. Currah, K. Hansen, C. P. Kurtzman, T. Laessøe & G. Rambold (eds.) 2003. Notes on ascomycete systematics. Nos 3580–3623. Myconet 9: 91–103.

[F33] Frey, E. 1933. Cladoniaceae (unter Ausschluss der Gattung Cladonia), Umbilicariaceae. In: Die Flechten (Dr. L. Rabenhorst’s Kryptogamenflora von Deutschland, Österreich und der Schweiz vol. 9 sect. 4 pt 1. Akademische Verlagsgesellschaft M. B. H.: Leipzig (reprinted 1971. Johnson Reprint Corporation: New York).

[H04] Haeckel, E. 1899–1904. Kunstformen der Natur. Bibliographisches Institut: Leipzig und Wien.

[H49] Hodgson, E. A. 1949. New Zealand Hepaticae (liverworts). VI. A review of the New Zealand species of the genus Frullania. Transactions and Proceedings of the Royal Society of New Zealand 77 (3): 361–389.

[M08] Marstaller, R. 2008. Moosgesellschaften am Südrand des Kyffhäusergebirges bei Bad Frankenhausen (Kyffhäuserkreis). 130. Beitrag zur Moosvegetation Thüringens. Mauritiana 20 (2): 289–348.

Umbilicaria section Glabrae

Petalled rocktripe Umbilicaria polyphylla, from here.

Belongs within: Umbilicariaceae.

The section Glabrae of the lichen genus Umbilicaria is characterised by a lobed or sinuate-edged thallus that lacks rhizines (Frey 1933).

Characters (from Frey 1933): Underside of thallus without rhizines. Thallus usually brown or greenish gray or brown-black above, black or reddish-gray or whitish below, edge usually somewhat roundly lobed or sinuate. Apothecia with grooved disc. Spores colourless.

Umbilicaria sect. Glabrae Frey 1931 F33
    |--U. arctica (Acharius) Nylander 1859 [=Gyrophora arctica Acharius 1803] F33
    |    |--U. a. var. arctica F33
    |    |--U. a. var. hyporrhiza Nylander 1866 F33
    |    `--U. a. var. plicata Nylander 1866 F33
    |--U. corrugata (de Candolle) Nylander 1861 (see below for synonymy) F33
    |    |--U. c. f. corrugata F33
    |    `--U. c. f. microphyllina (Nylander) Frey 1933 [=U. corrugata var. microphyllina Nylander 1860] F33
    |--U. deusta (Linnaeus) Baumgartner 1790 (see below for synonymy) F33
    |    |--U. d. f. deusta F33
    |    |--U. d. f. brotera (Acharius) Frey 1933 (see below for synonymy) F33
    |    `--U. d. f. subpapulosa (Nylander) Frey 1933 (see below for synonymy) F33
    |--U. erosa (Weber) Acharius 1794 (see below for synonymy) F33
    |    |--U. e. var. erosa F33
    |    |--U. e. var. subradians Nylander 1861 [=Gyrophora erosa var. subradians (Nylander) Fries 1871] F33
    |    `--U. e. var. torrefacta (Lightf.) Frey 1933 (see below for synonymy) F33
    |--U. hyperborea (Acharius) Hoffmann 1801 (see below for synonymy) F33
    `--U. polyphylla (Linnaeus) Hoffmann 1796 (see below for synonymy) F33
         |  i. s.: U. p. f. congregata (Turner) Leight. 1871 (see below for synonymy) F33
         |         U. p. f. coriacea (Fries) Frey 1933 [=Gyrophora polyphylla var. glabra f. coriacea Fries 1871] F33
         |         U. p. f. diffusa (Nylander) Frey 1933 (see below for synonymy) F33
         |         U. p. f. lacera (Hepp) Leight. 1871 [=Gyrophora polyphylla f. lacera Hepp 1860] F33
         |--U. p. var. polyphylla F33
         `--U. p. var. variegata Nylander 1861 [=Gyrophora polyphylla var. variegata (Nylander) Fries 1871] F33

Umbilicaria corrugata (de Candolle) Nylander 1861 [=U. proboscidea var. corrugata de Candolle in de Lamarck & de Candolle 1805, Gyrophora corrugata (de Candolle) Arnold 1875, G. deusta var. corrugata (de Candolle) Turner 1839, G. glabra var. corrugata (de Candolle) Acharius 1814, G. heteroidea var. corrugata (de Candolle) Acharius 1810, G. hyperborea var. corrugata (de Candolle) Fries 1871, G. proboscidea var. corrugata (de Candolle) Mudd 1861, Umbilicaria varia var. deusta f. corrugata (de Candolle) Leight. 1856; incl. U. polyphylloides Nylander 1860] F33

Umbilicaria deusta (Linnaeus) Baumgartner 1790 [=Lichen deustus Linnaeus 1753, Gyrophora deusta (Linnaeus) Acharius 1803, G. polyphylla var. deusta (Linnaeus) Rabenhorst 1845, Umbilicaria polyphylla var. deusta (Linnaeus) Fries 1831; incl. Lichen flocculosus Wulf. ex Jacquin 1789, Gyrophora aenea var. flocculosa (Wulf.) Schaerer 1818, G. deusta var. flocculosa (Wulf.) Acharius 1803, G. flocculosa (Wulf.) Körb. 1855, Umbilicaria aenea var. flocculosa (Wulf.) Schaerer 1836, U. flocculosa (Wulf.) Hoffmann 1796, U. polyphylla var. flocculosa (Wulf.) Schaerer 1850] F33

Umbilicaria deusta f. brotera (Acharius) Frey 1933 [=Gyrophora deusta var. brotera Acharius 1803, U. flocculosa f. brotera (Acharius) Nylander 1861; incl. G. flocculosa f. cribrata Mueller 1862, G. deusta f. cribrata (Mueller) Zahlbr. 1927] F33

Umbilicaria deusta f. subpapulosa (Nylander) Frey 1933 [=U. flocculosa f. subpapulosa Nylander 1861, Gyrophora deusta f. subpapulosa (Nylander) Zahlbr. 1927, G. polyphylla var. deusta f. subpapulosa (Nylander) Fries 1871; incl. Umbilicaria flocculosa f. squamulosa Harm. 1910] F33

Umbilicaria erosa (Weber) Acharius 1794 [=Lichen erosus Weber 1778, Gyrophora erosa (Weber) Acharius) 1803; incl. Gyrophora koldeweyi Körb. 1874, Lichen reticulatus Retz 1779] F33

Umbilicaria erosa var. torrefacta (Lightf.) Frey 1933 [=Lichen torrefactus Lightf. 1777, Gyrophora erosa var. torrefacta (Lightf.) Fries 1871, G. torrefacta (Lightf.) Cromb. 1894, Umbilicaria torrefacta (Lightf.) Schrader 1794; incl. G. erosa var. torrida Acharius 1804, G. torrida (Acharius) Röhling 1813, Umbilicaria erosa var. torrida (Acharius) Nylander 1861, U. torrida (Acharius) Nylander ex Stizenb. 1876] F33

Umbilicaria hyperborea (Acharius) Hoffmann 1801 [=Lichen hyperboreus Acharius 1794, Gyrophora aenea var. hyperborea (Acharius) Schaerer 1818, G. hyperborea (Acharius) Acharius 1803, Umbilicaria aenea var. hyperborea (Acharius) Schaerer 1826, U. polyphylla var. hyperborea (Acharius) Schaerer 1850, U. varia var. hyperborea (Acharius) Leight. 1856; incl. Gyrophora hyperborea var. pimaria Fries 1871, U. ustulata Wainio 1888, G. ustulata (Wainio) Dalla Torre & Sarnth. 1902] F33

