Neosuchia

Spectacled caiman Caiman crocodilus, copyright Berrucomons.


Belongs within: Metasuchia.
Contains: Thalattosuchia, Pholidosauridae, Crocodylidae.

The Neosuchia are a clade of crocodylomorphs that has been defined as including all taxa closer to Crocodylus niloticus than to Notosuchus terrestris. This clade includes all living crocodilians.

<==Neosuchia [Metamesosuchia] SF03
    |  i. s.: Susisuchus Salisbury, Frey et al. 2003 [Susisuchidae] SF03
    |           `--*S. anatoceps Salisbury, Frey et al. 2003 SF03
    |         Brillanceausuchus Michard et al. 1990 SF03, LB02
    |           `--B. babouriensis SF03
    |         Gilchristosuchus palatinus SF03
    |--Atoposauridae GPS06
    |    |--Theriosuchus pusillus GPS06
    |    |--Montsechosuchus depereti [=Alligatorium depereti] BC88
    |    `--Alligatorium [incl. Alligatorellus, Atoposaurus] BC88
    |         `--A. meyeri BC88
    `--+--+--Thalattosuchia GPS06
       |  `--Pholidosauridae GPS06
       `--+--Goniopholidae GPS06
          |    |--Sunosuchus SL01
          |    |--Eutretauranosuchus delfsi GPS06
          |    `--Goniopholis CRA04
          |         |--G. gracilidens [=Nannosuchus gracilidens] BC88
          |         `--G. paulistanus Roxo 1936 CRA04
          `--+--Shamosuchus Mook 1924 BC88, SF03 [incl. Paralligator BC88]
             |--Bernissartia Dollo 1885 GPS06, N85
             |    `--B. fagesii Dollo 1883 SF03
             `--Eusuchia GPS06
                  |--Borealosuchus formidabilis GPS06
                  |--Leidyosuchus BC88
                  |--Hylaeochampsa Owen 1874 GPS06, BC88
                  |    `--H. vectiana GPS06
                  |--Stomatosuchus Stromer 1925 [Stomatosuchidae] BC88
                  |    `--S. inermis SF03
                  `--Crocodylia BC88
                       |  i. s.: Diplocynodon RB05
                       |           |--D. darwini RRO00
                       |           |--D. hantoniensis RRO00
                       |           `--D. styriacus RB05
                       |         Pallimnarchus pollens F71
                       |         Rimasuchus Storrs 2003 LA04
                       |           `--*R. lloydi (Fourtau 1918) [=Crocodylus lloydi] LA04
                       |--+--Gavialidae AS09
                       |  |    |--Gavialis gangeticus GPS06
                       |  |    `--Tomistoma Müller 1846 AS09, LA04 [Tomistominae]
                       |  |         |--T. coppensi Pickford 1994 LA04
                       |  |         |--T. dowsoni Fourtau 1918 LA04
                       |  |         |--T. lusitanica Antunes 1967 LA04
                       |  |         `--T. schlegelii USDI77
                       |  `--Eosuchus lerichei GPS06
                       `--+--+--Crocodylidae GPS06
                          |  `--Osteolaemus tetraspis BC88, USDI77
                          |       |--O. t. tetraspis USDI77
                          |       `--O. t. osborni USDI77
                          `--Alligatoroidea D07
                               |  i. s.: Deinosuchus Colbert & Bird 1958 D07 [incl. Phobosuchus N85]
                               |           |--D. hatchery D07
                               |           |--D. riograndensis D07
                               |           `--D. rugosus D07
                               `--Alligatorinae ML14
                                    |--Navajosuchus mooki ML14
                                    `--+--+--Stangerochampsa mccabei AS09
                                       |  `--Alligator GPS06
                                       |       |--A. mississippiensis Daudin 1801 NS10
                                       |       `--A. sinensis SF03
                                       `--+--Paleosuchus BC88
                                          |    |--P. palpebrosus DS86
                                          |    `--P. trigonatus DS86
                                          `--Caiman BC88
                                               |  i. s.: C. sclerops GWN71
                                               |         C. yacare USDI77
                                               |--C. latirostris BC88
                                               `--+--C. crocodilus BC88
                                                  |    |--C. c. crocodilus B81
                                                  |    |--C. c. apaporiensis USDI77
                                                  |    `--C. c. fuscus B81
                                                  `--Melanosuchus niger BC88, DS86

Inorganic: Caiman crocodilus protominilorientalus Okamura 1987 O87

*Type species of generic name indicated

REFERENCES

[AS09] Alfaro, M. E., F. Santini, C. Brock, H. Alamillo, A. Dornburg, D. L. Rabosky, G. Carnevale & L. J. Harmon. 2009. Nine exceptional radiations plus high turnover explain species diversity in jawed vertebrates. Proceedings of the National Academy of Sciences of the USA 106 (32): 13410–13414.

