Ptilodontoidea

Skull of Ptilodus mediaevus, copyright Nobu Tamura.

Belongs within: Cimolodonta.

The Ptilodontoidea are a group of multituberculate mammals known from the Upper Cretaceous to the Eocene of Eurasia and North America, characterised by a very large, arcuate lower rear premolar protruding dorsally above the level of the molars (Kielan-Jaworowska & Hurum 2001).

Characters (from Kielan-Jaworowska & Hurum 2001): Dental formula 2.0.4.2/1.0.2.2; enamel usually microprismatic (gigantoprismatic in Cimolodon); I2 probably single-cusped, I3 placed on margin of premaxilla; lower incisor usually gracile, completely covered with enamel (robust with limited enamel band in Neoliotomus); P4 elongated, shaped as isosceles triangle in lateral view, protruding ventrally over level of anterior premolars and molars; p4 very large, arcuate, strongly protruding dorsally over level of molars; molars with ornamentation of grooves and ridges, cusps tending to coalesce. Snout wide, gently incurved in front of zygomatic arches. Frontals small, pointed posteriorly.

<==Ptilodontoidea
    |--Cimolodontidae K-JH01
    |    |--Anconodon Jepsen 1940 K-JH01
    |    |--Liotomus Cope 1884 K-JH01
    |    `--Cimolodon Marsh 1889 K-JH01
    |         `--C. electus K-JH01
    `--+--Neoliotomus Jepsen 1930 K-JH01
       `--+--Ptilodontidae [Chirogidae] K-JH01
          |    |--Kimbetohia Simpson 1936 K-JH01
          |    |--Prochetodon Jepsen 1940 K-JH01
          |    |--Baiotomeus Krause 1987 SFY02
          |    |    |--B. douglassi (Simpson 1935) SFY02
          |    |    |--B. lamberti Krause 1987 SFY02
          |    |    `--B. russelli Scott, Fox & Youzwyshyn 2002 SFY02
          |    `--Ptilodus Cope 1881 SFY02
          |         |--P. douglassi Simpson 1935 S35b
          |         |--P. fractus Dorr 1952 SFY02
          |         |--P. gidleyi Simpson 1935 S35b
          |         |--P. gnomus Scott, Fox & Youzwyshyn 2002 SFY02
          |         |--P. kummae Krause 1977 SFY02
          |         |--P. mediaevus Cope 1881 SFY02
          |         |--P. montanus Douglass 1908 [incl. P. admirabilis Hay 1930, P. gracilis Gidley 1909 (preoc.)] S35b
          |         |--P. sinclairi Simpson 1935 S35b
          |         |--P. trovessartianus S35b
          |         |--P. tsosiensis Sloan 1981 SFY02
          |         `--P. wyomingensis D07
          `--Neoplagiaulacidae K-JH01
               |--Xyronomys Rigby 1980 K-JH01
               |--Cernaysia Vianey-Liaud 1986 K-JH01
               |--Krauseia Vianey-Liaud 1986 K-JH01
               |--Mesodma Jepsen 1940 K-JH01
               |--Mimetodon Jepsen 1940 K-JH01
               |--Mesodmops Tong & Wang 1994 K-JH01
               |--Xanclomys Rigby 1980 K-JH01
               |--Neoplagiaulax Lemoine 1882 K-JH01
               |    `--N. hazeni K-JH01
               |--Parectypodus Jepsen 1930 K-JH01
               |    |--P. jepseni Simpson 1935 S35b
               |    |--P. laytoni K-JH01
               |    |--P. simpsoni S35b
               |    `--P. tardus S35b
               `--Ectypodus Matthew & Granger 1921 K-JH01
                    |--*E. musculus Matthew & Granger 1921 S35a
                    |--E. cochranensis S35b
                    |--E. grangeri Simpson 1935 S35b
                    |--E. russelli Simpson 1935 S35b
                    |--E. silberlingi Simpson 1935 S35b
                    `--E. tardus K-JH01

*Type species of generic name indicated

REFERENCES

[D07] Dixon, D. 2007. The Complete Illustrated Encyclopedia of Dinosaurs & Prehistoric Creatures. Hermes House: London.

[K-JH01] Kielan-Jaworowska, Z., & J. H. Hurum. 2001. Phylogeny and systematics of multituberculate mammals. Palaeontology 44 (3): 389–429.

[SFY02] Scott, C. S., R. C. Fox & G. P. Youzwyshyn. 2002. New earliest Tiffanian (late Paleocene) mammals from Cochrane 2, southwestern Alberta, Canada. Acta Palaeontologica Polonica 47: 691–704.

[S35a] Simpson, G. G. 1935a. The Tiffany fauna, Upper Paleocene. I.—Multituberculata, Marsupialia, Insectivora, and ?Chiroptera. American Museum Novitates 795: 1–19.

[S35b] Simpson, G. G. 1935b. New Paleocene mammals from the Fort Union of Montana. Proceedings of the United States National Museum 83 (2981): 221–244.

Cimolodonta

Lower jaw of Cimexomys, from Palaeos.

Belongs within: Multituberculata.
Contains: Ptilodontoidea.

The Cimolodonta are a group of multituberculate mammals known from the Early Cretaceous to the Eocene, including all multituberculates to survive the end of the Cretaceous (Kielan-Jaworowska & Hurum 2001).

Characters (from Kielan-Jaworowska & Hurum 2001): Enamel prismatic; dental formula 2.0.1–4.2/1.0.1–2.2; I1 absent, I2 single- or two-cusped, I3 small, single-cusped; p1 and p2 absent, p3, if present, peg-like, non-functional; p4 primitively arcuate with oblique ridges, single or no posterobuccal cusp present.