Umbilicaria polyphylla (Linnaeus) Hoffmann 1796 [=Lichen polyphyllus Linnaeus 1753, Gyrophora polyphylla (Linnaeus) Funck 1804; incl. G. glabra Acharius 1803, G. aenea var. glabra (Acharius) Schaerer 1826, G. polyphylla f. glabra (Acharius) Zahlbr. 1927, G. polyphylla var. glabra (Acharius) Fries 1871] F33

Umbilicaria polyphylla f. congregata (Turner) Leight. 1871 [=Gyrophora polyphylla f. congregata Turner 1839; incl. G. polyphylla var. conglobata Fries 1871, G. deusta var. conglobata (Fries) Zahlbr. 1927] F33

Umbilicaria polyphylla f. diffusa (Nylander) Frey 1933 [=Gyrophora glabra var. diffusa Nylander 1896, G. polyphylla f. diffusa (Nylander) Zahlbr. 1927] F33

*Type species of generic name indicated

REFERENCES

[F33] Frey, E. 1933. Cladoniaceae (unter Ausschluss der Gattung Cladonia), Umbilicariaceae. In Die Flechten (Dr. L. Rabenhorst’s Kryptogamenflora von Deutschland, Österreich und der Schweiz vol. 9) sect. 4, part 1. Akademische Verlagsgesellschaft M. B. H.: Leipzig (reprinted 1971. Johnson Reprint Corporation: New York).

Umbilicaria section Polymorphae

Umbilicaria cylindrica, from here.

Belongs within: Umbilicariaceae.
The section Polymorphae in the lichen genus Umbilicaria is characterised by a more or less distinctly reticulate thallus.

Characters (from Frey 1933): Upper side of thallus more or less distinctly reticulately ribbed, with ridges or bulges at least present in the middle clearly over the umbilicus, and with at least some rhizines present at edge when upper side is very smooth. Underside of thallus smooth, rhizines flat when cortex smoother, more or less distinctly belt-shaped, or at least partially fused to become more belt-shaped and fitted to underside. If rhizines completely absent on underside, then isolated rhizines present at the edge, or ribs above the umbilicus are whitish, at least much brighter than the other shallower ribs or ridges. Apothecia usually frequent, with flat or grooved disc. Spores colorless.

Umbilicaria sect. Polymorphae Frey 1931 F33
    |--U. cylindrica (Linnaeus) Delise ex Duby 1830 (see below for synonymy) F33
    |    |  i. s.: U. c. f. laciniata (Retzius) Frey 1933 (see below for synonymy) F33
    |    |         U. c. f. mesenteriformis (Wulf.) Frey 1933 (see below for synonymy) F33
    |    |         U. c. f. nudiuscula (Schaerer) Frey 1933 (see below for synonymy) F33
    |    |--U. c. var. cylindrica F33
    |    |--U. c. var. corrugatoides Frey 1933 F33
    |    |--U. c. var. delisei Nylander 1861 (see below for synonymy) F33
    |    |--U. c. var. denudata (Turner) Frey 1933 (see below for synonymy) F33
    |    |--U. c. var. fimbriata (Acharius) Nylander 1861 (see below for synonymy) F33
    |    `--U. c. var. tornata (Acharius) Nylander 1861 (see below for synonymy) F33
    |--U. proboscidea Schrader 1794 (see below for synonymy) F33
    |    |  i. s.: U. p. f. fimbriata (Turner) Leight. 1871 (see below for synonymy) F33
    |    |--U. p. var. proboscidea F33
    |    |--U. p. var. exasperata (Acharius) Nylander 1861 [=Gyrophora proboscidea var. exasperata Acharius 1803] F33
    |    `--U. p. var. pulla (Wulf.) Frey 1933 (see below for synonymy) F33
    `--U. virginis Schaerer 1841 (see below for synonymy) F33
         |--U. v. var. virginis F33
         `--U. v. var. meylani Frey 1933 F33

Umbilicaria cylindrica (Linnaeus) Delise ex Duby 1830 [=Lichen cylindricus Linnaeus 1753, Gyromium cylindricum (Linnaeus) Wahlenberg 1812, Gyrophora cylindrica (Linnaeus) Acharius 1803, Gyrophora polymorpha var. cylindrica (Linnaeus) Schaerer 1818, Gyrophora proboscidea var. cylindrica (Linnaeus) Mann 1825, Umbilicaria polymorpha var. cylindrica (Linnaeus) Schaerer 1826, U. proboscidea var. cylindrica (Linnaeus) Fries 1831; incl. Lichen crinitus Zoega ex Olafson & Povels 1775, Gyrophora cylindrica var. crinita (Zoega) Flot. 1850, Umbilicaria crinita (Zoega) Hoffmann 1794, Lichen proboscideus Hudson 1778] F33

Umbilicaria cylindrica var. delisei Nylander 1861 [=G. cylindrica f. delisei (Nylander) Lynge 1921, Gyrophora cylindrica var. delisei (Nylander) Sydow 1887, G. delisei (Nylander) Krempelh. 1871, Umbilicaria delisei (Nylander) Despr. ex Nylander 1861] F33

Umbilicaria cylindrica var. denudata (Turner) Frey 1933 [=Gyrophora proboscidea var. denudata Turner 1839, G. cylindrica f. denudata (Turner) Stein in Cohn 1879, G. cylindrica var. denudata (Turner) Mudd 1861] F33

Umbilicaria cylindrica var. fimbriata (Acharius) Nylander 1861 [=Gyrophora cylindrica var. fimbriata Acharius 1810, G. fimbriata (Acharius) Röhling 1813, Umbilicaria polymorpha var. cylindrica f. fimbriata (Acharius) Schaerer 1850] F33

Umbilicaria cylindrica f. laciniata (Retzius) Frey 1933 [=Lichen corneus var. laciniatus Retzius 1779, Gyrophora cylindrica var. laciniata (Retzius) Zahlbr. 1927, Umbilicaria cylindrica var. laciniata (Retzius) Wain. 1898; incl. G. cylindrica var. denticulata Acharius 1803, G. denticulata (Acharius) Floerke 1810, Umbilicaria polymorpha var. cylindrica f. denticulata (Acharius) Schaerer 1818] F33

Umbilicaria cylindrica f. mesenteriformis (Wulf.) Frey 1933 [=Lichen mesenteriformis Wulf. ex Jacquin 1781, Gyrophora cylindrica f. mesenteriformis (Wulf.) Anders 1928, G. cylindrica var. mesenteriformis (Wulf.) Flot. 1850] F33

Umbilicaria cylindrica f. nudiuscula (Schaerer) Frey 1933 [=Gyrophora polymorpha var. cylindrica f. nudiuscula Schaerer 1818, G. cylindrica var. nudiuscula (Schaerer) Zahlbr. 1927] F33

Umbilicaria cylindrica var. tornata (Acharius) Nylander 1861 [=Gyrophora tornata Acharius 1808, G. cylindrica var. tornata (Acharius) Arnold 1874] F33

Umbilicaria proboscidea Schrader 1794 non Lichen proboscideus Hudson 1778 [=Gyrophora polymorpha var. proboscidea (Schrader) Schaerer 1818, G. proboscidea (Schrader) Acharius 1803] F33

Umbilicaria proboscidea f. fimbriata (Turner) Leight. 1871 [=Gyrophora deusta var. fimbriata Turner 1839 non G. cylindrica var. fimbriata Acharius 1810, G. proboscidea var. fimbriata (Turner) Mudd 1861] F33

Umbilicaria proboscidea var. pulla (Wulf.) Frey 1933 [=Lichen pullus Wulf. ex Jacquin 1871; incl. U. proboscidea var. deplicans Nylander 1860, G. proboscidea f. deplicans (Nylander) Fries 1871] F33

Umbilicaria virginis Schaerer 1841 [=Gyrophora virginis (Schaerer) Frey 1929; incl. Umbilicaria rugifera Nylander 1861, Agyrophora rugifera (Nylander) Nylander 1896, Gyrophora rugifera (Nylander) Fries 1871, Umbilicaria stipitata Nylander 1861, Agyrophora stipitata (Nylander) Nylander 1896, Gyrophora stipitata (Nylander) Nylander ex Hue 1886-1887, G. tramnitziana Koerber 1871] F33

*Type species of generic name indicated

REFERENCES

[F33] Frey, E. 1933. Cladoniaceae (unter Ausschluss der Gattung Cladonia), Umbilicariaceae. In Die Flechten (Dr. L. Rabenhorst’s Kryptogamenflora von Deutschland, Österreich und der Schweiz vol. 9) sect. 4, part 1. Akademische Verlagsgesellschaft M. B. H.: Leipzig (reprinted 1971. Johnson Reprint Corporation: New York).

