Linyphiidae

Porrhomma pygmaeum, from Pavouci.


Belongs within: Araneoidea.
Contains: Lepthyphantes, Mughiphantes, Agyneta, Haplinis, Parafroneta, Linyphia, Laetesia, Diplocephalus, Erigone, Walckenaeria.

The Linyphiidae are a hyperdiverse family of small spiders, most of which build delicate sheet webs among vegetation. They include the Erigoninae, money spiders (sometimes treated as a separate family), very small spiders that are usually dark and shiny, and often have the frontal region of males modified with lobes, pits or setal fields (Jocqué & Dippenaar-Schoeman 2007). The Linyphiinae are united by the presence in males of the lamella characteristica, a sclerite of the embolic division of the pedipalp. Members of the Mynogleninae have characteristic cephalic pits just below the anterior lateral eyes, as well as numerous trichobothria on the metatarsi. The Micronetinae have a complex embolic division on the male pedipalp, often with the sperm duct dilated at the base of the embolus.

Characters (from Jocqué & Dippenaar-Schoeman 2007): Very small to small; eight eyes present; labium rebordered; chelicerae usually with stridulating file; numerous teeth, mostly three or four, on fang furrow; legs slender, provided with setae, especially on tibiae and metatarsi; autospasy at patella-tibia joint; tarsi cylindrical, with three tarsal claws; tarsus IV without comb; ecribellate; entelegyne; male palp with often U-shaped intersegmental paracymbium.

<==Linyphiidae
    |--Micronetinae TS06
    |    |--Lepthyphantes TS06
    |    |--Mughiphantes TS06
    |    |--Agyneta TS06
    |    |--Microneta Menge 1869 W76
    |    |    `--M. varia (Blackwall 1841) MKD01 [incl. M. nicholsoni V09]
    |    |--Ascetophantes Tanasevitch & Saaristo 2006 TS06
    |    |    `--*A. asceticus (Tanasevitch 1987) [=Lepthyphantes asceticus] TS06
    |    |--Fistulaphantes Tanasevitch & Saaristo 2006 TS06
    |    |    `--*F. canalis Tanasevitch & Saaristo 2006 TS06
    |    |--Spiralophantes Tanasevitch & Saaristo 2006 TS06
    |    |    `--*S. mirabilis Tanasevitch & Saaristo 2006 TS06
    |    |--Piniphantes Saaristo & Tanasevitch 1996 TS06
    |    |    `--P. himalayensis (Tanasevitch 1987) [=Lepthyphantes uzbekistanicus himalayensis] TS06
    |    |--Birgerius Saaristo 1973 MKD01
    |    |    `--*B. microps (Simon 1911) [=Centromerus microps] MKD01
    |    |--Claviphantes Tanasevitch & Saaristo 2006 TS06
    |    |    |--*C. bifurcatus (Tanasevitch 1987) [=Lepthyphantes bifurcatus] TS06
    |    |    `--C. bifurcatoides (Tanasevitch 1987) [=Lepthyphantes bifurcatoides] TS06
    |    |--Megalepthyphantes Wunderlich 1994 TS06
    |    |    |--M. nebulosoides (Wunderlich 1977) [=Lepthyphantes nebulosoides] TS06
    |    |    `--M. nebulosus (Sundevall 1830) TS06
    |    |--Martensinus Wunderlich 1973 TS06
    |    |    |--M. annulatus Wunderlich 1973 TS06
