Tephritoidea

Signal fly Rivellia sp., copyright Stephen Cresswell.


Belongs within: Schizophora.
Contains: Lonchaeidae, Richardiidae, Piophilidae, Ulidiidae, Tephritidae, Pyrgotidae.

The Tephritoidea are a group of acalyptrate flies united by the reduction or loss of sternite 6 in the male, the presence in females of anterior rod-like apodemes on sternites 4 to 7, and division of the female tergosternite 7 into two sections, with the anterior section forming a tubular oviscape and the posterior section comprising two pairs of longitudinal taeniae. Many tephritoids have strikingly patterned wings that are held outwards in display. Members include the Platystomatidae, signal flies, a varied group with often metallic-coloured bodies and strikingly patterned wings (Steyskal 1987).

<==Tephritoidea [Otitoidea] WT11
    |--+--Lonchaeidae WT11
    |  `--Richardiidae WT11
    `--+--Piophilidae WT11
       `--+--Ulidiidae WT11
          `--+--+--+--Tephritidae WT11
             |  |  `--+--Pyrgotidae WT11
             |  |     `--Tachiniscidae [Tachiniscinae] WT11
             |  |          |--Tachinisca cyaneiventris N10
             |  |          |--Neortalotrypeta bicolor N10
             |  |          |--Agnitrena igniceps N10
             |  |          |--Ischyropteron nigricaudatum N10
             |  |          `--Protortalotrypeta N10
             |  `--Ctenostylidae [Ctenostylinae, Lochmostyliinae] WT11
             |       |--Nepaliseta ashleyi K10, WT11
             |       |--Furciseta plaumanni K10
             |       |--*Ctenostylum rufum K10
             |       |--Sinolochmostylia K10
             |       |--Tauroscypson K10
             |       |    |--T. andina K10
             |       |    `--T. guiana K10
             |       |--Lochmostylia K10
             |       |    |--*L. borgmeieri K10
             |       |    `--L. lopesi K10
             |       `--Ramuliseta K10
             |            |--R. ashleyi K10
             |            |--R. lindneri O98
             |            `--R. palpifera O98
             `--Platystomatidae [Achiasidae] WT11
                  |  i. s.: Giraffomyia GE05
                  |         Lamprogaster C91
                  |           |--L. imperialis ZS10
                  |           |--L. semicyanea [=Dryomyza semicyanea] O98
                  |           `--L. violacea C91
                  |         Achias W10
                  |           |--A. australis ZS10
                  |           |--A. kimi O98
                  |           |--A. minax O98
                  |           `--A. rothschildi GE05
                  |         Plagiostenopterina W10
                  |         Euprosopia tenuicornis CM91
                  |         Elassogaster linearis CM91
                  |         Duomyia CM91
                  |         Lenophila dentipes CM74
                  |         Asyntona O98
                  |         Cleitamia astrolabei O98
                  |         Zygaenula O98
                  |         Pterogenia O98
                  |         Naupoda O98
                  |         Atopognathus complens O98
                  |         Trigonosoma decorum O98
                  |--Phasiamya O98 [Trapherinae W10]
                  |--Plastotephritinae W10
                  |--Scholastinae W10
                  |    |--Neohemigaster O98
                  |    |--Mesoctenia O98
                  |    `--Scholastes O98
                  |         |--S. foordi RJ93
                  |         `--S. lonchifer O98
                  `--Platystomatinae [Angitulinae] W10
                       |--Platystoma Meigen 1803 BR05
                       |--Himeroessa pretiosa W10, S87
                       |--Angitula nigra O98
                       |--Pogonortalis doclea O98, S87
                       |--Amphicnephes W10
                       |    |--A. pullus S87
                       |    `--A. stellatus W10
                       |--Senopterina W10 [=Stenopterina S87; incl. Bricinnia W10, Bricinniella W10]
                       |    |--S. brevipes W10
                       |    |--S. caerulescens W10
                       |    `--S. mexicana W10
                       `--Rivellia W10
                            |--R. basilaris BM76
                            |--R. bosci R13
                            |--R. flavimana W10
                            |--R. inaequata W10
                            |--R. occulta W10
                            |--R. syngenesiae WT11
                            `--R. trigona BM76

*Type species of generic name indicated

REFERENCES

[BM76] Bohart, R. M., & A. S. Menke. 1976. Sphecid Wasps of the World. University of California Press: Berkeley.