[BC88] Benton, M. J., & J. M. Clark. 1988. Archosaur phylogeny and the relationships of the Crocodylia. In: Benton, M. J. (ed.) The Phylogeny and Classification of the Tetrapods vol. 1. Amphibians, Reptiles, Birds pp. 295–338. Clarendon Press: Oxford.

[B81] Brooks, D. R. 1981. Hennig's parasitological method: a proposed solution. Systematic Zoology 30 (3): 229–249.

[CRA04] Carvalho, I. de S., L. C. B. Ribeiro & L. dos S. Avilla. 2004. Uberabasuchus terrificus sp. nov., a new Crocodylomorpha from the Bauru Basin (Upper Cretaceous), Brazil. Gondwana Research 7 (4): 975–1002.

[D07] Dixon, D. 2007. The Complete Illustrated Encyclopedia of Dinosaurs & Prehistoric Creatures. Hermes House: London.

[DS86] Dixon, J. R., & P. Soini. 1986. The Reptiles of the Upper Amazon Basin, Iquitos Region, Peru. Milwaukee Public Museum: Milwaukee.

[F71] Fletcher, H. O. 1971. Catalogue of type specimens of fossils in the Australian Museum, Sydney. Australian Museum Memoir 13: 1–167.

[GPS06] Gasparini, Z., D. Pol & L. A. Spalletti. 2006. An unusual marine crocodyliform from the Jurassic–Cretaceous boundary of Patagonia. Science 311: 70–73.

[GWN71] Gorman, G. C., A. C. Wilson & M. Nakanishi. 1971. A biochemical approach towards the study of reptilian phylogeny: evolution of serum albumin and lactic dehydrogenase. Systematic Zoology 20 (2): 167–185.

[LB02] Lapparent de Broin, F. de. 2002. Elosuchus, a new genus of crocodile from the Lower Cretaceous of the north of Africa. Comptes Rendus Palevol 1: 275–285.

[LA04] Llinás Agrasar, E. 2004. Crocodile remains from the Burdigalian (lower Miocene) of Gebel Zelten (Libya). Geodiversitas 26 (2): 309–321.

[ML14] Mitchell, K. J., B. Llamas, J. Soubrier, N. J. Rawlence, T. H. Worthy, J. Wood, M. S. Y. Lee & A. Cooper. 2014. Ancient DNA reveals elephant birds and kiwi are sister taxa and clarifies ratite bird evolution. Science 344 (6186): 898–900.

[NS10] Nesbitt, S. J., C. A. Sidor, R. B. Irmis, K. D. Angielczyk, R. M. H. Smith & L. A. Tsuji. 2010. Ecologically distinct dinosaurian sister group shows early diversification of Ornithodira. Nature 464: 95–98.

[N85] Norman, D. 1985 (reprinted 2000). The Illustrated Encyclopedia of Dinosaurs. Salamander Books: London.

[O87] Okamura, C. 1987. New facts: Homo and all Vertebrata were born simultaneously in the former Paleozoic in Japan. Original Report of the Okamura Fossil Laboratory 15: 347–573.

[RB05] Rage, J.-C., & S. Bailon. 2005. Amphibians and squamate reptiles from the late early Miocene (MN 4) of Béon 1 (Montréal-du-Gers, southwestern France). Geodiversitas 27 (3): 413–441.

[RRO00] Rossmann, T., M. Rauhe & F. Ortega. 2000. Studies on Cenozoic crocodiles: 8. Bergisuchus dietrichbergi Kuhn (Sebecosuchia: Bergisuchidae n. fam.) from the Middle Eocene of Germany, some new systematic and biological conclusions. Paläontologische Zeitschrift 74: 379–392.

[SF03] Salisbury, S. W., E. Frey, D. M. Martill & M. C. Buchy. 2003. A new crocodilian from the Lower Cretaceous Crato Formatioin of north-eastern Brazil. Palaeontographica Abteilung A: Paläozoologie—Stratigraphie 270: 3-47.

[SL01] Sereno, P. C., H. C. E. Larsson, C. A. Sidor & B. Gado. 2001. The giant crocodyliform Sarcosuchus from the Cretaceous of Africa. Science 294: 1516–1519.

[USDI77] USDI (United States Department of the Interior). 1977. Endangered and threatened wildlife and plants—republication of list of species. Federal Register 42: 36420–36431.

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