Cimolodonta [Boffiini]
    |--Barbatodon Râdulescu & Samson 1986 K-JH01
    |--Dakotamys Eaton 1995 K-JH01
    |    `--D. malcolmi K-JH01
    |--Bryceomys Eaton 1995 K-JH01
    |    `--B. fumosus K-JH01
    |--Paracimexomys Archibald 1982 K-JH01
    |    |--P. crossi K-JH01
    |    `--P. robinsoni K-JH01
    |--Cimexomys Sloan & van Valen 1965 K-JH0
    |    |--C. antiquus D07
    |    |--C. gratus D07
    |    |--C. gregoryi D07
    |    |--C. hausoi D07
    |    |--C. judithae K-JH01
    |    |--C. magnus D07
    |    `--C. minor D07
    `--+--Boffius Vianey-Liaud 1979 [Boffiidae] K-JH01
       |--Ptilodontoidea K-JH01
       `--Taeniolabidoidea K-JH97
            |--Cimolomyidae K-JH01
            |    |--Meniscoessus Cope 1882 K-JH01
            |    |--Cimolomys Marsh 1889 K-JH01
            |    |--Essonodon Simpson 1927 K-JH01
            |    `--Buginbaatar Kielan-Jaworowska & Sochava 1969 [Buginbaatarinae] K-JH01
            |         `--B. transaltaiensis Kielan-Jaworowska & Sochava 1969 K-JH97
            `--+--Djadochtatherioidea K-JH01
               |--Kogaionidae K-JH01
               |    |--Hainina Vianey-Liaud 1979 K-JH01
               |    `--Kogaionon Râdulescu & Samson 1996 K-JH01
               |--Eucosmodontidae K-JH01
               |    |--Eucosmodon Matthew & Granger 1921 K-JH01
               |    |--Stygimys Sloan & van Valen 1965 K-JH01
               |    `--Clemensodon Krause 1992 K-JH01
               |--Microcosmodontidae K-JH01
               |    |--Microcosmodon Jepsen 1930 K-JH01
               |    |--Pentacosmodon Jepsen 1940 K-JH01
               |    `--Acheronodon Archibald 1982 K-JH01
               `--Taeniolabididae K-JH01
                    |--Prionessus Matthew & Granger 1925 K-JH01
                    |--Sphenopsalis Matthew, Granger & Simpson 1928 K-JH01
                    |--Lambdopsalis Chow & Qi 1978 K-JH01
                    |    `--L. bulla K-JH97
                    |--Catopsalis Cope 1882 K-JH01
                    |    `--C. joyneri K-JH01
                    `--Taeniolabis Cope 1882 K-JH01
                         |--T. lamberti D07
                         |--T. sulcatus D07
                         `--T. taoensis K-JH97

*Type species of generic name indicated

REFERENCES

[D07] Dixon, D. 2007. The Complete Illustrated Encyclopedia of Dinosaurs & Prehistoric Creatures. Hermes House: London.

[K-JH97] Kielan-Jaworowska, Z., & J. H. Hurum. 1997. Djadochtatheria - a new suborder of multituberculate mammals. Acta Palaeontologica Polonica 42 (2): 201–242.

[K-JH01] Kielan-Jaworowska, Z., & J. H. Hurum. 2001. Phylogeny and systematics of multituberculate mammals. Palaeontology 44 (3): 389–429.

Eutriconodonta

Reconstruction of Repenomamus giganticus, copyright Nobu Tamura.

Belongs within: Mammalia.

The Eutriconodonta are a group of mammals known from the Jurassic and Cretaceous, characterised by triconodont molars with three laterally compressed cusps in a straight alignment (Luo et al. 2007).

Synapomorphies (from Krause et al. 2014): Dens not fused to axis; lumbar ribs not fused to vertebrae; tarsal spur absent; premaxilla with ventral internarial process present but not connected to nasal; single infraorbital foramen present; ventral opening of minor palatine foramen encircled by palatine and maxilla; postpalatine torus present; internal acoustic meatus cribriform plate absent; ear region with lateral flange narrowly separated from anterior paroccipital process (crista parotica); pterygoparoccipital foramen a laterally open notch; mallear manubrium robust; suture separating basioccipital and exoccipital absent; two lower and upper incisors in each quadrant; I3 with single cusp; seven lower postcanine teeth in each quadrant; four lower premolars in each quadrant; ultimate upper premolar (P5) with metacone present; penultimate lower premolar (p4) with paraconid (cusp b) absent or indistinct; ultimate lower premolar tooth (p5) with hypoconid absent; m1–2 with paracristid longitudinally oriented relative to longitudinal axis of molars; m1–2 with buccal curvature of protoconid (cusp a) at base level equivalent to curvature of paraconid (cusp b) and metaconid (cusp c); m1–2 with metacristid (protocristid), crest between protoconid (cusp a) and metaconid (cusp c) parallel to long axis of lower molars and lower jaw axis; m1 with main cusps of trigonid in single longitudinal row; m1–2 with labiolingual compression of primary functional cusps (at level of cusp base but above cingulid); M1–2 with stylar shelf absent; M1–2 with lingual cingulum discontinuous; dentary movement during power stroke of chewing cycle essentially dorsal (orthal); molar with prevallum/postvallid shearing absent.

<==Eutriconodonta
    |  i. s.: Phascolotherium Owen 1838 CM99
    |         ‘Kotatherium’ yadagirii Prasad & Manhas 1997 A02
    |--+--Amphilestes [Amphilestinae] KH14
    |  `--Gobiconodontidae LK-JC02
    |       |--Gobiconodon Trofimov 1978 KH14, CM99
    |       |    `--G. zofiae HM05
    |       `--Repenomamus Li, Wang et al. 2000 LC07, HM05 [Repenomamidae]
    |            |--R. giganticus Hu, Meng et al. 2005 HM05
    |            `--R. robustus HM05
    `--+--+--Jeholodens jenkinsi KH14, LC07
       |  `--Yanoconodon Luo, Chen et al. 2007 LC07
       |       `--*Y. allini Luo, Chen et al. 2007 LC07
       `--Triconodontidae [Triconodontinae] LK-JC02
            |--Priacodon KH14
            |--Trioracodon LK-JC02
            `--Astroconodon LK-JC02

*Type species of generic name indicated

REFERENCES

[A02] Averianov, A. O. 2002. Early Cretaceous “symmetrodont” mammal Gobiotheriodon from Mongolia and the classification of “Symmetrodonta”. Acta Palaeontologica Polonica 47 (4): 705–716.

[CM99] Cifelli, R. L., & S. K. Madsen. 1999. Spalacotheriid symmetrodonts (Mammalia) from the medial Cretaceous (upper Albian or lower Cenomanian) Mussentuchit local fauna, Cedar Mountain Formation, Utah, USA. Geodiversitas 21 (2): 167–214.

[HM05] Hu, Y., J. Meng, Y. Wang & C. Li. 2005. Large Mesozoic mammals fed on young dinosaurs. Nature 433: 149–152.

[KH14] Krause, D. W., S. Hoffmann, J. R. Wible, E. C. Kirk, J. A. Schultz, W. von Koenigswald, J. R. Groenke, J. B. Rossie, P. M. O'Connor, E. R. Seiffert, E. R. Dumont, W. L. Holloway, R. R. Rogers, L. J. Rahantarisoa, A. D. Kemp & H. Andriamialison. 2014. First cranial remains of a gondwanatherian mammal reveal remarkable mosaicism. Nature 515: 512–517.

[LC07] Luo, Z.-X., P. Chen, G. Li & M. Chen. 2007. A new eutriconodont mammal and evolutionary development in early mammals. Nature 446: 288–293.