Umbilicaria section Velleae

Hairy navel lichen Umbilicaria hirsuta, from here.

Belongs within: Umbilicariaceae.

The section Velleae in the genus Umbilicaria is a group of lichens with a more or less smooth surface.

Characters (from Frey 1933): Upper side of thallus smooth or finely cracked, never with net-like ribs or wavy ridges. Underside of thallus with rhizines or, failing this, having a cracked and warty areolar network that is visible to the naked eye. Rhizines flexible, simple or branched, may be lumpy and warty, with finely granular cortex, not flat strap-shaped and not clearly externally defined. Cortex between rhizines at least partially clearly warty-areolate; when rhizines are entirely absent, areolar network is clearer, stronger and more regular. Apothecia usually rare, if present, strongly grooved. Spores colourless.

Umbilicaria sect. Velleae Frey 1931 F33
    |--U. grisea (Swartz) Acharius 1794 (see below for synonymy) F33
    |    |--U. g. f. grisea F33
    |    `--U. g. f. subpapyria Frey 1933 F33
    |--U. hirsuta (Swartz) Acharius 1793 (see below for synonymy) F33
    |    |--U. h. var. hirsuta F33
    |    |--U. h. var. melanotricha Flot. 1850 F33
    |    |--U. h. var. papyria (Acharius) Nylander 1861 (see below for synonymy) F33
    |    `--U. h. var. pyrenaica Frey 1933 F33
    |--U. polyrrhiza (Linnaeus) Acharius 1794 (see below for synonymy) F33
    |    |--U. p. var. polyrrhiza F33
    |    `--U. p. var. luxurians (Acharius) Nylander 1861 (see below for synonymy) F33
    `--U. vellea (Linnaeus) Acharius 1794 (see below for synonymy) F33

Umbilicaria grisea (Swartz) Acharius 1794 [=Lichen griseus Swartz ex Westring 1793, Gyrophora grisea (Swartz) Swartz ex Billberg & Swartz 1819, G. hirsuta var. grisea (Swartz) Fries 1871, Umbilicaria vellea var. spodochroa f. grisea (Swartz) Schaerer 1850; incl. Lichen murinus Acharius 1798, Gyrophora depressa var. hirsuta f. murina (Acharius) Schaerer 1818, G. hirsuta var. murina (Acharius) Floerke 1810, G. murina (Acharius) Acharius 1803, Lecidea hirsuta var. murina (Acharius) Sprengel 1827, Umbilicaria depressa var. murina (Acharius) Duby 1830, U. murina (Acharius) de Candolle ex Lam & de Candolle 1805] F33

Umbilicaria hirsuta (Swartz) Acharius 1793 [=Lichen hirsutus Swartz ex Westring 1793, Gyrophora hirsuta (Swartz) Acharius 1803, Lecidea hirsuta (Swartz) Sprengl 1827, Umbilicaria depressa var. hirsuta (Swartz) Schaerer 1826, U. vellea var. hirsuta (Swartz) Fries 1831; incl. Gyrophora hirsuta var. vestita Fries 1871] F33

Umbilicaria hirsuta var. papyria (Acharius) Nylander 1861 [=Gyrophora hirsuta var. papyria Acharius 1803, G. grisea f. papyria (Acharius) Zahlbr. 1927, U. murina f. papyria (Acharius) Harmand 1910, U. vellea var. depressa f. papyria (Acharius) Schaerer 1850] F33

Umbilicaria polyrrhiza (Linnaeus) Acharius 1794 [=Lichen polyrrhizos Linnaeus 1753, Gyromium polyrrhizum (Linnaeus) Wahlenberg 1820, Gyrophora polyrrhiza (Linnaeus) Körb. 1859; incl. Gyrophora diabolica Zahlbr. ex Herre 1906, Lichen pellitus Acharius 1794, Gyrophora depressa var. spodochroa f. pellita (Acharius) Schaerer 1818, Gyrophora pellita (Acharius) Acharius 1803, Umbilicaria pellita (Acharius) Acharius 1794, U. varia var. pellita (Acharius) Leight. 1856] F33

Umbilicaria polyrrhiza var. luxurians (Acharius) Nylander 1861 [=Gyrophora pellita var. luxurians Acharius 1810, G. luxurians (Acharius) Röhling 1813, G. polyrrhiza f. luxurians (Acharius) Fries 1871] F33

Umbilicaria vellea (Linnaeus) Acharius 1794 [=Lichen velleus Linnaeus 1753, Gyrophora depressa var. spodochroa f. vellea (Linnaeus) Schaerer 1818, G. vellea (Linnaeus) Acharius 1803; incl. Umbilicaria vellea var. diffracta Nylander 1869, G. vellea var. diffracta (Nylander) Zahlbr. 1927, U. vellea var. depressa f. prolifera Schaerer 1850, G. vellea f. prolifera (Schaerer) Arnold 1878, G. vellerea Nylander ex Hue 1891, U. vellerea (Nylander) Minks 1900] F33

*Type species of generic name indicated

REFERENCES

[F33] Frey, E. 1933. Cladoniaceae (unter Ausschluss der Gattung Cladonia), Umbilicariaceae. In Die Flechten (Dr. L. Rabenhorst’s Kryptogamenflora von Deutschland, Österreich und der Schweiz vol. 9) sect. 4, part 1. Akademische Verlagsgesellschaft M. B. H.: Leipzig (reprinted 1971. Johnson Reprint Corporation: New York).

Umbilicaria subgenus Gyrophoropsis

Umbilicaria spodochroa, from here.

Belongs within: Umbilicariaceae.

Gyrophoropsis is a subgenus of the lichen genus Umbilicaria containing species with relatively large spores.

Characters (from Frey 1933): Spores rather large, usually about 12 μm long and 6 μm wide, rarely smaller, colorless or brownish, at least distal spores within the ascus two- or three-celled or multicellular with brick-like cells.

