Pleurotomarioidea

Trochotoma gansuensis, from Tong & Erwin (2001).

Belongs within: Vetigastropoda.
Contains: Pleurotomaria, Perotrochus.

The Pleurotomarioidea, slit shells, are a group of marine gastropods represented in the modern fauna only by the family Pleurotomariidae. Members of this group have a shell with a labral slit, followed by a corresponding selenizone (Bouchet et al. 2005). The Pleurotomariidae have a trochiform shell and are restricted in the modern fauna to deeper waters where they feed primarily on sponges.

<==Pleurotomarioidea
    |  i. s.: Mellarium Waterhouse 1960 W63c
    |           `--M. mutchi W63a
    |--Catantostoma Sandberger 1842 [Catantostomatidae, Catantostomatinae] BR05
    |    `--*C. clathratum Sandberger 1842 KC60
    |--Rhaphischisma Knight 1936 BR05 [=Rotellina de Koninck 1881 non Agassiz 1846 KC60; Rhaphischismatidae BR05]
    |    `--*R. planorbiformis (de Koninck 1881) [=*Rotellina planorbiformis] BR17
    |--Lancedellia Bandel 1991 [Lancedelliidae] BR17
    |    `--*L. costata (Zardini 1978) [=Paleunema costatum] BR17
    |--Stuorella Kittl 1891 KC60 [Stuorellidae BR17]
    |    `--*S. subconcava (Münster 1841) [=Trochus subconcavus] KC60
    |--Polytremariidae [Polytremariinae] BR05
    |    |--Polytremaria d’Orbigny 1850 BR05
    |    |    `--*P. catenata (de Koninck 1843) [=Pleurotomaria catenata] BR17
    |    `--Plocostoma Gemmellaro 1889 KC60
    |         |--*P. neumayri Gemmellaro 1890 KC60, TTE93 [=Pleurotomaria (*Plocostoma) neumayri KC60]
    |         |--P. josephina Gemmellaro 1890 TTE93
    |         `--P. piazzi Gemmellaro 1890 TTE93
    |--Zygitidae BR05
    |    |--Eirlysia Batten 1956 TE01
    |    |    |--*E. exquisita Batten 1956 KC60
    |    |    `--E. reticulata Batten 1958 W63b
    |    `--Zygites Kittl 1891 BR05
    |         |--Z. delphinula (Laube 1868) TE01 (see below for synonymy)
    |         |--Z. elegans Yin & Yochelson 1983 TTE93
    |         |--Z. laevigatus Tong & Erwin 2001 TE01
    |         `--Z. marmorea (Koken 1897) TTE93
    |--Trochotomidae [Ditremariinae] BR05
    |    |--Valfinia Cox 1958 [=Didymodon Fischer 1885 nec Hedw. 1801 (ICBN) nec Blake 1863] KC60
    |    |    |--*V. quinquecincta (Zieten 1832) [=Trochus quinquecinctus, *Didymodon quinquecinctus] KC60
    |    |    `--V. thurmanni KC60
    |    `--Trochotoma Eudes-Deslongchamps 1842 BR05 (see below for synonymy)
    |         |  i. s.: T. orbita Pan 1982 TTE93
    |         |--T. (Trochotoma) KC60
    |         |    |--*T. (T.) conuloides Eudes-Deslongchamps 1842 BR17
    |         |    `--T. (T.) bicarinata (d’Orbigny 1843) KC60, BR17 [=Ditremaria bicarinata BR17]
    |         `--T. (Discotoma Haber 1934) TE01
    |              |--T. (*D.) amata (d’Orbigny 1850) [=Ditremaria amata] TE01
    |              |--T. (D.) gansuensis Tong & Erwin 2001 TE01
    |              `--‘Discotoma’ planoconvexa TE01
    |--Portlockiellidae BR05
    |    |--Agniesella Cossmann 1909 [=Pleuroderma Perner 1907 non Tschudi 1837] KC60
    |    |    `--*A. aratula (Perner 1907) [=Pleurotomaria (*Pleuroderma) aratula] KC60
    |    |--Latischisma Thomas 1940 KC60
    |    |    `--L. globosa Thomas 1940 W63c
    |    |--Tapinotomaria Batten 1956 KC60
    |    |    |--*T. rugosa Batten 1956 KC60
    |    |    `--T. mirabilis Batten 1958 PE02
    |    |--Portlockiella Knight 1945 BR05 [incl. Portlockia deKoninck 1881 non M’Coy 1846 KC60]
    |    |    |--*P. kentuckyensis Knight 1945 KC60
    |    |    |--‘Portlockia’ kamenkinsis G31
    |    |    `--‘Portlockia’ rotundata G31
    |    `--Shansiella Yin 1932 W63c
    |         |--*S. altispiralis Yin 1932 W63c
    |         |--S. carbonaria (Norwood & Pratten 1855) B79