[LK-JC02] Luo, Z.-X., Z. Kielan-Jaworowska & R. L. Cifelli. 2002. In quest for a phylogeny of Mesozoic mammals. Acta Palaeontologica Polonica 47: 1–78.

Dasyurinae

Tiger quolls Dasyurus maculatus, copyright J.-L. Klein & M.-L. Hubert.

Belongs within: Dasyuridae.

The Dasyurinae is a group of carnivorous marsupials including the quolls (marsupial cats), mulgaras and related taxa. Members of this group include the largest living carnivorous marsupials.

Dasyurinae
    |--+--Dasykaluta rosamondae CB-E04
    |  `--Parantechinus Tate 1947 CB-E04, R64
    |       `--*P. apicalis [=Phascogale apicalis] R64
    |--+--‘Pseudantechinus’ ningbing CB-E04
    |  `--Myoictis Gray 1858 CB-E04, LA02
    |       `--*M. melas (Müller 1840) LA02
    `--+--Pseudantechinus Tate 1947 CB-E04, R64
       |    |--*P. macdonnellensis [=Phascogale macdonnellensis] R64
       |    |--P. roryi CWS05
       |    `--P. woolleyae CB-E04
       |--Dasycercus Peters 1875 CB-E04, LA02
       |    |--*D. cristicauda (Krefft 1867) LA02
       |    |--D. blythi TL70
       |    |--D. byrnei CB-E04
       |    |--D. hillieri LA02
       |    `--D. worboysi Dawson, Muirhead & Wroe 1999 LA02
       `--+--‘Antechinus’ bilarni Johnson 1954 R64 [=Parantechinus bilarni CB-E04]
          `--+--+--Neophascogale lorentzi CB-E04
             |  `--Phascolosorex CB-E04
             |       |--P. doriae CB-E04
             |       `--P. dorsalis CB-E04
             `--+--Sarcophilus Cuvier 1837 CB-E04, LA02 [Sarcophilini]
                |    |--*S. laniarius (Owen 1838) LA02
                |    |--S. harrisii (Boitard 1841) K92 [incl. S. ursinus S61]
                |    `--S. moornaensis Crabb 1982 LA02
                `--Dasyurus Geoffroy 1796 CB-E04, LA02
                     |  i. s.: D. bowlingi A14
                     |--D. hallucatus CB-E04 [=Satanellus hallucatus TL70]
                     `--+--+--*D. maculatus (Kerr 1792) LA02, CB-E04, LA02 [=Viverra maculata G91]
                        |  `--D. dunmalli Bartholomai 1971 LA02
                        `--+--D. viverrinus (Shaw 1800) CB-E04, R64 (see below for synonymy)
                           `--+--D. albopunctatus CB-E04
                              `--+--D. geoffroii Gould 1841 CB-E04, BD-D09
                                 `--D. spartacus CB-E04

Dasyurinae incertae sedis:
  Dasyurops D02
  Dasyurinus D02
  Archerium Wroe & Mackness 2000 LA02
    `--*A. chinchillaensis Wroe & Mackness 2000 LA02
  Dasyuroides Spencer 1896 LA02
    |--*D. byrnei Spencer 1896 LA02
    `--D. achilpatna Archer 1982 LA02
  Ganbulanyi Wroe 1998 LA02
    `--*G. djadjinguli Wroe 1998 LA02
  Glaucodon Stirton 1957 LA02
    `--*G. ballaratensis Stirton 1957 LA02

Dasyurus viverrinus (Shaw 1800) CB-E04, R64 [=Didelphis viverrina R64; incl. Mustela quoll Zimmermann 1777 (nom. inv.) R64, Dasyurus quoll R64]

*Type species of generic name indicated

REFERENCES

[A14] Anderson, W. 1914. Notes on the occurrence of the sand-rock containing bones of extinct species of marsupials (emu, kangaroo, wombat, etc.) on King Island, Bass Strait, Tasmania. Records of the Australian Museum 10 (9): 275-283.

[BD-D09] Beveridge, I. & M.-C. Durette-Desset. 2009. A new species of the nematode genus Copemania (Nematoda: Trichostrongylida), parasitic in the western quoll, Dasyurus geoffroii and short-nosed bandicoot, Isoodon obesulus from south-western Western Australia. Records of the Western Australian Museum 25 (3): 345-349.

[CB-E04] Cardillo, M., O. R. P. Bininda-Emonds, E. Boakes & A. Purvis. 2004. A species-level phylogenetic supertree of marsupials. Journal of Zoology 264: 11-31.

[CWS05] Cooper, N. K., P. C. Withers & T. Stewart. 2005. Description of sternal glands in Western Australian Sminthopsis and Ningaui (Marsupialia: Dasyuridae). Journal of the Royal Society of Western Australia 88 (2): 45-49.

[D02] Dyck, S. van. 2002. Morphology-based revision of Murexia and Antechinus (Marsupialia: Dasyuridae). Memoirs of the Queensland Museum 48: 239-330.

[G91] Groves, C. P. 1991. A Theory of Human and Primate Evolution, revised ed. Clarendon Press: Oxford.

[K92] Klompen, J. S. H. 1992. Phylogenetic relationships in the mite family Sarcoptidae (Acari: Astigmata). Miscellaneous Publications, Museum of Zoology, University of Michigan 180: i-vi, 1-154.

[LA02] Long, J., M. Archer, T. Flannery & S. Hand. 2002. Prehistoric Mammals of Australia and New Guinea: One Hundred Million Years of Evolution. University of New South Wales Press: Sydney.

[R64] Ride, W. D. L. 1964. A list of mammals described from Australia between the years 1933 and 1963 (comprising newly proposed names and additions to the Australian faunal list). Australian Mammal Society Bulletin 7 (Suppl.): 1-15.

[S61] Sharman, G. B. 1961. The mitotic chromosomes of marsupials and their bearing on taxonomy and phylogeny. Australian Journal of Zoology 9 (1): 38-60.

[TL70] Turnbull, W. D. & E. L. Lundelius, Jr. 1970. The Hamilton fauna: a Late Pliocene mammalian fauna from the Grange Burn, Victoria, Australia. Fieldiana Geology 19: 1-163.

Macropodini

Yellow-footed rock wallaby Petrogale xanthopus, copyright Talexbikes.

Belongs within: Macropodinae.
Contains: Macropus.

The Macropodini is a clade including the 'true' wallabies and kangaroos. Members of this clade have high-crowned molars, reflecting a change in habit to grazing.