Umbilicaria subg. Gyrophoropsis (Elenkin & Sav.) Zahlbr. 1927 [=Gyrophoropsis Elenkin & Sav. 1911] F33
    |--U. cinereorufescens (Schaerer) Frey 1931 (see below for synonymy) F33
    |--U. crustulosa (Acharius) Frey 1931 (see below for synonymy) F33
    |    |--U. c. var. crustulosa F33
    |    `--U. c. var. badiofusca Frey 1933 F33
    |--U. depressa (Acharius) Duby 1830 [=Gyrophora crustulosa var. depressa Acharius 1810] F33
    |--U. ruebeliana (DR. & Frey) Frey 1931 [=Gyrophora ruebeliana DR. & Frey 1929] F33
    |    |--U. r. f. ruebeliana F33
    |    `--U. r. f. rhizinosa Frey 1933 F33
    `--U. spodochroa (Acharius) Frey 1931 (see below for synonymy) F33
         |--U. s. var. spodochroa F33
         `--U. s. var. lapponica Frey 1933 F33

Umbilicaria cinereorufescens (Schaerer) Frey 1931 [=U. vellea var. spodochroa f. cinereorufescens Schaerer 1850, Gyrophora vellea f. cinereorufescens (Schaerer) Arnold 1878; incl. G. mammulata, G. cirrhosa f. mammulata (Acharius) Zahlbr. 1928, G. depressa var. mammulata (Acharius) Dalla Torre & Sarnth. 1902, G. spodochroa var. mammulata (Acharius) Arnold 1878, G. vellea var. mammulata (Acharius) Arnold 1878, G. tylorhiza Nylander ex Hue 1886-1887] F33

Umbilicaria crustulosa (Acharius) Frey 1931 [=Gyrophora crustulosa Acharius 1810, G. depressa var. crustulosa (Acharius) Dalla Torre & Sarnth. 1902, G. spodochroa var. crustulosa (Acharius) Arnold 1878, Umbilicaria spodochroa var. crustulosa (Acharius) Nylander 1861; incl. U. vellea var. spodochroa f. abortiva Schaerer 1850, Gyrophora cirrhosa f. abortiva (Schaerer) Zahlbr. 1927, G. vellea f. abortiva (Schaerer) Arnold 1850, Umbilicaria vellea var. depressa f. abortiva (Schaerer) Schaerer 1850, G. cirrhosa var. depressa (Acharius) Anders 1928, G. depressa (Acharius) Röhling 1813, G. spodochroa var. depressa (Acharius) Acharius 1810, G. crustulosa var. eximia Nylander 1875, G. depressa var. spodochroa f. leprosa Schaerer 1818, G. crustulosa f. leprosa (Schaerer) Zahlbr. 1927, G. spodochroa var. crustulosa f. leprosa (Schaerer) Arnold 1878, G. vellea f. leprosa (Schaerer) Zahlbr. 1927, Umbilicaria vellea var. cinereorufescens f. leprosa (Schaerer) Schaerer 1850, U. vellea var. spodochroa f. rupta Schaerer 1850, G. vellea f. rupta (Schaerer) Arnold 1878, U. vellea var. depressa f. rupta (Schaerer) Schaerer 1850, U. vellerea f. rupta (Schaerer) Dalla Torre & Sarnth. 1902, U. vellea var. depressa f. subpedicellata Schaerer 1850, Gyrophora vellea f. subpedicellata (Schaerer) Zahlbr. 1927, U. vellea var. depressa f. velleiformis Schaerer 1850, G. vellea var. velleiformis (Schaerer) Zahlbr. 1927] F33

Umbilicaria spodochroa (Acharius) Frey 1931 [=Lichen spadochrous Acharius 1798, Gyrophora spodochroa (Acharius) Acharius 1810; incl. Umbilicaria vellea var. cinereorufescens f. aenea Schaerer 1850, G. vellea f. aenea (Schaerer) Arnold 1878, U. cirrhosa Hoffmann 1790, G. cirrhosa (Hoffmann) Wain. 1888] F33

*Type species of generic name indicated

REFERENCES

[F33] Frey, E. 1933. Cladoniaceae (unter Ausschluss der Gattung Cladonia), Umbilicariaceae. In Die Flechten (Dr. L. Rabenhorst’s Kryptogamenflora von Deutschland, Österreich und der Schweiz vol. 9) sect. 4, part 1. Akademische Verlagsgesellschaft M. B. H.: Leipzig (reprinted 1971. Johnson Reprint Corporation: New York).

Baeomycetaceae

Baeomyces rufus, photographed by Bernd Haynold.

Belongs within: Lecanoromycetes.

The Baeomycetaceae are a family of lichens containing green algae, found growing on soil, rocks or similar substrates (Cannon & Kirk 2007).

Characters (from Cannon & Kirk 2007): Thallus varied, usually crustose or squamulose. Ascomata sessile or shortly stipitate, sometimes clustered, formed on specialised generally non-lichenised thalline branches, flat or convex, pink or brown, the wall of interwoven hyphae. Interascal tissue of simple or sparingly branched paraphyses, often swollen at the apices. Hymenial gel J+ or J-. Asci thin-walled, not thickened at the apex, with a J+ or J- apical pore. Ascospores hyaline, simple or transversely septate. Anamorph pycnidial.

Baeomycetaceae [Baeomycetales]
    |--Phyllobaeis EB03
    `--Baeomyces Persoon 1794 F33
         |--B. subg. Eubaeomyces Mueller 1862 F33
         |    `--B. roseus Persoon 1794 (see below for synonymy) F33
         |         |--B. r. var. roseus F33
         |         `--B. r. var. complicatus Erichsen ex Frey 1933 F33
         `--B. subg. Sphyridium (Flotow) Mueller 1862 [=Sphyridium Flotow 1843] F33
              |--B. placophyllus Acharius 1803 (see below for synonymy) F33
              |--B. rufus (Hudson) Rebent. 1804 (see below for synonymy) F33
              |    |  i. s.: B. r. f. exilis Erichsen in Frey 1933 F33
              |    |         B. r. f. fuscorufescens (Vainio) Frey 1933 [=B. fuscorufescens Vainio 1922] F33
              |    |         B. r. f. lignorum Acharius 1814 F33
              |    |         B. r. f. parasiticus (Fries) Zahlbr. 1927 (see below for synonymy) F33
              |    |         B. r. f. rupestris (Persoon) Harmand 1895 (see below for synonymy) F33
              |    |         B. r. f. sessilis (de Candolle) Nylander 1860 (see below for synonymy) F33
              |    |         B. r. f. sorediatus Frey 1933 F33
              |    |         B. r. f. subsquamulosus (Nylander) Frey 1933 (see below for synonymy) F33
              |    |         B. r. f. terrestris (Fries) Zahlbr. 1927 (see below for synonymy) F33
              |    |--B. r. var. rufus F33
              |    |--B. r. var. callianthus (Lettau) Lettau 1919 [=B. callianthus Lettau 1912] F33
              |    `--B. r. var. carneus (Flk.) Nylander 1860 [=B. carneus Flk. 1821] F33
              |         |--B. r. var. c. f. carneus F33
              |         `--B. r. var. c. f. polycephalus (Fries) Zahlbr. 1927 (see below for synonymy) F33
              `--B. speciosus (Körb.) Lindau 1913 [=Sphyridium speciosum Körb. ex Stein in Cohn 1879] F33

Baeomyces placophyllus Acharius 1803 [=Biatora placophylla (Acharius) Fries 1831, Lecidea placophylla (Acharius) Link 1833, Lichen placophyllus (Acharius) de Lamarck 1813, Ludovicia placophylla (Acharius) Trèvis. 1857, Patellaria placophylla (Acharius) Sprengel 1827, Sphyridium placophyllum (Acharius) Fries 1861] F33

Baeomyces roseus Persoon 1794 [=Lichen baeomyces Linnaeus 1781; incl. Baeomyces ericetorum (Linnaeus) de Candolle in de Lamarck & de Candolle 1805] F33

