Reed sac spider Clubiona phragmites, from Pavouci.

Belongs within: Entelegynae.

Clubiona is a large, cosmopolitan genus of sac spiders in which the fourth pair of legs is longer than the remainder.

Characters (from Chen & Huang 2012): Yellow or brownish; carapace concave and gradually narrowing towards cephalic region; thoracic groove distinct; abdomen cylindrical. Eyes homogeneous, arranged in two parallel transverse rows occupying most of head width; posterior eye row straight or slightly procurved, wider than anterior row. Male palp with retrolateral tibial apophysis; retrobasal cymbial spur absent. Legs prograde; tarsi with two claws; leg I as long as or shorter than leg II, leg IV longest.

<==Clubiona Latreille 1804 CH12 [Clubioninae]
    |--*C. pallidula (Clerck 1757) CH12
    |--C. asrevida Ono 1992 CH12
    |--C. bandoi Hayashi 1995 CH12
    |--C. barbara Lucas 1846 E12
    |--C. brevipes V09
    |--C. brevispina Huang & Chen in Chen & Huang 2012 CH12
    |--C. cambridgei H09
    |--C. canadensis Emerton 1890 PS08
    |--C. chathamensis H09
    |--C. clima M83
    |--C. compta Koch 1839 K55
    |--C. corrugata Bösenberg & Strand 1906 [incl. C. coerulescens sinensis Hu 1979] CH12
    |--C. corticalis S95
    |--C. cycladata E07
    |--C. damirkovaci Deeleman-Reinhold 2001 CH12
    |--C. deletrix P.-Cambridge 1885 [incl. C. maculata Song & Chen 1979, C. reichlini Schenkel 1944] CH12
    |--C. diversa Cambridge 1862 BBM02
    |--C. elaphines B96
    |--C. exilipes Lucas 1846 E12
    |--C. expansa Huang & Chen in Chen & Huang 2012 CH12
    |--C. femorocalcarata Huang & Chen in Chen & Huang 2012 CH12
    |--C. frisia Wunderlich & Schuett 1995 BBM02
    |--C. genevensis Koch 1866 PO08
    |--C. hilaris Simon 1878 K55
    |--C. hystrix CH12
    |--C. insulana Ono 1989 CH12
    |--C. japonica Koch 1878 CH12
    |--C. japonicola Bösenberg & Strand 1906 [incl. C. parajaponicola Schenkel 1963] CH12
    |--C. jucunda (Karsch 1879) (see below for synonymy) CH12
    |--C. kayashimai Ono 1994 CH12
    |--C. kigabensis Strand 1915 J98
    |--C. kuanshanensis Ono 1994 CH12
    |--C. kurilensis Bösenberg & Strand 1906 CH12
    |--C. kurosawai Ono 1986 CH12
    |--C. lutescens Westring 1851 BBM02
    |--C. mandibularis Lucas 1846 E12
    |--C. moesta Banks 1896 PO08
    |--C. moralis Song & Zhu in Song et al. 1991 CH12
    |--C. neglecta V09
    |--C. oblonga Lucas 1846 E12
    |--C. ornata Lucas 1846 E12
    |--C. pallipes Lucas 1846 E12
    |--C. parvula Lucas 1846 E12
    |--C. peculiaris H09
    |--C. phragmites Koch 1843 BBM02
    |--C. pteronetoides CH12
    |--C. reclusa Cambridge 1863 BBM02
    |--C. rufipes Lucas 1846 E12
    |--C. sapporensis Hayashi 1986 CH12
    |--C. subnotabilis Strand 1907 J98
    |--C. subrostrata Zhang & Hu 1991 CH12
    |--C. subtilis Koch 1867 BBM02
    |--C. taiwanica Ono 1994 [incl. C. bonicula Ono 1994] CH12
    |--C. tanikawai Ono 1989 CH12
    |--C. terrestris Westring 1851 SL14
    |--C. trivialis Koch 1843 BBM02
    |--C. viridula Ono 1989 CH12
    |--C. wolongica Zhu & An 1999 CH12
    |--C. yaginumai Hayashi 1989 CH12
    |--C. yangmingensis Hayashi & Yoshida 1993 CH12
    |--C. yoshidai Hayashi 1989 CH12
    `--C. zilla CH12

Clubiona jucunda (Karsch 1879) [=Liocranum jucundum; incl. C. mantis Bösenberg & Strand 1906, C. sakatensis Saito 1939] CH12

*Type species of generic name indicated


[BBM02] Bonte, D., L. Baert & J.-P. Maelfait. 2002. Spider assemblage structure and stability in a heterogeneous coastal dune system (Belgium). Journal of Arachnology 30 (2): 331–343.

[B96] Brunet, B. 1996. Spiderwatch: A Guide to Australian Spiders. Reed New Holland: Sydney.

[CH12] Chen, S.-H., & W.-J. Huang. 2012. The Spider Fauna of Taiwan—Araneae: Miturgidae, Anyphaenidae, Clubionidae. National Taiwan Normal University.

[E07] Elliott, M. G. 2007. Annotated catalogue of the Pompilidae (Hymenoptera) of Australia. Zootaxa 1428: 1–83.

[E12] Evenhuis, N. L. 2012. Publication and dating of the Exploration Scientifique de l'Algérie: Histoire Naturelle des Animaux Articulés (1846–1849) by Pierre Hippolyte Lucas. Zootaxa 3448: 1–61.

[H09] Hogg, H. R. 1909. Some New Zealand and Tasmanian Arachnidae. Transactions and Proceedings of the New Zealand Institute 42: 273–283.

[J98] Jäger, P. 1998. Das Typenmaterial der Spinnentiere (Arachnida: Acari, Amblypygi, Araneae, Opiliones, Pseudoscorpiones, Scorpiones, Uropygi) aus dem Museum Wiesbaden. Jahrbuecher des Nassauischen Vereins fuer Naturkunde 119: 81–91.

[K55] Kraus, O. 1955. Spinnen von Korsika, Sardinien und Elba (Arach., Araneae). Senckenbergiana Biologica 36: 371–394.

[M83] Martin, N. A. 1983. Miscellaneous observations on a pasture fauna: An annotated species list. DSIR Entomology Division Report 3: 1–98.

[PS08] Pinzón, J., & J. Spence. 2008. Performance of two arboreal pitfall trap designs in sampling cursorial spiders from tree trunks. Journal of Arachnology 36 (2): 280–286.

[PO08] Pluess, T., I. Opatovsky, E. Gavish-Regev, Y. Lubin & M. H. Schmidt. 2008. Spiders in wheat fields and semi-desert in the Negev (Israel). Journal of Arachnology 36 (2): 368–373.

[SL14] Samu, F., G. Lengyel, É. Szita, A. Bidló & P. Ódor. 2014. The effect of forest stand characteristics on spider diversity and species composition in deciduous-coniferous mixed forests. Journal of Arachnology 42 (2): 135–141.

[S95] Simon, E. 1895. Arachnides recueillis par Mr G. Potanine en Chine et en Mongolie (1876-1879). Izviestĭia Imperatorskoi Akademĭi Nauk” (Bulletin de l’Academie Imperiale des Sciences de St. Pétersbourg), Ser. 5, 4: 331–345.

[V09] Verdcourt, B. (ed.) 2009. Additions to the Wild Fauna and Flora of the Royal Botanic Gardens, Kew. XXVI. Miscellaneous records. Kew Bulletin 64 (1): 183–194.

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