<==Macropodini
    |  i. s.: Baringa Flannery & Hann 1984 LA02
    |           `--*B. nelsonensis Flannery & Hann 1984 LA02
    |--+--Lagorchestes CB-E04
    |  |    |--L. asomatus Finlayson 1943 R64
    |  |    |--L. conspicillatus CB-E04
    |  |    |--L. hirsutus CB-E04
    |  |    `--L. leporides CB-E04
    |  `--+--Setonix brachyurus (Quoy & Gaimard 1830) CB-E04, BD-D09
    |     `--+--Onychogalea CB-E04
    |        |    |--O. unguifera CB-E04
    |        |    `--+--O. fraenata CB-E04
    |        |       `--O. lunata CB-E04
    |        `--+--Macropus CB-E04
    |           |--Kurrabi Flannery & Archer 1984 LA02
    |           |    |--*K. mahoneyi Flannery & Archer 1984 LA02
    |           |    |--K. merriwaensis Flannery & Archer 1984 LA02
    |           |    `--K. pelchenorum Flannery, Rich et al. 1992 LA02
    |           `--Wallabia Trouessart 1905 CB-E04, LA02
    |                |--*W. bicolor (Desmarest 1804) LA02 [=Protemnodon bicolor S61; incl. Macropus ualabatus S61]
    |                |--W. eugenii TL70
    |                |--W. indra (De Vis 1895) LA02
    |                `--W. kitcheneri Flannery 1989 LA02
    `--+--Thylogale Gray 1837 LA02
       |    |  i. s.: T. eugenii R64 [=Protemnodon eugenii S61]
       |    |           |--T. e. eugenii R64
       |    |           `--T. e. decres Troughton 1941 R64
       |    |--*T. billardieri (Desmarest 1822) LA02, CB-E04 [=Protemnodon billardierii S61]
       |    |--T. ignis Flannery, Rich et al. 1992 LA02
       |    |--T. christenseni Hope 1981 LA02
       |    `--+--T. brunii CB-E04
       |       `--+--T. stigmatica CB-E04
       |          `--T. thetis CB-E04
       `--Petrogale CB-E04
            |  i. s.: P. herberti Thomas 1926 CO03
            |         P. sharmani COA02
            |--+--P. brachyotis CB-E04
            |  `--+--P. burbidgei CB-E04
            |     `--P. concinna CB-E04 [=Peradorcas concinna BP87]
            `--+--P. lateralis Gould 1842 CB-E04, S11
               |--P. persephone CB-E04
               |--P. rothschildi CB-E04
               |--P. xanthopus CB-E04
               `--+--P. inornata CB-E04
                  |--P. penicillata CB-E04
                  |    |--P. p. penicillata S61
                  |    `--P. p. pearsoni S61
                  `--+--P. assimilis CB-E04
                     `--P. godmani CB-E04

*Type species of generic name indicated

REFERENCES

[BD-D09] Beveridge, I. & M.-C. Durette-Desset. 2009. A new species of the nematode genus Copemania (Nematoda: Trichostrongylida), parasitic in the western quoll, Dasyurus geoffroii and short-nosed bandicoot, Isoodon obesulus from south-western Western Australia. Records of the Western Australian Museum 25 (3): 345-349.

[BP87] Burton, J. A., & B. Pearson. 1987. Collins Guide to the Rare Mammals of the World. Collins: London.

[CO03] Cameron, S. L., & P. J. O’Donoghue. 2003. Trichostome ciliates from Australian marsupials. II. Polycosta gen. nov. (Litostomatea: Polycostidae fam. nov.) European Journal of Protistology 39: 83-99.

[COA02] Cameron, S. L., P. J. O’Donoghue & R. D. Adlard. 2002. Species diversity within Macropodinium (Litostomatea: Trichostomatia): endosymbiotic ciliates from Australian macropodid marsupials. Memoirs of the Queensland Museum 48 (1): 49-69.

[CB-E04] Cardillo, M., O. R. P. Bininda-Emonds, E. Boakes & A. Purvis. 2004. A species-level phylogenetic supertree of marsupials. Journal of Zoology 264: 11-31.

[LA02] Long, J., M. Archer, T. Flannery & S. Hand. 2002. Prehistoric Mammals of Australia and New Guinea: One Hundred Million Years of Evolution. University of New South Wales Press: Sydney.

[R64] Ride, W. D. L. 1964. A list of mammals described from Australia between the years 1933 and 1963 (comprising newly proposed names and additions to the Australian faunal list). Australian Mammal Society Bulletin 7 (Suppl.): 1-15.

[S61] Sharman, G. B. 1961. The mitotic chromosomes of marsupials and their bearing on taxonomy and phylogeny. Australian Journal of Zoology 9 (1): 38-60.

[S11] Smales, L. R. 2011. New species and new locality records of the nematode genus Labiosimplex (Strongylida: Chabertiidae) from macropodid marsupials in Western Australia. Records of the Western Australian Museum 26 (2): 183-190.

[TL70] Turnbull, W. D. & E. L. Lundelius, Jr. 1970. The Hamilton fauna: a Late Pliocene mammalian fauna from the Grange Burn, Victoria, Australia. Fieldiana Geology 19: 1-163.

Peramelemorphia

New Guinea spiny bandicoot Echymipera kalubu, photographed by Michael Pennay.

Belongs within: Marsupialia.

The Peramelemorphia includes the bandicoots, a group of long-snouted Australian marsupials. Most species are insectivorous or carnivorous, though the now-extinct pig-footed bandicoot Chaeropus ecaudatus may have been a grazer (Long et al. 2002.

Characters (from Long et al. 2002): Lower molars with very large, un-notched anterior cingulid; upper molars with large adjacent stylar cusps C and D, and buccally shifted centrocrista. Hind feet syndactylous (at least in Perameloidea), with second and third toes reduced and joined by a web of skin).