Baeomyces rufus (Hudson) Rebent. 1804 [=Lichen rufus Hudson 1762, Ba. rupestris var. rufus (Hudson) Acharius 1810, Sphyridium byssoides var. rufum Fries 1874; incl. Lichen byssoides Linnaeus 1767, Ba. byssoides (Linnaeus) Gartner, Meyer & von Schreber 1801, Ba. rufus f. byssoides Oliv. 1900, Ba. rupestris var. byssoides (Linnaeus) Acharius 1810, Biatora byssoides (Linnaeus) Fries 1845, Lecidea byssoides (Linnaeus) Link 1833, Sphyridium byssoides (Linnaeus) Beltram. 1858, Lichen cinereofuscus Jacquin 1762, Li. fungiformis Scopoli 1772, Stereocaulon fungiforme (Scopoli) Schaerer 1833, Sphyridium fungiforme (Scopoli) Flotow 1843] F33

Baeomyces rufus var. carneus f. polycephalus (Fries) Zahlbr. 1927 [=Sphyridium byssoides f. polycephalum Fries 1874, B. byssoides var. carneus f. polycephalus; incl. S. carneum f. botryosum Flot. 1849] F33

Baeomyces rufus f. parasiticus (Fries) Zahlbr. 1927 [=Sphyridium byssoides f. parasiticus Fries 1874] F33

Baeomyces rufus f. rupestris (Persoon) Harmand 1895 [=Ba. rupestris Persoon 1803, Biatora byssoides f. rupestris (Persoon) Fries 1831, Sphyridium byssoides f. rupestre Rabenhorst 1870, S. fungiforme f. rupestre (Persoon) Körb. 1855] F33

Baeomyces rufus f. sessilis (de Candolle) Nylander 1860 [=B. sessilis de Candolle in de Lamarck & de Candolle 1840; incl. B. sessilis var. rupicola de Candolle in de Lamarck & de Candolle 1840, B. sessilis var. sphagnicola de Candolle in de Lamarck & de Candolle 1840] F33

Baeomyces rufus f. subsquamulosus (Nylander) Frey 1933 [=B. rufus var. subsquamulosus Nylander 1877, Sphyridium byssoides var. subsquamulosum (Nylander) Arnold 1884] F33

Baeomyces rufus f. terrestris (Fries) Zahlbr. 1927 [=Sphyridium byssoides f. terrestre Fries 1874] F33

*Type species of generic name indicated

REFERENCES

Cannon, P. F., & P. M. Kirk. 2007. Fungal Families of the World. CABI.

[EB03] Eriksson, O. E., H. O. Barah, R. S. Currah, K. Hansen, C. P. Kurtzman, G. Rambold & T. Laessøe (eds.) 2003. Outline of Ascomycota—2003. Myconet 9: 1-89.

[F33] Frey, E. 1933. Cladoniaceae (unter Ausschluss der Gattung Cladonia), Umbilicariaceae. In Die Flechten (Dr. L. Rabenhorst’s Kryptogamenflora von Deutschland, Österreich und der Schweiz vol. 9) sect. 4, part 1. Akademische Verlagsgesellschaft M. B. H.: Leipzig (reprinted 1971. Johnson Reprint Corporation: New York).

Ectolechiaceae

Sporopodium xantholeucum, photographed by André Aptroot.

Belongs within: Lecanoromycetidae.

The Ectolechiaceae are a mostly tropical family of lichens containing green algae.

Characters (from Cannon & Kirk 2007): Thallus usually foliose or crustose. Ascomata apothecial, pale, convex, without a well-developed margin. Interascal tissue of very narrow anastomosing pseudoparaphysis-like hyphae, variously pigmented, the apices thin-walled. Asci with a J+ apical cap and a very well-developed ocular chamber, with an outer J+ gelatinous layer, mostly 1- or 2-spored. Ascospores muriform, usually thin-walled. Sometimes with cephalodia. Anamorphs pycnidial, often with campylidia.

<==Ectolechiaceae
    |--Badimia EB03
    |--Badimiella EB03
    |--Barubria EB03
    |--Kantvilasia EB03
    |--Lasioloma EB03
    |--Loflammia EB03
    |--Logilvia EB03
    |--Lopadium EB03
    |--Sporopodiopsis EB03
    |--Tapellaria EB03
    |--Calopadia EB03 [incl. Cyrta E99, Pyrenotrichum E99]
    |    `--‘*Cyrta’ licaniae E99
    `--Sporopodium EB03 [incl. Acleistomyces E99]
         `--S. xantholeucum [incl. Acleistomyces zolleriae] E99

*Type species of generic name indicated

REFERENCES

Cannon, P. F., & P. M. Kirk. 2007. Fungal Families of the World. CABI.

[E99] Eriksson, O. E. (ed.) 1999. Notes on ascomycete systematics. Nos 2440-2755. Myconet 2: 1-41.

[EB03] Eriksson, O. E., H. O. Barah, R. S. Currah, K. Hansen, C. P. Kurtzman, G. Rambold & T. Laessøe (eds.) 2003. Outline of Ascomycota—2003. Myconet 9: 1-89.

Stereocaulon botryosum

Stereocaulon botryosum, from here.

Belongs within: Stereocaulaceae.

Stereocaulon botryosum is a species of lichen found in northern Eurasia and North America.

Characters (from Frey 1933, translated using Google Translate; all errors my own): Dusty thallus often present, both among strongly developed podetia as well as particularly at the edge of encrusted forms. Podetia in dense patches, gradating at the lower, younger edges into a crusty thallus. Entire surface usually smooth(?). Individual well-developed podetia 2.5(-3) cm high, very firmly stuck to the rock, often radiating from a common center, but often growing side by side like grass. Larger podetia often thickened at the base to 4 mm, brown to brown-black, whitish upwards, with numerous tree-like branches, the outermost strong podetien usually very cauliflower-like. Phyllocladia slightly bluish or grayish white, clustered very tightly, warty, rarely slightly elongated scaly, or widened, and then slightly, usually 0.2 to 0.3 mm wide. Apothecia occasionally on extended branches, otherwise somewhat outstanding, terminal, often grouped together, larger apothecia divided into many sections(?), dark brown, first flat and surrounded by a fine, soon vanishing edge, 1.5-3 mm wide. Hyphae of the central strand (2-)3-4(-5) μm thick, fairly tightly parallel, more or less dense, often somewhat bonded; those against the base of podetia finer, smoother, less glued, with thin walls; those against the points of the podetia thick, rough and more glued, rather thich-walled. Average area of ​​the cavities are, however, more than half of the entire thickness. The external hyphae form a fairly clearly defined cylinder, 30-50 μm thick, the outermost hyphae 7-10 μm thick, thick-walled. Felt to 60 μm thick. Reaction KOH-. Cortex of phyllocladia contiguous, about 15 μm thick, often thinner, hyphae 3.5 μm thick, dimly discernible. Gonidia in layer about 50 μm thick, 7-10 μm in size, Cystococcus. Cephalodia hidden in tomentum or grape-like and warty, less than 0.5 mm, purple, or whitish dusted, often on the surface of the pad and then larger, up to 2.5 mm wide. Mostly containing Stigonema, but also Nostoc. Underside of apothecium covered with a dense felt, which often reaches up to the edge of the disc, not uncommonly small phyllocladia are woven into this tomentum. In places, this felt is dense and forms a loose palisade. However, elsewhere this felt is very loose, consisting of highly branched and confused hyphae, to 150 μm thick. Where the felt is designed more like a palisade, the hyphae are up to 8 μm thick and moderately thick-walled; in the loose, but thicker parts of the tomentum, hyphae are only 4-5 μm and moderately thin-walled. Central strand breaking off suddenly under the central cone; packing hyphae of the latter tightly glued, thick-walled, indistinct, only the 2-2.5 μm wide lumina of the cells clearly visible. Central cone granulated. Hypothecium quite significantly confined, 40 μm high; parathecium indistinct; hymenium 60-65 μm high, the hyphae 2-1.5 μ thick, J colors only the asci blue, not the hymenial gelatine. Paraphyses 1-1.5 μm thick, mostly loose, head cells to 4.5 μ thick, yellow-brown, asci 50-55 × 8-10 μm, spores often poorly developed, 20-31 × 4-5 μm. Pycnidia often numerous, 0.09 to 0.12 mm, brown spots on the tips of the phyllocladia, wall pale brown, colorless in the lower part. Conidia 5-7 × 0.6-0.8 μ, cylindrical.