    |    |    `--M. micronetiformis Wunderlich 1973 TS06
    |    |--Indophantes Saaristo & Tanasevitch 2003 TS06
    |    |    |--I. agamus Tanasevitch & Saaristo 2006 TS06
    |    |    |--I. bengalensis Saaristo & Tanasevitch 2003 TS06
    |    |    `--I. digitulus (Thaler 1987) [=Lepthyphantes digitulus] TS06
    |    |--Anguliphantes Saaristo & Tanasevitch 1996 TS06
    |    |    |--A. dybowskii (P.-Cambridge 1873) MKD01
    |    |    |--A. karpinskii (P.-Cambridge 1873) MKD01
    |    |    `--A. nepalensis (Tanasevitch 1987) [=Lepthyphantes nepalensis] TS06
    |    |--Palliduphantes Saaristo & Tanasevitch 2001 TS06
    |    |    |--P. ericaeus (Blackwall 1853) BBM02
    |    |    |--P. pallidus (Cambridge 1871) BBM02
    |    |    `--P. theosophicus (Tanasevitch 1987) [=Lepthyphantes theosophicus] TS06
    |    |--Himalaphantes Tanasevitch 1992 TS06
    |    |    |--H. grandiculus (Tanasevitch 1987) [=Lepthyphantes grandiculus] TS06
    |    |    |--H. magnus (Tanasevitch 1987) [=Lepthyphantes magnus] TS06
    |    |    `--H. martensi (Thaler 1987) [=Lepthyphantes martensi] TS06
    |    `--Tenuiphantes Saaristo & Tanasevitch 1996 TS06
    |         |--T. alacris (Blackwall 1853) MKD01
    |         |--T. altimontanus Tanasevitch & Saaristo 2006 TS06
    |         |--T. crassus Tanasevitch & Saaristo 2006 TS06
    |         |--T. mengei (Kulczyński 1887) MKD01
    |         |--T. plumipes (Tanasevitch 1987) [=Lepthyphantes plumipes] TS06
    |         |--T. tenuis (Blackwall 1852) PVD10
    |         `--T. zimmermanni (Betkau 1890) BBM02
    |--Mynogleninae PVD10
    |    |--Haplinis PVD10
    |    |--Parafroneta PVD10
    |    |--Cassafroneta Blest 1979 PVD10
    |    |    `--*C. forsteri Blest 1979 PVD10
    |    |--Hyperafroneta Blest 1979 PVD10
    |    |    `--*H. obscura Blest 1979 PVD10
    |    |--Poecilafroneta Blest 1979 PVD10
    |    |    `--*P. caudata Blest 1979 PVD10
    |    |--Protoerigone Blest 1979 PVD10
    |    |    |--*P. otagoa Blest 1979 PVD10
    |    |    `--P. obtusa Blest 1979 PVD10
    |    |--Megafroneta Blest 1979 PVD10
    |    |    |--*M. elongata Blest 1979 PVD10
    |    |    |--M. dugdaleae Blest & Vink 2002 PVD10
    |    |    `--M. gigas Blest 1979 PVD10
    |    |--Metafroneta Blest 1979 PVD10
    |    |    |--*M. sinuosa Blest 1979 PVD10
    |    |    |--M. minima Blest 1979 PVD10
    |    |    `--M. subversa Blest & Vink 2002 PVD10
    |    |--Mynoglenes Simon 1905 [incl. Afroneta Holm 1968, Paralinyphia Bryant 1933] W76
    |    |    |--M. diloris M83
    |    |    |--M. insolens Simon 1905 F64 [incl. Linyphia antipodiana Berland 1925 H39]
    |    |    |--M. major FF99
    |    |    |--M. marrineri Hogg 1909 W76
    |    |    |--M. simplicipalpis Wunderlich 1976 W76
    |    |    `--M. titan FF99
    |    |--Novafroneta Blest 1979 PVD10
    |    |    |--*N. vulgaris Blest 1979 PVD10
    |    |    |--N. annulipes Blest 1979 PVD10