<==Peramelemorphia
    |--Yarala Muirhead & Filan 1995 [Yaralidae, Yaraloidea] LA02
    |    |--*Y. burchfieldi Muirhead & Filan 1995 LA02
    |    `--Y. kida MJ11
    `--Perameloidea [Peramelidae] LA02
         |--Chaeropus ecaudatus CB-E04
         `--+--Thylacomyidae LA02
            |    |--Ischnodon Stirton 1955 LA02
            |    |    `--*I. australis Stirton 1955 LA02
            |    |--Macrotis CB-E04
            |    |    |--M. lagotis CB-E04
            |    |    `--M. leucura CB-E04
            |    `--Thylacomys nigripes WJ26
            `--+--+--Isoodon CB-E04
               |  |    |--I. macrourus (Gould 1842) CB-E04, K92
               |  |    `--+--I. auratus CB-E04
               |  |       `--I. obesulus (Shaw 1797) CB-E04, BD-D09
               |  `--Perameles Geoffroy 1803 CB-E04, LA02
               |       |  i. s.: P. allinghamensis Archer 1976 LA02
               |       |         P. bowensis Muirhead, Dawson & Archer 1997 LA02
               |       |         P. eremiana CB-E04
               |       |         P. wombeyensis Broom 1896 F71
               |       |--P. bougainville CB-E04
               |       `--+--*P. nasuta Geoffroy 1804 LA02, CB-E04, LA02
               |          `--P. gunnii CB-E04
               `--Peroryctidae LA02
                    |--Rhynchomeles prattorum LA02, BP87
                    |--Peroryctes CB-E04
                    |    |--P. broadbenti CB-E04
                    |    |--P. longicaudata TL70
                    |    `--P. raffrayana CB-E04
                    |--Microperoryctes LA02
                    |    |--M. longicauda CB-E04
                    |    |--M. murina CB-E04
                    |    `--M. papuensis CB-E04
                    `--Echymipera Lesson 1842 LA02
                         |--*E. kalubu (Lesson 1828) [=Perameles kalubu] R64
                         |--E. clara CB-E04
                         |--E. davidi CB-E04
                         |--E. doreyana TL70
                         |--E. echinista CB-E04
                         `--E. rufescens CB-E04
                              |--E. r. rufescens R64
                              `--E. r. australis Tate 1948 R64

*Type species of generic name indicated

REFERENCES

[BD-D09] Beveridge, I. & M.-C. Durette-Desset. 2009. A new species of the nematode genus Copemania (Nematoda: Trichostrongylida), parasitic in the western quoll, Dasyurus geoffroii and short-nosed bandicoot, Isoodon obesulus from south-western Western Australia. Records of the Western Australian Museum 25 (3): 345-349.

[BP87] Burton, J. A., & B. Pearson. 1987. Collins Guide to the Rare Mammals of the World. Collins: London.

[CB-E04] Cardillo, M., O. R. P. Bininda-Emonds, E. Boakes & A. Purvis. 2004. A species-level phylogenetic supertree of marsupials. Journal of Zoology 264: 11-31.

[F71] Fletcher, H. O. 1971. Catalogue of type specimens of fossils in the Australian Museum, Sydney. Australian Museum Memoir 13: 1-167.

[K92] Klompen, J. S. H. 1992. Phylogenetic relationships in the mite family Sarcoptidae (Acari: Astigmata). Miscellaneous Publications, Museum of Zoology, University of Michigan 180: i-vi, 1-154.

[LA02] Long, J., M. Archer, T. Flannery & S. Hand. 2002. Prehistoric Mammals of Australia and New Guinea: One Hundred Million Years of Evolution. University of New South Wales Press: Sydney.

[MJ11] Meredith, R. W., J. E. Janečka, J. Gatesy, O. A. Ryder, C. A. Fisher, E. C. Teeling, A. Goodbla, E. Eizirik, T. L. L. Simão, T. Stadler, D. L. Rabosky, R. L. Honeycutt, J. J. Flynn, C. M. Ingram, C. Steiner, T. L. Williams, T. J. Robinson, A. Burk-Herrick, M. Westerman, N. A. Ayoub, M. S. Springer & W. J. Murphy. 2011. Impacts of the Cretaceous terrestrial revolution and KPg extinction on mammal diversification. Science 334: 521-524.

[R64] Ride, W. D. L. 1964. A list of mammals described from Australia between the years 1933 and 1963 (comprising newly proposed names and additions to the Australian faunal list). Australian Mammal Society Bulletin 7 (Suppl.): 1-15.

[TL70] Turnbull, W. D. & E. L. Lundelius, Jr. 1970. The Hamilton fauna: a Late Pliocene mammalian fauna from the Grange Burn, Victoria, Australia. Fieldiana Geology 19: 1-163.

[WJ26] Wood Jones, F. 1926. The R. M. Johnston Memorial Lecture, 1925. The mammalian toilet and some considerations arising from it. Papers and Proceedings of the Royal Society of Tasmania 1925: 14-62.

Vombatomorphia

Southern hairy-nosed wombat Lasiorhinus latifrons, from Blanchetown Allsorts.

Belongs within: Diprotodontia.
Contains: Diprotodontidae.

The Vombatomorphia include the living wombats (Vombatidae) and their fossil relatives. These include such diverse forms as the marsupial lions (Thylacoleonidae), the rhinoceros-like Diprotodontidae, and the large clawed herbivores of the Palorchestidae.

Vombatomorphia
    |--Thylacoleonidae [Wakaleoninae] LA02
    |    |--Priscileo Rauscher 1987 LA02
    |    |    |--*P. pitikantensis Rauscher 1987 LA02
    |    |    `--P. roskellyae Gillespie 1997 LA02
    |    `--+--Wakaleo Clemens & Plane 1974 LA02
    |       |    |--*W. vanderleueri Clemens & Plane 1974 LA02
    |       |    |--W. alcootaensis Archer & Rich 1982 LA02
    |       |    `--W. oldfieldi Clemens & Plane 1974 LA02
    |       `--Thylacoleo Gervais 1852 [Thylacoleoninae] LA02
    |            |--*T. carnifex Owen 1858 LA02
    |            |--T. crassidentatus Bartholomai 1962 LA02
    |            |--T. hilli Pledge 1977 LA02
    |            |--T. oweni McCoy 1876 F71
    |            `--T. robustus Krefft 1872 F71
    `--+--Ilariidae LA02
       |    |--Kuterintja Pledge 1987 LA02
       |    |    `--*K. ngama Pledge 1987 LA02
       |    `--Ilaria Tedford & Woodburne 1987 LA02
       |         |--*I. illumidens Tedford & Woodburne 1987 LA02
       |         `--I. lawsoni Tedford & Woodburne 1987 LA02
       |--+--Wynyardiidae LA02
       |  |    |--Muramura Pledge 1987 LA02
       |  |    |    `--*M. williamsi Pledge 1987 LA02
       |  |    |--Namilamadeta Rich & Archer 1979 LA02
       |  |    |    `--*N. snideri Rich & Archer 1979 LA02
       |  |    `--Wynyardia Spencer 1901 LA02
       |  |         `--*W. bassiana Spencer 1901 LA02
       |  `--Diprotodontoidea LA02
       |       |--Diprotodontidae LA02
       |       `--Palorchestidae LA02
       |            |--Propalorchestes Murray 1986 LA02
       |            |    |--*P. novaculacephalus Murray 1986 LA02
       |            |    `--P. ponticulus Murray 1990 LA02
       |            `--Palorchestes Owen 1873 LA02
       |                 |--*P. azael Owen 1873 LA02
       |                 |--P. anulus Black 1997 LA02
       |                 |--P. painei Woodburne 1967 LA02
       |                 |--P. parvus De Vis 1895 LA02
       |                 |--P. rephaim Ramsay 1885 F71
       |                 `--P. selestiae Mackness 1995 LA02
       `--Vombatidae [Phascolomidae, Vombatoidea] LA02
            |  i. s.: Wombatula Troughton 1941 R64
            |           `--*W. gillespiei (De Vis 1900) [=Phascolomys gillespiei] R64
            |--Rhizophascolonus Stirton, Tedford & Woodburne 1967 LA02
            |    `--*R. crowcrofti Stirton, Tedford & Woodburne 1967 LA02
            `--+--Warendja Hope & Wilkinson 1982 LA02
               |    `--*W. wakefieldi Hope & Wilkinson 1982 LA02
               `--+--Vombatus Geoffroy 1803 LA02
                  |    |--*V. ursinus (Shaw 1800) LA02
                  |    `--V. hacketti (Glauert 1910) LA02
                  `--+--Ramsayia Tate 1951 LA02
                     |    |--*R. magna (Owen 1872) LA02
                     |    `--R. lemleyi (Archer 1976) LA02
                     `--+--Lasiorhinus Gray 1863 LA02
                        |    |--*L. latifrons (Owen 1845) LA02
                        |    |    |--L. l. latifrons R64
                        |    |    `--L. l. barnardi Longman 1939 R64
                        |    |--L. angustidens (De Vis 1891) LA02
                        |    |--L. gillespiei USDI77
                        |    `--L. krefftii CB-E04
                        `--+--Phascolonus Owen 1872 LA02
                           |    `--*P. gigas (Owen 1858) LA02 [=Phascolomys (Phascolonus) gigas F71]
                           `--Phascolomys LA02
                                |--P. curvirostris Owen 1885 F71
                                |--P. medius Owen 1872 LA02
                                |--P. ursinus A14
                                `--‘Phascolomis’ vombatus Leach 1815 R64