<==Stereocaulon botryosum Acharius 1810 (see below for synonymy) F33
    |  i. s.: S. b. f. confluens (Magnusson 1926) Frey 1933 F33
    |         S. b. f. congestum (Magnusson 1926) Frey 1933 F33
    |         S. b. f. depressum (Magnusson 1926) Frey 1933 F33
    |         S. b. f. dissolutum (Magnusson) Frey 1933 [=S. fastigiatum f. dissolutum Magnusson 1926] F33
    |         S. b. f. finmarkicum (Magnusson) Frey 1933 F33
    |         S. b. f. globuliferum (Magnusson 1926) Frey 1933 F33
    |         S. b. f. irregulare (Magnusson 1926) Frey 1933 F33
    |         S. b. f. simplicior (Magnusson 1926) Frey 1933 F33
    |         S. b. f. spathuliferum (Vainio) Frey 1933 [=S. spathuliferum Vainio 1909] F33
    |--S. b. f. botryosum F33
    `--S. b. var. subincrustatum Frey 1933 F33

Stereocaulon botryosum Acharius 1810 [incl. S. fastigiatum Anzi 1860, S. evolutum f. fastigiatum (Anzi) Anders 1928, S. evolutum var. fastigiatum (Anzi) Fries 1874, S. paschale var. pulvinatum Schaerer 1833, S. corallinum var. pulvinatum (Schaerer) Schaerer 1850] F33

*Type species of generic name indicated

REFERENCES

[F33] Frey, E. 1933. Cladoniaceae (unter Ausschluss der Gattung Cladonia), Umbilicariaceae. In Die Flechten (Dr. L. Rabenhorst’s Kryptogamenflora von Deutschland, Österreich und der Schweiz vol. 9) sect. 4, part 1. Akademische Verlagsgesellschaft M. B. H.: Leipzig (reprinted 1971. Johnson Reprint Corporation: New York).

Physciaceae

Heterodermia leucomelos, from here.
Belongs within: Lecanoromycetidae.

The Physciaceae are a family of lichenised fungi associated with green algae, often found growing on nitrogen-rich substrates.

Characters (from Cannon & Kirk 2007): Thallus crustose, foliose or fruticose. Ascomata apothecial, sessile, occasionally immersed, usually with a distinct margin. Interascal tissue composed of ± branched and anastomosing paraphyses, often with swollen and/or pigmented apices. Asci with a J+ apical cap, with a conical or funnel-shaped more weakly staining apical cushion, usually with a J+ gelatinized outer layer. Ascospores usually septate, usually dark brown with thickened walls and septa. Anamorphs pycnidial.

<==Physciaceae
    |--Amandinea EB03
    |--Anaptychia ciliaris EB03, RS99
    |--Australiaena EB03
    |--Awasthia EB03
    |--Dermatiscum EB03
    |--Dermiscellum EB03
    |--Dimelaena EB03
    |--Diploicia EB03
    |--Diplotomma EB03
    |--Dirinaria EB03
    |--Hafellia EB03
    |--Hyperphyscia EB03
    |--Mischoblastia EB03
    |--Mobergia EB03
    |--Phaeophyscia EB03
    |    |--P. decolor (Kashiwadani) Essl. 1978 [=Physcia decolor Kashiwadani 1975] S03
    |    `--P. hispidula J87
    |--Phaeorrhiza EB03
    |--Physciella EB03
    |--Physconia EB03
    |--Pyxine EB03
    |--Redonia EB03
    |--Rinodina EB03
    |--Rinodinella EB03
    |--Santessonia namibensis EB03, E99
    |--Tornabea EB03
    |--Buellia EB03
    |    |--*B. disciformis E99
    |    `--B. aethalea E99
    |--Gassicurtia EB03
    |--Physcia EB03
    |    |--P. aipolia E99
    |    |--P. alba J87
    |    |--P. caesia F33
    |    `--P. tribacia F33
    `--Heterodermia EB03
         |--H. chondroidea Web. & Awasthi 1971 S03
         |--H. circinalis J87
         |--H. comosa J87
         |--H. diademata BB05
         |--H. japonica BB05
         |--H. leucomelos J87
         |    |--H. l. ssp. leucomelos J87
         |    `--H. l. ssp. boryi J87
         `--H. lutescens J87

Nomen illegitimum: Buellia parasema E99

*Type species of generic name indicated

REFERENCES

[BB05] Banik, D. & P. Bujarbarua. 2005. Lichens on Agapetes D. Don ex G. Don (Ericaceae) in India. Bulletin of the Botanical Survey of India 47 (1-4): 59-62.

Cannon, P. F., & P. M. Kirk. 2007. Fungal Families of the World. CABI.

[E99] Eriksson, O. E. (ed.) 1999. Notes on ascomycete systematics. Nos 2440-2755. Myconet 2: 1-41.

[EB03] Eriksson, O. E., H. O. Barah, R. S. Currah, K. Hansen, C. P. Kurtzman, G. Rambold & T. Laessøe (eds.) 2003. Outline of Ascomycota – 2003. Myconet 9: 1-89.

[F33] Frey, E. 1933. Cladoniaceae (unter Ausschluss der Gattung Cladonia), Umbilicariaceae. In Die Flechten (Dr. L. Rabenhorst’s Kryptogamenflora von Deutschland, Österreich und der Schweiz vol. 9) sect. 4, part 1. Akademische Verlagsgesellschaft M. B. H.: Leipzig (reprinted 1971. Johnson Reprint Corporation: New York).

[J87] Judd, W. S. 1987. Floristic study of Morne La Visite and Pic Macaya National Parks, Haiti. Bulletin of the Florida State Museum – Biological Sciences 32 (1): 1-136.

[RS99] Rossman, A. Y., G. J. Samuels, C. T. Rogerson & R. Lowen. 1999. Genera of Bionectriaceae, Hypocreaceae and Nectriaceae (Hypocreales, Ascomycetes). Studies in Mycology 42: 1-248.

[S03] Sinha, G. P. 2003. Additions to the lichen flora of India from Sikkim. Bulletin of the Botanical Survey of India 45 (1-4): 221-224.

Teloschistales

Elegant sunburst lichen Xanthoria elegans, from here.
Belongs within: Lecanoromycetidae.

The Teloschistales is a clade of lichenised fungi associated with green algae.