    |    |    |--N. gladiatrix Blest 1979 PVD10
    |    |    |--N. nova Blest & Vink 2003 PVD10
    |    |    |--N. parmulata Blest 1979 PVD10
    |    |    `--N. truncata Blest & Vink 2003 PVD10
    |    |--Promynoglenes Blest 1979 PVD10
    |    |    |--*P. nobilis Blest 1979 PVD10
    |    |    |--P. grandis Blest 1979 PVD10
    |    |    |--P. minuscula Blest & Vink 2003 PVD10
    |    |    |--P. minuta Blest & Vink 2002 PVD10
    |    |    |--P. parvula Blest 1979 PVD10
    |    |    `--P. silvestris Blest 1979 PVD10
    |    |--Pseudafroneta Blest 1979 PVD10
    |    |    |--*P. incerta (Bryant 1935) PVD10
    |    |    |--P. frigida Blest 1979 PVD10
    |    |    |--P. lineata Blest 1979 PVD10
    |    |    |--P. maxima Blest 1979 NS00
    |    |    |--P. pallida Blest 1979 PVD10
    |    |    |--P. perplexa Blest 1979 PVD10
    |    |    `--P. prominula Blest 1979 NS00
    |    `--Metamynoglenes Blest 1979 PVD10
    |         |--*M. incurvata Blest 1979 PVD10
    |         |--M. absurda Blest & Vink 2002 PVD10
    |         |--M. attenuata Blest 1979 PVD10
    |         |--M. flagellata Blest 1979 PVD10
    |         |--M. gracilis Blest 1979 PVD10
    |         |--M. helicoides PVD10
    |         |--M. magna Blest 1979 PVD10
    |         `--M. ngongotaha Blest & Vink 2002 PVD10
    |--Linyphiinae W76
    |    |--Mioxena C90
    |    |--Tennesseellum formicum (Emerton 1882) WCH11
    |    |--Gongylidium rufipes (Linnaeus 1758) K01, BBM02
    |    |--Linyphia W76
    |    |--Laetesia PVD10
    |    |--Microlinyphia Gerhardt 1928 W76
    |    |    `--M. pusilla (Sundevall 1830) MKD01 [=Linyphia pusilla W76]
    |    |--Ostearius Hull 1911 PVD10
    |    |    `--O. melanopygius (Pickard-Cambridge 1879) W76 (see below for synonymy)
    |    |--Scironis Bishop & Crosby 1938 C90
    |    |    |--S. sima C90
    |    |    `--S. tarsalis C90
    |    |--Ringina F64
    |    |    |--R. antarctica (Hickman 1939) [=Drapetisca antarctica] F64
    |    |    `--R. crozetensis Tambs-Lyche 1954 F64
    |    |--Dismodicus K01
    |    |    |--D. bifrons (Blackwall 1841) MKD01
    |    |    `--D. elevatus K01
    |    |--Novalaetesia Millidge 1988 PVD10
    |    |    |--*N. anceps Millidge 1988 PVD10
    |    |    `--N. atra Blest & Vink 2003 PVD10
    |    |--Dunedinia Millidge 1988 PVD10
    |    |    |--*D. denticulata Millidge 1988 PVD19
    |    |    |--D. decolor Millidge 1988 PVD10
    |    |    `--D. pullata Millidge 1988 PVD10
    |    |--Laperousea de Dalmas 1917 PVD10
    |    |    |--*L. blattifera (Urquhart 1887) PVD10 [=Linyphia blattifer NS00]
    |    |    |--L. cupidinea (Simon 1908) W76
    |    |    `--L. quindecimpunctata (Urquhardt 1893) W76
    |    |--Stemonyphantes [Stemonyphantinae] C90
    |    |    |--S. conspersus (Koch 1879) MKD01
    |    |    |--S. lineatus (Linnaeus 1758) K02
    |    |    `--S. sibiricus (Grube 1861) MKD01