*Type species of generic name indicated

REFERENCES

[A14] Anderson, W. 1914. Notes on the occurrence of the sand-rock containing bones of extinct species of marsupials (emu, kangaroo, wombat, etc.) on King Island, Bass Strait, Tasmania. Records of the Australian Museum 10 (9): 275-283.

[CB-E04] Cardillo, M., O. R. P. Bininda-Emonds, E. Boakes & A. Purvis. 2004. A species-level phylogenetic supertree of marsupials. Journal of Zoology 264: 11-31.

[F71] Fletcher, H. O. 1971. Catalogue of type specimens of fossils in the Australian Museum, Sydney. Australian Museum Memoir 13: 1-167.

[LA02] Long, J., M. Archer, T. Flannery & S. Hand. 2002. Prehistoric Mammals of Australia and New Guinea: One Hundred Million Years of Evolution. University of New South Wales Press: Sydney.

[R64] Ride, W. D. L. 1964. A list of mammals described from Australia between the years 1933 and 1963 (comprising newly proposed names and additions to the Australian faunal list). Australian Mammal Society Bulletin 7 (Suppl.): 1-15.

[USDI77] USDI (United States Department of the Interior). 1977. Endangered and threatened wildlife and plants—republication of list of species. Federal Register 42: 36420-36431.

Dasyuromorphia

Thylacine Thylacinus cynocephalus, from here.

Belongs within: Marsupialia.
Contains: Dasyuridae.

The Dasyuromorphia is a group of Australian carnivorous marsupials including the marsupial mice and cats (Dasyuridae), numbat Myrmecobius fasciatus and thylacines (Thylacinidae).

Characters (from Long et al. 2002): Small number of incisors (I⁴/₃); intestinal caecum absent.

<==Dasyuromorphia [Dasyura, Dasyurini, Dasyuroidea]
    |  i. s.: Ankotarinja Archer 1976 LA02
    |           `--*A. tirarensis Archer 1976 LA02
    |         Apoktesis Campbell 1976 LA02
    |           `--*A. cuspis Campbell 1976 LA02
    |         Dasylurinja Archer 1982 LA02
    |           `--*D. kokuminola Archer 1982 LA02
    |         Keeuna Archer 1976 LA02
    |           `--*K. woodburnei Archer 1976 LA02
    |         Mayigriphus Wroe 1997 LA02
    |           `--*M. orbus Wroe 1997 LA02
    |         Wakamartha Archer & Rich 1979 LA02
    |           `--*W. tasselli Archer & Rich 1979 LA02
    |--Djarthia Godthelp, Wroe & Archer 1999 LA02
    |    `--*D. murgonensis Godthelp, Wroe & Archer 1999 LA02
    `--+--+--Dasyuridae CB-E04
       |  `--Myrmecobius [Myrmecobiidae, Myrmecobiinae] CB-E04
       |       `--M. fasciatus CB-E04
       |            |--M. f. fasciatus BP87
       |            `--M. f. rufus BP87
       `--Thylacinidae [Thylacininae] CB-E04
            |--Badjacinus Muirhead & Wroe 1998 LA02
            |    `--*B. turnbulli Muirhead & Wroe 1998 LA02
            |--Maximucinus Wroe 2001 LA02
            |    `--*M. muirheadae Wroe 2001 LA02
            |--Muribacinus Wroe 1996 LA02
            |    `--*M. gadiyuli Wroe 1996 LA02
            |--Mutpuracinus Murray & Megirian 2000 LA02
            |    `--*M. archibaldi Murray & Megirian 2000 LA02
            |--Nimbacinus Muirhead & Archer 1990 LA02
            |    |--*N. dicksoni Muirhead & Archer 1990 LA02
            |    `--N. richi Murray & Megirian 2000 LA02
            `--+--Ngamalacinus Muirhead 1997 LA02
               |    `--*N. timmulvaneyi Muirhead 1997 LA02
               `--+--Wabulacinus Muirhead 1997 LA02
                  |    `--*W. ridei Muirhead 1997 LA02
                  |--Tyarrpecinus Murray & Megirian 2000 LA02
                  |    `--*T. rothi Murray & Megirian 2000 LA02
                  `--Thylacinus Temminck 1824 LA02
                       |--*T. cynocephalus (Harris 1808) LA02
                       |--T. macknessi Muirhead 1992 LA02
                       |--T. megiriani Murray 1997 LA02
                       |--T. potens Woodburne 1967 LA02
                       `--T. spelaeus F71

Inorganic: Myrmecobius fasciatus protominilorientalus Okamura 1987 O87

*Type species of generic name indicated

REFERENCES

[BP87] Burton, J. A., & B. Pearson. 1987. Collins Guide to the Rare Mammals of the World. Collins: London.