<==Teloschistales [Teloschistineae]
    |--Fuscideaceae EB03
    |    |--Fuscidea lygaea LK04
    |    |--Lettauia EB03
    |    |--Maronea EB03
    |    |    |--M. chilensis LK04
    |    |    `--M. constans LK04
    |    |--Orphniospora EB03
    |    |--Ropalospora EB03
    |    `--Sarrameana EB03
    `--+--Megalosporaceae EB03
       |    |--Austroblastenia EB03
       |    |--Megaloblastenia EB03
       |    `--Megalospora LK04
       |         |--M. sulphurata E99
       |         `--M. tuberculosa LK04
       `--Teloschistaceae EB03
            |  i. s.: Cephalophysis EB03
            |         Fulgensia EB03
            |         Ioplaca EB03
            |         Seirophora EB03
            |         Xanthodactylon EB03
            |         Xanthopeltis EB03
            |         Caloplaca ML04
            |           |--C. crocea J87
            |           `--C. gomerana ML04
            |         Josefpoeltia Kondratyuk & Kärnefelt 1997 EB03, E99
            |           |--*J. boliviensis Kondratyuk & Kärnefelt 1997 E99
            |           `--J. sorediosa Kondratyuk & Kärnefelt 1997 E99
            |--+--Letrouitia LK04 [Letrouitiaceae]
            |  |    `--L. domingensis LK04
            |  `--Xanthoria LK04
            |       |--X. elegans E99
            |       |--X. fallax E99
            |       |--X. parietina ML04
            |       `--X. polycarpa LH04
            `--+--Xanthomendoza Kondratyuk & Kärnefelt 1997 LK04, E99
               |    |--*X. mendozae (Räs.) Kondratyuk & Kärnefelt 1997 [=Xanthoria mendozae] E99
               |    `--X. fallax LK04
               `--Teloschistes LK04
                    |--T. chrysophthalmus E99
                    |--T. exilis LK04
                    `--T. flavicans J87

*Type species of generic name indicated

REFERENCES

[E99] Eriksson, O. E. (ed.) 1999. Notes on ascomycete systematics. Nos 2440-2755. Myconet 2: 1-41.

[EB03] Eriksson, O. E., H. O. Barah, R. S. Currah, K. Hansen, C. P. Kurtzman, G. Rambold & T. Laessøe (eds.) 2003. Outline of Ascomycota – 2003. Myconet 9: 1-89.

[J87] Judd, W. S. 1987. Floristic study of Morne La Visite and Pic Macaya National Parks, Haiti. Bulletin of the Florida State Museum – Biological Sciences 32 (1): 1-136.

[LH04] Liu, Y. J. & B. D. Hall. 2004. Body plan evolution of ascomycetes, as inferred from an RNA polymerase II phylogeny. Proceedings of the National Academy of Sciences of the USA 101: 4507-4512.

[LK04] Lutzoni, F., F. Kauff, C. J. Cox, D. McLaughlin, G. Celio, B. Dentinger, M. Padamsee, D. Hibbett, T. Y. James, E. Baloch, M. Grube, V. Reeb, V. Hofstetter, C. Schoch, A. E. Arnold, J. Miadlikowska, J. Spatafora, D. Johnson, S. Hambleton, M. Crockett, R. Shoemaker, G.-H. Sung, R. Lücking, T. Lumbsch, K. O'Donnell, M. Binder, P. Diederich, D. Ertz, C. Gueidan, K. Hansen, R. C. Harris, K. Hosaka, Y.-W. Lim, B. Matheny, H. Nishida, D. Pfister, J. Rogers, A. Rossman, I. Schmitt, H. Sipman, J. Stone, J. Sugiyama, R. Yahr & R. Vilgalys. 2004. Assembling the fungal tree of life: progress, classification, and evolution of subcellular traits. American Journal of Botany 91 (10): 1446-1480.

[ML04] Miadlikowska, J., & F. Lutzoni. 2004. Phylogenetic classification of peltigeralean fungi (Peltigerales, Ascomycota) based on ribosomal RNA small and large subunits. American Journal of Botany 91 (3): 449-464.

Collematineae

Jelly lichen Leptogium cyanescens, from here.
Belongs within: Lecanoromycetidae.

The Collematineae are a clade of mostly cyanobacterium-associated lichenised fungi. Most species of Collematineae have gelatinous or homoiomerous fruiting bodies (Miadlikowska & Lutzoni 2004).

Characters (from Miadlikowska & Lutzoni 2004): Lichen-forming ascomycetes. Mostly bipartite with cyanobacteria, rarely tripartite. Thalli non- or gelatinous, squamulose, foliose, or subfruticose, rarely crustose; homoiomerous or heteromerous. Cortex present on both sides of the thallus or lower side ecorticated and sometimes with dark prothallus. Vegetative propagules (soredia, isidia, or phyllidia) common. Apothecia immersed or sessile; lecideine, biatorine, or lecanorine. Paraphyses simple or branched, sometimes strongly gelatinous. Ascospores aseptate, one- to multiseptate, or muriform, colorless. Asci with an amyloid external wall layer, I+ blue. Secondary substances uncommon or absent.

<==Collematineae
    |--Pannariaceae ML04
    |    |--Degelia EB03
    |    |--Erioderma EB03
    |    |--Fuscoderma EB03
    |    |--Fuscopannaria EB03
    |    |--Leioderma EB03
    |    |--Moelleropsis humida EB03, ML04
    |    |--Pannaria conoplea ML04
    |    |--Parmeliella atlantica EB03, RS99
    |    |--Psoroma hypnorum ML04
    |    |--Psoromidium EB03
    |    |--Santessoniella EB03
    |    `--Siphulastrum ML04
    `--+--Placynthiaceae ML04
       |    |--Hertella EB03
       |    |--Koerberia EB03
       |    |--Leptochidium EB03
       |    |--Placynthiopsis EB03
       |    |--Placynthium nigrum ML04
       |    |--Polychidium EB03
       |    `--Vestergrenopsis EB03
       `--Collemataceae ML04
            |--Homothecium EB03
            |--Leciophysma EB03
            |--Leightoniella EB03
            |--Physma EB03
            |--Ramalodium EB03
            |--Staurolemma EB03
            |--Collema flaccidum ML04
            `--Leptogium ML04 [incl. Pericoccis Clements in Clements & Shear 1931 RS99]
                 |--L. azureum J87
                 |--L. cyanescens ML04
                 |--L. gelatinosum ML04
                 `--‘Hypocrea’ leptogiicola Cooke & Massee 1890 (see below for synonymy) RS99

‘Hypocrea’ leptogiicola Cooke & Massee 1890 [=Broomella leptogiicola (Cooke & Massee) Saccardo 1891, B. lichenicola (l. c.), *Pericoccis leptogiicola (Cooke & Massee) Clements in Clements & Shear 1931] RS99

*Type species of generic name indicated

REFERENCES

[EB03] Eriksson, O. E., H. O. Barah, R. S. Currah, K. Hansen, C. P. Kurtzman, G. Rambold & T. Laessøe (eds.) 2003. Outline of Ascomycota – 2003. Myconet 9: 1-89.

[J87] Judd, W. S. 1987. Floristic study of Morne La Visite and Pic Macaya National Parks, Haiti. Bulletin of the Florida State Museum – Biological Sciences 32 (1): 1-136.

[ML04] Miadlikowska, J., & F. Lutzoni. 2004. Phylogenetic classification of peltigeralean fungi (Peltigerales, Ascomycota) based on ribosomal RNA small and large subunits. American Journal of Botany 91 (3): 449-464.

[RS99] Rossman, A. Y., G. J. Samuels, C. T. Rogerson & R. Lowen. 1999. Genera of Bionectriaceae, Hypocreaceae and Nectriaceae (Hypocreales, Ascomycetes). Studies in Mycology 42: 1-248.

Peltigeraceae

Membranous dog-lichen Peltigera membranacea, from here.
Belongs within: Peltigerineae.

The Peltigeraceae are a family of generally cyanobacterium-associated lichenised fungi, found worldwide but particularly in temperate regions.