    |    |--Drapetisca Menge 1866 PVD10
    |    |    |--D. alteranda Chamberlin 1909 DH14
    |    |    |--D. australis Forster 1955 PVD10
    |    |    |--D. bicruris (Tu & Li 2006) DH14
    |    |    `--D. socialis (Sundevall 1833) DH14
    |    |--Caviphantes Oi 1960 [incl. Lessertiella Dumintrescu & Miller 1962] C90
    |    |    |--C. dobrogicus C90
    |    |    |--C. pseudosaxetorum C90
    |    |    |--C. samensis C90
    |    |    `--C. saxetorum C90
    |    |--Diploplecta Millidge 1988 PVD10
    |    |    |--*D. communis Millidge 1988 PVD10
    |    |    |--D. adjacens Millidge 1988 PVD10
    |    |    |--D. duplex Millidge 1988 PVD10
    |    |    |--D. nuda Millidge 1988 PVD10
    |    |    |--D. opaca Millidge 1988 PVD10
    |    |    |--D. proxima Millidge 1988 PVD10
    |    |    `--D. simplex Millidge 1988 PVD10
    |    |--Maorineta Millidge 1988 PVD10
    |    |    |--*M. tibialis Millidge 1988 PVD10
    |    |    |--M. acerba Millidge 1988 PVD10
    |    |    |--M. gentilis Millidge 1988 PVD10
    |    |    |--M. minor Millidge 1988 PVD10
    |    |    |--M. mollis Millidge 1988 PVD10
    |    |    |--M. sulcata Millidge 1988 PVD10
    |    |    `--M. tumida Millidge 1988 PVD10
    |    `--Porrhomma Simon 1884 H39
    |         |--P. antarctica Hickman 1939 H39
    |         |--P. egeria Simon 1884 RL11
    |         |--P. fuegianum H39
    |         |--P. longjiangense Zhu & Wang 1983 MKD01
    |         |--P. microphthalmum (Cambridge 1871) BBM02
    |         |--P. microps (Roewer 1931) RL11
    |         |--P. myops Simon 1884 RL11
    |         |--P. pygmaeum (Blackwall 1834) RL11
    |         `--P. rakanum Yaginuma & Saito 1981 MKD01
    `--Erigoninae [Erigonidae] W76
         |--Eperigone tridentata M83
         |--Diplocephalus MKD01
         |--Erigone MKD01
         |--Tunagyna debilis JVHN90
         |--Walckenaeria D90
         |--Delorrhipis fronticornis Simon 1884 MKD01
         |--Sciastes Bishop & Crosby 1938 W76
         |--Aphileta Hull 1920 W76
         |--Ceratinops D90
         |--Eridantes D90
         |--Minyriolus pusillus (Wider 1834) D90, MKD01
         |--Ceratinopsis interventa Chamberlin 1948 D90, MKD01
         |--Islandiana D90
         |--Halorates [incl. Catabrithorax] D90
         |--Spirembolus D90
         |--Sisicottus D90
         |--Diplocentria [incl. Scotoussa] D90
         |--Scylaceus D90
         |--Eogonatium P92
         |--Microctenonyx Dahl 1886 PVD10
         |    `--M. subitaneus (Pickard-Cambridge 1875) PVD10
         |--Araeoncus Simon 1884 PVD10
         |    `--*A. humilis (Blackwall 1841) PVD10
         |--Holminaria Eskov 1991 MKD01
         |    `--H. sibirica Eskov 1991 [incl. Birgerius triangulus Tao, Li & Zhu 1995] MKD01
         |--Uusitaloia Marusik, Koponen & Danilov 2001 MKD01
         |    `--*U. transbaicalica Marusik, Koponen & Danilov 2001 MKD01
         |--Lessertia Smith 1908 PVD10