[CB-E04] Cardillo, M., O. R. P. Bininda-Emonds, E. Boakes & A. Purvis. 2004. A species-level phylogenetic supertree of marsupials. Journal of Zoology 264: 11-31.

[F71] Fletcher, H. O. 1971. Catalogue of type specimens of fossils in the Australian Museum, Sydney. Australian Museum Memoir 13: 1-167.

[LA02] Long, J., M. Archer, T. Flannery & S. Hand. 2002. Prehistoric Mammals of Australia and New Guinea: One Hundred Million Years of Evolution. University of New South Wales Press: Sydney.

[O87] Okamura, C. 1987. New facts: Homo and all Vertebrata were born simultaneously in the former Paleozoic in Japan. Original Report of the Okamura Fossil Laboratory 15: 347-573.

Macropodiformes

Northern bettong Bettongia tropica, from the Queensland Government.

Belongs within: Diprotodontia.
Contains: Macropodidae.

The Macropodiformes are a group of mostly herbivorous marsupials (though the Hypsiprymnodontidae are omnivorous), including the wallabies and kangaroos. Members of this group have the hind feet enlarged for rapid, often bounding locomotion.

Characters (from Long et al. 2002, as Macropodoidea): Dentary bearing masseteric canal. Hind foot with fourth digit hypertrophied.

Macropodiformes [Potoroinae]
    |--Balbaridae [Balbarinae] LA02
    |    |--Wururoo Cooke 1997 LA02
    |    |    `--*W. dayamayi Cooke 1997 LA02
    |    |--Balbaroo Flannery, Archer & Plane 1983 LA02
    |    |    |--*B. camfieldensis Flannery, Archer & Plane 1983 LA02
    |    |    |--B. fangaroo Cooke 2000 LA02
    |    |    `--B. gregoriensis Flannery, Archer & Plane 1983 LA02
    |    |--Ganawamaya Cooke 1992 LA02
    |    |    |--*G. acris Cooke 1992 LA02
    |    |    |--G. aediculis Cooke 1992 LA02
    |    |    `--G. ornata Cooke 1992 LA02
    |    `--Nambaroo Flannery & Rich 1986 LA02
    |         |--*N. tarrinyeri Flannery & Rich 1986 LA02
    |         |--N. couperi Cooke 1997 LA02
    |         |--N. novus Flannery & Rich 1986 LA02
    |         `--N. saltavus Flannery & Rich 1986 LA02
    `--+--Hypsiprymnodontidae LA02
       |    |--Hypsiprymnodon Ramsay 1876 [Hypsiprymnodontinae] LA02
       |    |    |--*H. moschatus Ramsay 1876 [=Pleopus moschatus] LA02
       |    |    `--H. bartholomaii Flannery & Archer 1987 LA02
       |    `--Propleopinae LA02
       |         |--Jackmahoneya Ride 1993 LA02
       |         |    `--*J. toxoniensis Ride 1993 LA02
       |         |--Ekaltadeta Archer & Flannery 1985 LA02
       |         |    |--*E. ima Archer & Flannery 1985 LA02
       |         |    `--E. jamiemulvaneyi Wroe 1996 LA02
       |         `--Propleopus Longman 1924 LA02
       |              |--*P. oscillans (De Vis 1888) LA02 [=Triclis oscillans F71]
       |              |--P. chillagoensis Archer, Bartholomai & Marshall 1978 LA02
       |              `--P. wellingtonensis Archer & Flannery 1985 LA02
       `--Macropodoidea MJ11
            |--Macropodidae LA02
            `--Potoroidae LA02
                 |--Caloprymnus campestris LA02, CB-E04
                 |--Aepyprymnus rufescens LA02
                 |--Borungaboodie Prideaux 1999 LA02
                 |    `--*B. hatcheri Prideaux 1999 LA02
                 |--Milliyowi Flannery, Rich et al. 1992 LA02
                 |    `--*M. bunganditj Flannery, Rich et al. 1992 LA02
                 |--Potorous CB-E04
                 |    |--P. longipes CB-E04
                 |    |--P. morgani Finlayson 1938 R64
                 |    |--P. platyops CB-E04
                 |    `--P. tridactylus CB-E04
                 |         |--P. t. tridactylus F71
                 |         |--P. t. antiquus Broom 1896 F71
                 |         |--P. t. apicalis BP87
                 |         |--P. t. benormi Courtney 1963 R64
                 |         `--P. t. gilberti BP87
                 `--Bettongia Gray 1837 [Bettongini] LA02
                      |--*B. gaimardi (Desmarest 1822) LA02
                      |--B. anhydra Finlayson 1957 [=B. penicillata anhydra] B08
                      |--B. cuniculus S61
                      |--B. lesueur CB-E04
                      |    |--B. l. lesueur B32
                      |    `--B. l. grayi B32
                      |--B. moyesi Flannery & Archer 1987 LA02
                      |--B. penicillata B08
                      |    |--B. p. penicillata R64
                      |    `--B. p. francisca Finlayson 1957 R64
                      |--B. pusilla McNamara 1997 LA02
                      `--B. tropica BP87

Macropodiformes incertae sedis:
  Galanarla Flannery, Archer & Plane 1983 LA02
    `--*G. tessellata Flannery, Archer & Plane 1983 LA02
  Gumardee Flannery, Archer & Plane 1983 LA02
    `--*G. pascuali Flannery, Archer & Plane 1983 LA02
  Palaeopotorous Flannery & Rich 1986 LA02
    `--*P. priscus Flannery & Rich 1986 LA02
  Purtia Case 1984 LA02
    `--*P. mosaicus Case 1984 LA02
  Wakiewakie Woodburne 1984 LA02
    `--*W. lawsoni Woodburne 1984 LA02

*Type species of generic name indicated

REFERENCES

[B08] Baynes, A. 2008. The original non-volant land mammal fauna of Faure Island, Shark Bay, Western Australia. Records of the Western Australian Museum Supplement 75: 25-31.

[B32] Bourne, G. 1932. Sarcosporidia. Journal of the Royal Society of Western Australia 19: 1-8.

[BP87] Burton, J. A., & B. Pearson. 1987. Collins Guide to the Rare Mammals of the World. Collins: London.

[CB-E04] Cardillo, M., O. R. P. Bininda-Emonds, E. Boakes & A. Purvis. 2004. A species-level phylogenetic supertree of marsupials. Journal of Zoology 264: 11-31.