Characters (from Cannon & Kirk 2007): Lichenized with cyanobacteria or green algae (then often with cyanobacterial cephalodia). Thallus usually large and spreading, foliose, lobed, usually green or dark greyish, matt or shiny, sometimes with cephalodia, corticate only on the upper surface, the lower surface often tomentose and with conspicuous rhizoids. Ascomata apothecial, formed on the upper surface, laminar or marginal, with a thin covering layer which breaks apart at an early stage to expose the hymenium, flat or saddleshaped, brown or black. Interascal tissue of simple conglutinate paraphyses, sometimes swollen at the tip. Asci fissitunicate with a thickened apex, a J+ apical ring and a more weakly J+ gelatinized outer layer. Ascospores hyaline to brown, elongate, transversely septate, sometimes ornamented. Anamorphs pycnidial, poorly known.

<==Peltigeraceae
    |--Solorina [Solorinaceae] ML04
    |    |--S. crocea (Linnaeus) Ach. 1810 [=Lichen croceus Linnaeus 1753] M60
    |    `--S. saccata ML04
    `--Peltigera ML04
         |  i. s.: P. austroamericana J87
         |         P. coloradoensis Gyelnik 1930 M60
         |         P. didactyla RS99
         |         P. meridiana M60
         |         P. neopolydactyla ML04
         |--P. (subg. Peltidea) nigripunctata Bitt. 1909 [incl. P. nigripunctata f. farinosa Gyelnik 1932] M60
         `--P. subg. Peltigera M60
              |  i. s.: P. dolichorhiza Nyl. 1888 [=P. polydactyla var. dolichorhiza Nyl. 1860, P. dolichorrhiza] M60
              |           |--P. d. var. dolichorhiza [incl. P. dolichorhiza f. javanica Gyelnik 1930] M60
              |           |--P. d. var. nana (Nyl.) Murray 1960 (see below for synonymy) M60
              |           `--P. d. var. oceanica [=P. oceanica Gyelnik 1931] M60
              |         P. frigida M60
              |         P. horizontalis M60
              |           |--P. h. var. horizontalis M60
              |           `--P. h. var. muscorum (Schleich) Schaer. 1833 [=P. muscorum Schleich 1823] M60
              |                |--P. h. var. m. f. muscorum M60
              |                `--P. h. var. m. f. albido-pruinosa Murray 1960 M60
              |         P. lairdii Dodge & Rudolph 1955 M60
              |         P. scutata (Dicks) Duby 1830 [=P. polydactyla var. scutata Müll. Arg. 1896] M60
              |         P. virescens (Stnr.) Gyelnik 1932 (see below for synonymy) M60
              |--+--P. scabrosa ML04
              |  `--+--P. polydactylum (Neck.) Hoffm. 1790 ML04, M60 (see below for synonymy)
              |     |    |--P. p. var. polydactylum M60
              |     |    |    |--P. p. var. p. f. polydactylum M60
              |     |    |    `--P. p. var. p. f. microcarpa M60
              |     |    `--P. p. var. magyarica (Gyelnik) Murray 1960 [=P. magyarica Gyelnik 1937] M60
              |     `--+--P. occidentalis ML04
              |        `--P. hymenina ML04 [=P. polydactyla var. hymenina M60]
              `--+--P. hydrothyria Miadlikowska & Lutzoni 2000 [=Hydrothyria venosa] ML04
                 `--+--+--P. venosa ML04
                    |  `--+--P. leucophlebia ML04
                    |     `--+--P. malacea (Ach.) Funck 1827 ML04, M60 [=Peltidea malacea Ach. 1814 M60]
                    |        `--+--P. aphthosa ML04
                    |           `--P. britannica ML04
                    |--+--P. elisabethae ML04
                    |  `--+--P. neckeri ML04
                    |     `--P. polydactyloides Nyl. 1863 ML04, M60 (see below for synonymy)
                    |--P. retifoveata ML04
                    `--+--+--P. monticola ML04
                       |  `--P. ponojensis ML04
                       `--+--P. continentalis ML04
                          |--P. kristinssonii ML04
                          `--+--+--+--P. extenuata ML04
                             |  |  `--P. lambinonii ML04
                             |  `--+--P. laciniata ML04
                             |     `--P. rufescens Humb. 1793 ML04, M60 (see below for synonymy)
                             `--+--P. cinnamomea ML04
                                `--+--P. degenii ML04
                                   |--P. membranacea ML04
                                   `--+--P. canina (Linnaeus) Willd. 1787 ML04, M60 [=Lichen caninus Linnaeus 1753 M60]
                                      |    |--P. c. var. canina [incl. P. c. var. albescens, P. c. var. ulorrhiza] M60
                                      |    `--P. c. var. spuria (Ach.) Schaer 1833 (see below for synonymy) M60
                                      |         |--P. c. var. s. f. spuria M60
                                      |         `--P. c. var. s. f. sorediata Schaer. 1850 M60
                                      `--P. praetextata (Flk.) Wain 1899 ML04, M60 (see below for synonymy)

Peltigera canina var. spuria (Ach.) Schaer 1833 [=Lichen spurius Ach. 1798, P. rufescens var. spuria Kitk 1871; incl. P. pusilla Müller 1896, P. canina var. pusilla Bab. in Hooker 1855] M60

Peltigera dolichorhiza var. nana (Nyl.) Murray 1960 [=P. polydactyla var. nana, P. nana Wainio 1913, P. nana var. philippina Gyelnik 1936 (nom. illeg.); incl. P. nana f. nervosa Gyelnik apud Zahlbr. 1941] M60

Peltigera polydactyloides Nyl. 1863 ML04, M60 [=P. polydactyla var. polydactyloides M60; incl. P. crassoides Gyeln. 1930 M60]

Peltigera polydactylum (Neck.) Hoffm. 1790 ML04, M60 [=Lichen polydactylon Neck. 1771 M60, Peltidea polydactyla Hooker 1844 M60; incl. P. polydactyla f. lophyra M60, P. polydactyla f. minor Krmph. 1870 M60]

Peltigera praetextata (Flk.) Wain 1899 ML04, M60 [=Peltidea ulorrhiza var. praetextata Flk. apud Sommerf. 1826 M60; incl. P. nitens f. zeelandica Gyeln. 1929 M60]

Peltigera rufescens Humb. 1793 ML04, M60 [=Lichen caninus var. rufescens Weis. 1770 M60, P. canina var. rufescens (Weis.) Mudd 1861 M60; incl. P. rufescens var. spuria Hooker 1867 non Lichen spurius Ach. 1798 M60]

Peltigera virescens (Stnr.) Gyelnik 1932 [=P. rufescens var. virescens Steiner apud Zahlbr. & Zederb. 1907; incl. P. pellucida f. dilacerata Gyeln. apud Zahlbr. 1941, P. degeni f. tasmaniae Gyel. 1929, P. virescens var. tasmaniae Gyelnik apud Zahlbruckner 1941, P. tereziana Gyelnik 1928] M60

*Type species of generic name indicated

REFERENCES

Cannon, P. F., & P. M. Kirk. 2007. Fungal Families of the World. CABI.

[J87] Judd, W. S. 1987. Floristic study of Morne La Visite and Pic Macaya National Parks, Haiti. Bulletin of the Florida State Museum – Biological Sciences 32 (1): 1-136.

[ML04] Miadlikowska, J., & F. Lutzoni. 2004. Phylogenetic classification of peltigeralean fungi (Peltigerales, Ascomycota) based on ribosomal RNA small and large subunits. American Journal of Botany 91 (3): 449-464.

[M60] Murray, J. 1960. Studies on New Zealand lichens. Part III.—The family Peltigeraceae. Transactions of the Royal Society of New Zealand 88 (3): 381-399.

[RS99] Rossman, A. Y., G. J. Samuels, C. T. Rogerson & R. Lowen. 1999. Genera of Bionectriaceae, Hypocreaceae and Nectriaceae (Hypocreales, Ascomycetes). Studies in Mycology 42: 1-248.