         |    `--*L. dentichelis (Simon 1884) PVD10
         |--Mermessus Pickard-Cambridge 1899 PVD10
         |    |--M. denticulatus (Banks 1898) PO08
         |    `--M. fradeorum (Berland 1932) PVD10
         |--Maso JVHN90
         |    |--M. gallicus Simon 1894 BBM02
         |    `--M. sundevalli (Westring 1851) MKD01
         |--Tapinocyba D90
         |    |--T. insecta (Koch 1869) BBM02
         |    `--T. pallens RKD02
         |--Scotinotylus D90
         |    |--‘Scironis’ autor C90
         |    `--S. protervus (Koch 1879) MKD01
         |--Collinsia D90
         |    |--C. inerrans (Cambridge 1855) BBM02
         |    `--C. submissa (Koch 1879) MKD01
         |--Micryphantes Koch 1833 P92, PL73 [Micryphantidae]
         |    |--*M. ovatus Koch 1833 [=Meioneta ovata] PL73
         |    `--M. rurestris PL73
         |--Grammonota D90
         |    |--G. angusta JDW90
         |    |--G. inornata Emerton 1882 WCH11
         |    `--G. pictilis JDW90
         |--Hypselistes JVHN90
         |    |--H. florens JVHN90
         |    |--H. jacksoni (P.-Cambridge 1902) MKD01
         |    `--H. semiflavus (Koch 1879) MKD01
         |--Pocadicnemis C90
         |    |--P. americana Millidge 1976 PS08
         |    |--P. juncea Locket & Millidge 1953 PG08
         |    `--P. pumila (Blackwall 1841) MKD01
         |--Savignia MKD01
         |    |--S. borea MKD01
         |    |--S. eskovi Marusik, Koponen & Danilov 2001 MKD01
         |    |--S. frontata (Blackwall 1833) BH02
         |    `--S. nenilini Marusik 1988 MKD01
         |--Ceraticelus D90
         |    |--C. atriceps JDW90
         |    |--C. emertoni (Pickard-Cambridge 1874) MF11
         |    |--C. fissiceps JVHN90
         |    `--C. orientalis Eskov 1987 MKD01
         |--Oedothorax Bertkau 1883 TS06
         |    |--O. agrestis (Blackwall 1853) MKD01
         |    |--O. apicatus (Blackwall 1850) K08
         |    |--O. fuscus (Blackwall 1834) PG08 [=Gongylidium fuscus V09]
         |    |--O. gibbosus (Blackwall 1841) [incl. O. tuberosus] J02
         |    `--O. retusus (Westring 1851) MF11
         `--Ceratinella D90
              |--C. brevipes (Westring 1851) MKD01
              |--C. brevis (Wider 1834) BBM02
              |--C. rotunda K01
              |--C. scabrosa (P.-Cambridge 1863) BBM02
              `--C. sydneyensis Wunderlich 1976 W76

Linyphiidae incertae sedis:
  Pityohyphantes MKD01
    |--P. costatus (Hentz 1850) WA02
    |--P. subarticus Chamberlin & Ivie 1943 PS08
    `--P. phrygianus (Koch 1836) MKD01
  Drepanotylus uncatus (Cambridge 1873) K02
  Tallusia experta (Cambridge 1871) K02
  Bolyphantes MF11
    |--B. alticeps (Sundevall 1833) MF11
    |--B. index BV00
    `--B. luteolus (Blackwall 1833) K02
  Minicia marginella (Wider 1834) K02
  Taranucnus setosus (O.P.-Cambridge 1863) K02
  Hilaira MKD01
    |--H. gibbosa Tanasevitch 1982 MKD01
    |--H. jamalensis Eskov 1981 MKD01