[F71] Fletcher, H. O. 1971. Catalogue of type specimens of fossils in the Australian Museum, Sydney. Australian Museum Memoir 13: 1-167.

[LA02] Long, J., M. Archer, T. Flannery & S. Hand. 2002. Prehistoric Mammals of Australia and New Guinea: One Hundred Million Years of Evolution. University of New South Wales Press: Sydney.

[MJ11] Meredith, R. W., J. E. Janečka, J. Gatesy, O. A. Ryder, C. A. Fisher, E. C. Teeling, A. Goodbla, E. Eizirik, T. L. L. Simão, T. Stadler, D. L. Rabosky, R. L. Honeycutt, J. J. Flynn, C. M. Ingram, C. Steiner, T. L. Williams, T. J. Robinson, A. Burk-Herrick, M. Westerman, N. A. Ayoub, M. S. Springer & W. J. Murphy. 2011. Impacts of the Cretaceous terrestrial revolution and KPg extinction on mammal diversification. Science 334: 521-524.

[R64] Ride, W. D. L. 1964. A list of mammals described from Australia between the years 1933 and 1963 (comprising newly proposed names and additions to the Australian faunal list). Australian Mammal Society Bulletin 7 (Suppl.): 1-15.

[S61] Sharman, G. B. 1961. The mitotic chromosomes of marsupials and their bearing on taxonomy and phylogeny. Australian Journal of Zoology 9 (1): 38-60.

Diprotodontidae

Mounted skeleton of Zygomaturus trilobus, from the Museum Victoria.

Belongs within: Vombatomorphia.

The Diprotodontidae are a group of sometimes large herbivorous marsupials known from the late Oligocene to the Pleistocene of Australia.

Characters (from Long et al. 2002): Medium-sized to large browsing quadrupeds. Dental formula I^1-3/₁, C¹/₀, P³/₃, M^1-4/_1-4; molars simple, transversely lophodont but lacking longitudinal links. Lower jaw with masseteric foramen absent.

<==Diprotodontidae [Nototheriidae]
    |--Diprotodontinae [Nototheriinae] LA02
    |    |  i. s.: Nototherium Owen 1845 (n. d.) LA02
    |    |           `--*N. inerme Owen 1845 (n. d.) LA02
    |    |--+--Pitikantia Stirton 1967 LA02
    |    |  |    `--*P. dailyi Stirton 1967 LA02
    |    |  `--Ngapakaldia Stirton 1967 LA02
    |    |       |--*N. tedfordi Stirton 1967 LA02
    |    |       `--N. bonythoni Stirton 1967 LA02
    |    `--+--Bematherium Tedford 1967 LA02
    |       |    `--*B. angulum Tedford 1967 LA02
    |       `--+--Pyramios Woodburne 1967 LA02
    |          |    `--*P. alcootense Woodburne 1967 LA02
    |          `--+--+--Meniscolophus Stirton 1955 LA02
    |             |  |    `--*M. mawsoni Stirton 1955 LA02
    |             |  `--Euowenia De Vis 1891 [=Owenia De Vis 1887 non Gray 1855] LA02
    |             |       |--*E. grata (De Vis 1887) [=*Owenia grata] LA02
    |             |       `--E. robusta De Vis 1891 LA02
    |             `--+--Euryzygoma Longman 1921 LA02
    |                |    `--*E. dunense (De Vis 1888) LA02 [=Nototherium dunense F71]
    |                `--Diprotodon Owen 1838 LA02
    |                     |--*D. optatum Owen 1838 LA02
    |                     |--D. annextans McCoy 1861 LA02
    |                     |--D. australis F71
    |                     |--D. bennettii Krefft 1873 F71
    |                     |--D. loderi Krefft 1873 LA02
    |                     `--D. minor Huxley 1862 LA02
    `--Zygomaturinae LA02
         |--Raemeotherium Rich, Archer & Tedford 1978 LA02
         |    `--*R. yatkolai Rich, Archer & Tedford 1978 LA02
         |--Silvabestius Black & Archer 1997 LA02
         |    |--*S. johnnilandi Black & Archer 1997 LA02
         |    `--S. michaelbirti Black & Archer 1997 LA02
         `--+--Alkwertatherium Murray 1990 LA02
            |    `--*A. webborum Murray 1990 LA02
            `--+--Nimbadon Hand, Archer et al. 1993 LA02
               |    |--*N. lavarackorum Hand, Archer et al. 1993 LA02
               |    `--N. whitelawi Hand, Archer et al. 1993 LA02
               `--+--+--Kolopsoides Plane 1967 LA02
                  |  |    `--*K. cultridens Plane 1967 LA02
                  |  `--Plaisiodon Woodburne 1967 LA02
                  |       `--*P. centralis Woodburne 1967 LA02
                  `--+--Neohelos Stirton 1967 LA02
                     |    |--*N. tirarensis Stirton 1967 LA02
                     |    |--N. scottorrorum (Hand, Archer et al. 1993) LA02
                     |    `--N. stirtoni Murray, Megirian et al. 2000 LA02
                     `--+--Kolopsis Woodburne 1967 LA02
                        |    |--*K. torus Woodburne 1967 LA02
                        |    |--K. rotundus Plane 1967 LA02
                        |    |--K. watutense (Anderson 1937) [=Nototherium watutense] LA02
                        |    `--K. yperus Murray, Megirian & Wells 1993 LA02
                        `--+--+--Hulitherium Flannery & Plane 1986 LA02
                           |  |    `--*H. tomasettii Flannery & Plane 1986 LA02
                           |  `--Maokopia Flannery 1992 LA02
                           |       `--*M. ronaldi Flannery 1992 LA02
                           `--Zygomaturus Macleay 1857 LA02
                                |--*Z. trilobus Macleay 1858 (see below for synonymy) LA02
                                |--Z. creedii Krefft 1873 F71
                                |--Z. gilli Stirton 1967 LA02
                                |--Z. keanei Stirton 1967 LA02
                                |--Z. macleayi Krefft 1921 LA02
                                `--Z. nimborensia Hardjasasmita 1985 LA02

*Zygomaturus trilobus Macleay 1858 [incl. Nototherium mitchelli Owen 1845, N. tasmanicum Scott 1911, N. tasmaniense Noetling 1912, N. victoriae Owen 1872] LA02

*Type species of generic name indicated

REFERENCES

[F71] Fletcher, H. O. 1971. Catalogue of type specimens of fossils in the Australian Museum, Sydney. Australian Museum Memoir 13: 1-167.

[LA02] Long, J., M. Archer, T. Flannery & S. Hand. 2002. Prehistoric Mammals of Australia and New Guinea: One Hundred Million Years of Evolution. University of New South Wales Press: Sydney.