Sugar cane armyworm Leucania stenographa, copyright Simon Hinkley & Ken Walker.

Belongs within: Xyleninae.

The Hadeninae are a group of moths whose larvae often feed on grasses (Poaceae); the oblong, smooth eggs are often laid in clusters in a gel. Adults have hairy, unlashed eyes and unspined tibiae (Nielsen & Common 1991). Members include the Glottulini, a small group of colourful moths found in the Malesian region whose larvae feed on bulbs of Amaryllidaceae. Leucania is a genus of cryptically coloured moths with fore wings bearing striate brown shading on a tan background that rest by day among dry grasses.

Hadeninae M99
    `--Glottulini [Glottulidae, Glottulinae] ZK11
         |--Chasmina cygnus WM66
         `--Diaphone capillamentum ZK11

Hadeninae incertae sedis:
  Metopiora sanguinata (Lucas 1892) M99
  Dasygaster padockina NC91
  Tiracola plagiata NC91
  Hadena P01
    |--H. amanda Swinhoe 1901 S01
    |--H. ochroleuca F92
    |--H. lateritia P01
    |--H. tritici R13
    `--H. turbulenta S96
  Persectania NC91
    |--P. atristriga M83
    |--P. aversa M83
    |--P. ewingii NC91
    `--P. steropastis L27
  Pseudaletia C70
    |--P. adultera [=Cirphis adultera] S72
    |--P. convecta C70
    |--P. separata H74
    `--P. unipuncta LY07
  Leucania M99
    |--L. albicosta [=Cirphis albicosta] T69
    |--L. albipuncta F92
    |--L. alopa T27
    |--L. amblycasis T69
    |--L. atristriga T27
    |--L. comma F92
    |--L. commoides T69
    |--L. cruegeri MC13
    |--L. griseipennis T27
    |--L. insueta T69
    |--L. juncicola T69
    |--L. latiuscula T69
    |--L. linda T69
    |--L. linita T69
    |--L. littoralis F92
    |--L. loreyi F92
    |--L. lithargyrea F92
    |--L. miasticta T69
    |--L. mollis L27
    |--L. multilinea T69
    |--L. oaxacana T69
    |--L. obsoleta F92
    |--L. obtusa Guenée 1852 M99
    |--L. pallens F92
    |--L. phragmitidicola T69
    |--L. pseudargyria T69
    |--L. purdii T27
    |--L. scirpicola T69
    |--L. semivittata T27
    |--L. separata F92
    |--L. solita H01
    |--L. stenographa Lower 1900 M99
    |--L. unipunctata B88
    |--L. ursula T69
    `--L. vitellina T69

*Type species of generic name indicated


[B88] Bouček, Z. 1988. Australasian Chalcidoidea (Hymenoptera): A biosystematic revision of genera of fourteen families, with a reclassification of species. CAB International: Wallingford (UK).

[C70] Common, I. F. B. 1970. Lepidoptera (moths and butterflies). In: CSIRO. The Insects of Australia: A textbook for students and research workers pp. 765–866. Melbourne University Press.

[F92] Fan Z. 1992. Key to the Common Flies of China 2nd ed. Science Press: Beijing.

[H01] Hampson, G. F. 1901. The Lepidoptera-Phalaenae of the Bahamas. Annals and Magazine of Natural History, series 7, 7: 246–261.

[H74] Helson, G. A. H. 1974. Insect Pests: Identification, life history, and control of pests of farms, horticulture, gardens, and public health. A. R. Shearer, Government Printer: Wellington (New Zealand).

[L27] Lindsay, S. 1927. A list of the Lepidoptera of Deans Bush, Riccarton, Canterbury. Transactions and Proceedings of the New Zealand Institute 57: 693–696.

[LY07] Liu, J., X. Yang, Y. Zhang & Z. Liu. 2007. Effects of exogenous juvenile hormone III on the ixodid tick, Haemaphysalis longicornis Neumann (Acari: Ixodidae). In: Morales-Malacara, J. B., V. M. Behan-Pelletier, E. Ueckermann, T. M. Pérez, E. G. Estrada-Venegas & M. Badii (eds) Acarology XI: Proceedings of the International Congress pp. 209–213. Instituto de Biología and Faculdad de Ciencias, Universidad Nacional Autónoma de México, Sociedad Latinoamericana de Acarología: México.

[MC13] Majer, J. D., S. K. Callan, K. Edwards, N. R. Gunawardene & C. K. Taylor. 2013. Baseline survey of the terrestrial invertebrate fauna of Barrow Island. Records of the Western Australian Museum Supplement 83: 13–112.

[M83] Martin, N. A. 1983. Miscellaneous observations on a pasture fauna: an annotated species list. DSIR Entomology Division Report 3: 1–98.

[M99] Matthews, M. 1999. Heliothine Moths of Australia: A guide to bollworms and related noctuid groups. CSIRO Publishing.

[NC91] Nielsen, E. S., & I. F. B. Common. 1991. Lepidoptera (moths and butterflies). In: CSIRO. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 2 pp. 817–915. Melbourne University Press: Carlton (Victoria).

[P01] Pável, J. 1901. Lepkék [Lepidopteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 171–177. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[R13] Reuter, O. M. 1913. Lebensgewohnheiten und Instinkte der Insekten bis zum Erwachen der sozialen Instinkte. R. Friedländer & Sohn: Berlin.

[S96] Smith, D. R. 1996. Trigonalyidae (Hymenoptera) in the eastern United States: seasonal flight activity, distribution, hosts. Proceedings of the Entomological Society of Washington 98 (1): 109–118.

[S72] Southcott, R. V. 1972. Revision of the larvae of the tribe Callidosomatini (Acarina: Erythraeidae) with observations on post-larval instars. Australian Journal of Zoology, Supplementary Series 13: 1–84.

[S01] Swinhoe, C. 1901. New genera and species of eastern and Australian moths (continued). Annals and Magazine of Natural History, series 7, 7: 489–501.

[T27] Thomson, G. M. 1927. The pollination of New Zealand flowers by birds and insects. Transactions and Proceedings of the New Zealand Institute 57: 106–125.

[T69] Treat, A. E. 1969. The genus Dicrocheles: three new species and a reappraisal. Acarologia 11 (4): 630–656.

[WM66] Wallace, A. R., & F. Moore. 1866. List of lepidopterous insects collected at Takow, Formosa, by Mr. R. Swinhoe. Proceedings of the Zoological Society of London 1866: 355–365.

[ZK11] Zahiri, R., I. J. Kitching, J. D. Lafontaine, M. Mutanen, L. Kaila, J. D. Holloway & N. Wahlberg. 2011. A new molecular phylogeny offers hope for a stable family level classification of the Noctuoidea (Lepidoptera). Zoologica Scripta 40 (2): 158–173.

Last updated: 21 February 2021.

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