    `--H. nubigena Hull 1911 K02
  Dicymbium nigrum (Blackwall 1834) V09, BBM02
  Entelecara MKD01
    |--E. acuminata V09
    |--E. congenera (P.-Cambridge 1879) MKD01
    |--E. erythropus (Westring 1851) BH02
    `--E. sombra (Chamberlin & Ivie 1947) MKD01
  Gonatium rubens (Blackwall 1833) MKD01 [=Neriene rubens V09]
  Cnephalocotes obscurus (Blackwall 1834) MKD01
  Monocephalus fuscipes (Blackwall 1836) BH02
  Erigonella hiemalis (Blackwall 1841) BH02
  Panamomops sulcifrons (Wider 1834) BH02
  Syedra gracilis (Menge 1869) BH02
  Diplostyla concolor (Wider 1834) MKD01, MF11
  Floronia bucculenta (Clerck 1757) BBM02
  Parapelecopsis nemoralis (Cambridge 1884) BBM02
  Styloctetor MKD01
    |--S. lehtineni Marusik & Tanasevitch 1999 MKD01
    `--S. romanus (Cambridge 1872) BBM02
  Tapinopa longidens (Wider 1834)BBM02
  Saaristoa abnormis (Blackwall 1841) V09, BBM02
  Gnathonarium MKD01
    |--G. dentatum (Wider 1834) MKD01 [=Gongylidium dentatum V09]
    `--G. suppositum (Kulczyński 1885) MKD01
  Tiso vagans (Blackwall 1834) V09, BBM02
  Troxochrus V09
    |--T. nasutus K08
    `--T. scabriculus (Westring 1851) J02 (see below for synonymy)
  Metopobactrus prominulus (Cambridge 1872) BBM02
  Kaestneria MKD01
    |--K. dorsalis (Wider 1834) MKD01 [=Bathyphantes dorsalis V09]
    `--K. pullata (P.-Cambridge 1863) MKD01
  Prinerigone vagans (Audouin 1826) MF11
  Thypochrestus digitatus (Cambridge 1872) BBM02
  Trichopterna cito (Cambridge 1872) BBM02
  Lepthothrix hardyi (Blackwall 1850) BBM02
  Trichoncoides piscator (Simon 1884) PO08
  Trichoncus MKD01
    |--T. hackmani SS02
    |--T. hyperboreus Eskov 1992 MKD01
    `--T. vasconicus Denis 1944 MKD01
  ‘Theridium’ pumilio Urquhart 1886 NS00 [=Theridion pumilio M83]
  Neriene MKD01
    |--N. clathrata (Sundevall 1830) BBM02 [=Linyphia clathrata V09]
    |--N. hammeni (van Helsdingen 1963) MKD01
    |--N. japonica (Oi 1960) [=Neolinyphia japonica] MKD01
    `--N. radiata (Walckenaer 1841) MKD01
  Macrargus MKD01
    |--M. carpenteri (Cambridge 1894) K02
    |--M. multesimus (P.-Cambridge 1875) MKD01
    `--M. rufus (Wider 1830) SL14 [=Tmeticus rufus V09]
  Centromerita V09
    |--C. bicolor (Blackwall 1833) BBM02 [=Tmeticus bicolor V09]
    `--C. concinna (Thorell 1875) K02
  Bathyphantes MKD01
    |--B. eumenis (Koch 1879) MKD01
    |    |--B. e. eumenis RL11
    |    `--B. e. buchari (Koch 1879) RL11
    |--B. extricates (Pickard-Cambridge 1876) PO08
    |--B. gracilis (Blackwall 1841) PG08
    |--B. parvulus (Westring 1851) MKD01
    `--B. setiger P.-Cambridge 1894 MKD01
  Gongylidiellum PO08
    |--G. murcidum RKD02
    `--G. vivum (Cambridge 1875) BBM02
  Poeciloneta MKD01
    |--P. globosa (Wider 1841) BH02
    `--P. variegata (Blackwall 1841) MKD02
  Sintula PO08
    |--S. corniger RKD02
    `--S. spiniger SS02
  Maro MKD01
    |--M. flavescens (P.-Cambridge 1873) MKD01
    |--M. lepidus Casemir 1961 K02
