Female orchard spider Leucauge venusta, copyright Kaldari.

Belongs within: Araneoidea.
Contains: Meta, Tetragnatha.

The Tetragnathidae are a cosmopolitan family of long-legged orb-weaving spiders. Many, but not all, species have characteristically long, well-developed chelicerae. Male papi are relatively simple and lack a median apophysis (Paquin et al. 2010). The femora are often provided with dorsal trichobothria though these are absent in the genus Meta (Scharff & Coddington 1997).

Characters (from Jocqué & Dippenaar-Schoeman 2007): Small to very large; eight eyes present; legs with three tarsal claws; ecribellate; entelegyne or secondarily haplogyne; male genitalia comparatively simple, with large paracymbium and often several cymbial processes, conductor and embolus coiled apically; median apophysis absent.

<==Tetragnathidae [Meteae, Pachygnathidae]
    |--+--Meta BS09
    |  `--Metellina BS09
    |       |--M. mengei (Blackwall 1870) BBM02
    |       |--M. merianae (Scopoli 1763) GA11
    |       `--M. segmentata (Clerck 1757) BS09, BH02 [=Meta segmentata V09]
    `--+--Leucauge Darwin in White 1841 BS09, PVD10 [Leucauginae]
       |    |--L. argyra Walckenaer 1842 WA02
       |    |--L. celebesiana (Walck. 1837) (see below for synonymy) R13
       |    |--L. decorate (Blackwell 1864) HU08
       |    |--L. dromedaria (Thorell 1881) PVD10 (see below for synonymy)
       |    |--L. festiva JD-S07
       |    |--L. grata R13 (see below for synonymy)
       |    |--L. magnifica S94
       |    |--L. mariana Taczanowski 1881 WA02
       |    `--L. venusta Walckenaer 1842 WA02
       `--+--Tetragnatheae SC97
          |    |--Tetragnatha BS09
          |    `--Cyrtognatha SC97
          `--Pachygnatheae SC97
               |--Pachygnatha SC97
               |    |--*P. clercki Sundevall 1823 SC97
               |    |--P. autumnalis Keyserling 1884 SC97
               |    |--P. degeeri Sundevall 1829 VPB11
               |    |--P. furcillata SC97
               |    |--P. listeri V09
               |    `--P. superba Rainbow 1897 R97
               `--Glenognatha Simon 1887 SC97, HD90
                    |--G. centralis Chamberlin 1925 HD90
                    |--G. emertoni HD90
                    |--G. foxi (McCook 1894) WCH11
                    |--G. heleios Hormiga & Döbel 1990 HD90
                    |--G. iviei Levi 1980 HD90
                    |--G. minuta Banks 1898 HD90
                    `--G. mira Bryant 1945 HD90

Tetragnathidae incertae sedis:
  Eugnatha KK90
    |--E. cylindracea Koch & Keyserling 1887 KK90
    |--E. cylindrica KK90
    `--E. rubriventris (Doleschall 1857) [=Tetragnatha rubriventris; incl. T. lupata Koch 1872] KK90
  Eucta Sim. 1881 KK90
    |--E. anguilla (Thorell 1877) [=Tetragnatha anguilla] KK90
    `--E. caudifera Koch & Keyserling 1887 KK90
  Tylorida ventralis FF99
  Plesiometa argyra GE05
  Metleucauge B11
    |--M. kompirensis (Bösenberg & Strand 1906) B11
    `--M. yunohamensis (Bösenberg & Strand 1906) B11
  Dolichognatha [Diphyeae, Dolichognathinae] JD-S07
  Dyschiriognatha JD-S07
  Doryonychus F95
  Nanometinae [Nanometidae] PVD10
    |--Orsiella lagenifera FF99
    |--Nanometa Simon 1908 PVD10
    |    `--*N. gentilis (Simon 1908) PVD10
    |--Eryciniolia Strand 1912 PVD10
    |    `--*E. purpurapunctata (Urquhart 1889) PVD10 (see below for synonymy)
    `--Orsinome Thorell 1890 PVD10
         |--*O. vethi [=Pachygnatha vethi] S99
         |--O. arborea (Urquhart 1890) [=Tetragnatha arborea] S99
         |--O. australis Simon 1899 S99
         `--O. lagenifera (Urquhart 1888) PVD10 [=Linyphia lagenifera NS00]
  Chrysometa lancetilla L05
  Argyroepeira Em. 1884 SC97, T94
    |--A. argyra C01
    |--A. culta K92
    |--A. nigrotrivittata (Doleschall 1859) [=Epeira nigrotrivittata] T94
    `--A. ventralis K92
  Azilieae SC97
  Memoratrix rydei Petrunkevitch 1942 S93
  Eometa S93
    |--E. aberrans Petrunkevitch 1958 S93
    |--E. longipes Petrunkevitch 1958 S93
    |--E. robusta Petrunkevitch 1958 S93
    `--E. samlandica Petrunkevitch 1958 S93
  Priscometa tenuipes Petrunkevitch 1958 S93
  Macryphantes cowdeni Selden 1990 S93
  Homalometa P92

*Eryciniolia purpurapunctata (Urquhart 1889) PVD10 [=Linyphia purpurapunctata NS00; incl. L. nitidulum Urquhart 1890 NS00, Erycina violacea Urquhart 1891 NS00]

Leucauge celebesiana (Walck. 1837) [=Tetragnatha celebesiana, Argyroepeira celebesiana; incl. Meta decorata Blackw. 1864] R13

Leucauge dromedaria (Thorell 1881) PVD10 [=Argyroepeira dromadaria S99; incl. Nephila argentatum Urquhart 1887 NS00]

Leucauge grata R13 [=Epeira grata R13, Argyroepeira grata R13; incl. E. coccinea Doleschall 1857 R13, Meta coccinea KK90]

*Type species of generic name indicated


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Last updated: 9 January 2022.

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