[BR05] Bouchet, P., & J.-P. Rocroi. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1–2): 1–397.

[CM74] Colless, D. H., & D. K. McAlpine. 1974. Diptera (flies). In: CSIRO. The Insects of Australia: A textbook for students and research workers. Supplement 1974 pp. 91–96. Melbourne University Press.

[CM91] Colless, D. H., & D. K. McAlpine. 1991. Diptera (flies). In: CSIRO. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 2 pp. 717–786. Melbourne University Press: Carlton (Victoria).

[C91] CSIRO. 1991. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 1. Melbourne University Press: Carlton (Victoria).

[GE05] Grimaldi, D., & M. S. Engel. 2005. Evolution of the Insects. Cambridge University Press: New York.

[K10] Korneyev, V. A. 2010. Ctenostylidae (ctenostylid flies). In: Brown, B. V., A. Borkent, J. M. Cumming, D. M. Wood, N. E. Woodley & M. A. Zumbado (eds) Manual of Central American Diptera vol. 2 pp. 963–969. NRC Research Press: Ottawa.

[N10] Norrbom, A. L. 2010. Tephritidae (fruit flies, moscas de frutas). In: Brown, B. V., A. Borkent, J. M. Cumming, D. M. Wood, N. E. Woodley & M. A. Zumbado (eds) Manual of Central American Diptera vol. 2 pp. 909–954. NRC Research Press: Ottawa.

[O98] Oosterbroek, P. 1998. The Families of Diptera of the Malay Archipelago. Brill: Leiden.

[R13] Reuter, O. M. 1913. Lebensgewohnheiten und Instinkte der Insekten bis zum Erwachen der sozialen Instinkte. R. Friedländer & Sohn: Berlin.

[RJ93] Ross, A. J., & E. A. Jarzembowski. 1993. Arthropoda (Hexapoda; Insecta). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 363–426. Chapman & Hall: London.

[S87] Steyskal, G. C. 1987. Platystomatidae. In: McAlpine, J. F. (ed.) Manual of Nearctic Diptera vol. 2 pp. 809–812. Research Branch, Agriculture Canada.

[W10] Whittington, A. E. 2010. Platystomatidae (signal flies). In: Brown, B. V., A. Borkent, J. M. Cumming, D. M. Wood, N. E. Woodley & M. A. Zumbado (eds) Manual of Central American Diptera vol. 2 pp. 903–907. NRC Research Press: Ottawa.

[WT11] Wiegmann, B. M., M. D. Trautwein, I. S. Winkler, N. B. Barr, J.-W. Kim, C. Lambkin, M. A. Bertone, B. K. Cassel, K. M. Bayless, A. M. Heimberg, B. M. Wheeler, K. J. Peterson, T. Pape, B. J. Sinclair, J. H. Skevington, V. Blagoderov, J. Caravas, S. N. Kutty, U. Schmidt-Ott, G. E. Kampmeier, F. C. Thompson, D. A. Grimaldi, A. T. Beckenbach, G. W. Courtney, M. Friedrich, R. Meier & D. K. Yeates. 2011. Episodic radiations in the fly tree of life. Proceedings of the National Academy of Sciences of the USA 108 (14): 5690–5695.

[ZS10] Zborowski, P., & R. Storey. 2010. A Field Guide to Insects in Australia 3rd ed. Reed New Holland: Sydney.

Last updated: 12 November 2019.

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