    |--M. minutus Cambridge 1906 K02
    `--M. sublestus Falconer 1915 K02
  Pelecopsis MKD01
    |--P. baicalensis Marusik, Koponen & Danilov 2001 MKD01
    |--P. inedita (Pickard-Cambridge 1875) PO08
    |--P. janus Jocqué 1984 J02
    |--P. mengei (Simon 1884) K02
    `--P. parallella (Wider 1834) MKD01 [=Lophocarenum parallelum V09]
  Micrargus SL14
    |--M. apertus RKD02
    |--M. herbigradus (Blackwall 1854) SL14
    `--M. subaequalis (Westring 1851) BH02
  Centromerus MKD01
    |--C. aequalis SS02
    |--C. arcanus (Cambridge 1873) K02
    |--C. clarus (Koch 1879) MKD01
    |--C. levitarsis (Simon 1884) K02
    |--C. prudens (Cambridge 1873) BBM02
    |--C. satyrus D93
    |    |--C. s. satyrus D93
    |    `--C. s. paradoxus D93
    `--C. sylvaticus (Blackwall 1841) BBM02
  Troglohyphantes strandi PN02
  Prolinyphia marginata JVHN90
  Aulacocyba subitanea F75
  Sphecozone longipes (Strand 1908) [=Hypselistoides longipes] J98
  Bathylinyphia Eskov 1992 MKD01
    `--*B. maior (Kulczyński 1885) (see below for synonymy) MKD01
  Lasiargus MKD01
    |--L. hirsutus (Menge 1869) MKD01
    |--L. pilipes (Kulczyński 1908) [=Metopobactrus pilipes; incl. L. laricetorum Eskov 1989] MKD01
    `--L. zhui Eskov & Marusik 1994 MKD01
  Silometopoides Eskov 1990 MKD01
    |--*S. pampia (Chamberlin 1943) [=Minyriolus pampia] MKD01
    |--S. asiaticus (Eskov in Eskov & Marusik 1995) [=Silometopus asiaticus] MKD01
    |--S. koponeni (Eskov & Marusik 1994) [=Silometopus koponeni] MKD01
    |--S. mongolensis Eskov & Marusik 1992 MKD01
    |--S. sibiricus (Eskov 1989) [=Silometopus sibiricus] MKD01
    |--S. sphagnicola Eskov & Marusik 1992 MKD01
    |--S. tibialis (Heimer 1987) [=Metopobactrus tibialis] MKD01
    `--S. yodoensis (Oi 1960) [=Lophomma yodoensis] MKD01
  Silometopus MKD01
    |--S. ambiguus (Pickard-Cambridge 1905) PG08
    `--S. sachalinensis (Eskov & Marusik 1994) [=Silometopoides sachalinensis] MKD01
  Sauron Eskov in Eskov & Marusik 1995 [incl. Bothriopterna Miller & Zitnanska 1976 (n. n.)] MKD01
    |--*S. fissocornis Eskov 1995 MKD01
    `--S. rayi (Simon 1881) [=Metopobactrus rayi; incl. Trichopterna fatrensis Miller 1966] MKD01
  Asiceratinops kolymensis Eskov 1992 MKD01
  Asthenargoides kurenstchikovi Eskov 1993 MKD01
  Pacifiphantes MKD01
  Perlongipalpus pinipumilis Eskov & Marusik 1991 MKD01
  Procerocymbium sibiricum Eskov 1989 [=P. sibiricus] MKD01
  Abacoproeces saltuum (Koch 1872) MKD01
  Abiskoa abiskoensis (Holm 1945) MKD01
  Acartauchenius scurrilis (P.-Cambridge 1872) MKD01
  Concavocephalus rubens Eskov 1989 MKD01
  Dactylopisthes video (Chamberlin & Ivie 1947) MKD01
  Decipiphantes decipiens (Koch 1879) MKD01
  Epibellowia pacifica (Eskov & Marusik 1992) MKD01
  Erigonidium graminicola (Sundevall 1830) MKD01
  Erigonoplus sibiricus Eskov & Marusik 1997 MKD01
  Estrandia grandaeva (Keyserling 1886) MKD01
  Hylyphantes nigritus (Simon 1881) MKD01
  Hypomma bituberculatum (Wider 1834) MKD01 [=Dicyphus bituberculatus V09]
  Incestophantes MKD01
    |--I. calcaratus (Emerton 1909) PS08
    |--I. incestus (Koch 1879) MKD01
    `--I. kochiellus (Strand 1900) MKD01
  Ivielum sibiricum Eskov 1988 MKD01
  Nenilinium asiaticum Eskov 1988 MKD01
  Obscuriphantes pseudoobscurus (Marusik, Hippa & Koponen 1996) MKD01
  Oreonetides MKD01
    |--O. abnormis [=Tmeticus abnormis] V09
    `--O. helsdingeni Eskov 1984 MKD01
  Parawubanoides MKD01
    |--P. marusiki (Tanasevitch 1987 MKD01
    `--P. unicornis (P.-Cambridge 1873) MKD01
  Praestigia MKD01
  Thaleria sukatchevae Eskov & Marusik 1992 MKD01
  Tibioplus diversus (Koch 1897) MKD01
  Typhochrestoides baikalensis Eskov 1990 MKD01
  Ummeliata sibirica (Eskov 1980) MKD01
  Walckenaerianus aimakensis Wunderlich 1995 MKD01
  Wubanoides uralensis (Pakhorukov 1981) MKD01
    |--W. u. uralensis RL11
    `--W. u. lithodytes Schikora 2004 RL11
  Yakutopus xerophilus Eskov 1990 MKD01
  Zornella cultrigera (Koch 1879) MKD01
  Dubiaranea P-GGJ08
  Thaumatancus PO08
  Alioranus pastoralis (Pickard-Cambridge 1872) PO08
  Mecopisthes monticola Bosmans 1993 PO08
  Australolinyphia Wunderlich 1976 W76
    `--*A. remota Wunderlich 1976 W76
  Knischatiria Wunderlich 1976 W76
    `--*K. abnormis Wunderlich 1976 W76
  Anacotyle setoensis Oi 1960 W76
  Helophora insignis (Blackwall 1841) MF11
  Improphantes improbulus (Simon 1929) RL11
  Labulla machadoi Hormiga & Schaff 2005 F05
  Allomengea D90
  Wubana D90
  Hylophantes graminicola [=Gongylidium graminicolum] V09
  Leptoroptrum robustum V09
  Baryphyma duffeyi JD-S07
  Proelauna humicola JD-S07
  Callitrichia JD-S07
  Frontinellina JD-S07
  Frontinella E95
    |--F. communis SG14
    `--F. pyramitela E95
  Impulsor S93
    |--I. mutilus Petrunkevitch 1958 S93
    `--I. neglectus Petrunkevitch 1942 S93
  Liticen setosus Petrunkevitch 1942 S93
  Mystagogus S93
    |--M. dubius Petrunkevitch 1942 S93
    `--M. glaber Petrunkevitch 1942 S93
  Custodela cheiracantha (Koch & Berendt 1854) S93
  Malleator niger Petrunkevitch 1942 S93
  Meditrina circumvallata Petrunkevitch 1942 S93
  Obnisus tenuipes Petrunkevitch 1942 S93
  Antopia P92
  Viocurus P92
  Malepellis extincta Petrunkevitch 1971 P92
  Palaeolinyphia P92

*Bathylinyphia maior (Kulczyński 1885) [=Bathyphantes maior, Bathylinyphia major (l. c.), Neriene major (l. c.); incl. Bathyphantes japonicus Oi 1980] MKD01

Ostearius melanopygius (Pickard-Cambridge 1879) W76 [incl. Erigone atriventer Urquhart 1887 NS00, Neriene analis Simon 1894 W76]

Troxochrus scabriculus (Westring 1851) J02 [=T. scabrosa (l. c.) BBM02; incl. T. cirrifrons (Cambridge 1871) BBM02, T. scabriculus cirrifrons J02]

*Type species of generic name indicated

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Last updated: 14 January 2022.

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