    |         |--S. conica Batten 1958 W63c
    |         `--S. tabulata Batten 1958 W63c
    |--Pleurotomariidae [Pleurotomariae, Pleurotomariinae] A88
    |    |  i. s.: ‘Holopea’ obscura Hynda 1986 D94
    |    |--Pleurotomaria BR05
    |    |--Perotrochus A88
    |    |--Angyomphalus Cossmann 1916 P84
    |    |    `--*A. radians (deKoninck 1843) [=Euomphalus radians, Trepospira (*Angyomphalus) radians] KC60
    |    |--Mikadotrochus Lindholm 1927 HPA88
    |    |    |--*M. beyrichi (Hilger 1877) [=Pleurotomaria beyrichi] KC60
    |    |    `--M. hirasei HPA88
    |    |--Conotomaria Cox 1959 HPA88
    |    |    `--*C. mailleana (d’Orbigny 1843) [=Pleurotomaria mailleana] KC60
    |    |--Entemnotrochus Fischer 1885 HPA88
    |    |    `--*E. adansonianus (Crosse & Fischer 1861) [=Pleurotomaria adansoniana] KC60
    |    |--Bathrotomaria Cox 1956 KC60
    |    |    `--*B. reticulata (Sowerby 1821) [=Trochus reticulatus] KC60
    |    |--Obornella Cox 1959 KC60
    |    |    `--*O. plicopunctata (Eudes-Deslongchamps 1849) [=Pleurotomaria plicopunctata] KC60
    |    |--Chelotia Bayle in Fischer 1885 KC60
    |    |    `--*C. concava (Deshayes 1832) [=Pleurotomaria concava] KC60
    |    |--Pyrgotrochus Fischer 1885 KC60
    |    |    |--*P. bitorquatus (Eudes-Deslongchamps 1849) [=Pleurotomaria bitorquata] KC60
    |    |    `--P. luciensis C60
    |    `--Pleurotrochus Sherzer & Grabau 1909 S11, KC60
    |         |--P. atlanticus Rios & Matthews 1968 S11
    |         `--P. notialis Leme & Penna 1969 S11
    `--Phymatopleuridae W63c
         |--Dictyomaria capillaria (Conrad 1842) TTE93
         |--Paragoniozona Nelson 1947 KC60
         |    `--*P. nodolirata Nelson 1947 KC60
         |--Borestus Thomas 1940 [incl. Platypleurotomaria Wanner 1942] KC60
         |    `--*B. wrighti Thomas 1940 KC60
         |--Callitomaria Batten 1956 KC60
         |    `--*C. stanislavi Batten 1956 KC60
         |--Eymarella Cossmann 1897 [=Echetus Koken 1896 non Krøyer 1863] KC60
         |    `--*E. subscalariformis [=Pleurotomaria subscalariformis] KC60
         |--Lamellospira Batten 1958 KC60
         |    `--*L. conica Batten 1958 KC60
         |--Phymatopleura Girty 1939 BR05 [=Orestes Girty 1912 nec Blackinstone & Fryer 1880 nec Redtenbacher 1906 KC60]
         |    |--*P. nodosus (Girty 1911) [=*Orestes nodosus] BR17
         |    `--P. brazoensis W63c
         |--Euryalox Cossmann 1896 TE01 [=Sagana Koken 1896 non Walker 1855 KC60]
         |    |--*E. juvavica (Koken 1894) TE01 [=Pleurotomaria juvavica KC60, *Sagana juvavica KC60]
         |    `--E. subcancellata [=Pleurotomaria subcancellata d’Orbigny non Morris 1845] TE01
         |--Leptomaria Eudes-Deslongchamps 1864 TTE93, KC60
         |    |--*L. amoena (Eudes-Deslongchamps 1849) [=Pleurotomaria amoena] KC60
         |    `--L. sublevis Traub 1979 TTE93
         |--Codinella Kittl 1899 TE01
         |    |--*C. generelli (Stoppani 1858) [=Trochus generelli] TE01
         |    |--C. leda (Kittl 1894) TE01
         |    `--C. mammiformis TE01
         |--Glyptotomaria Knight 1945 KC60
         |    |--*G. (Glyptotomaria) apiarium Knight 1945 KC60
         |    `--G. (Dictyotomaria Knight 1945) KC60
         |         `--G. (*D.) scitula (Meek & Worthen 1861) [=Pleurotomaria scitula] KC60
         `--Discotomaria Batten 1956 P85
              |--*D. basisulcata Batten 1956 KC60
              |--D. klobukowskii (Mansuy 1912) P85, B79 (see below for synonymy)
              |--D. turritus (Delpey 1942) P85, B79 [=Euomphalus turritus P85, Discotropis turritus B79]
              `--D. xingwenensis Pan 1985 P85

Discotomaria klobukowskii (Mansuy 1912) P85, B79 [=Euomphalus klobukowskii P85, Discotropis klobukowskii B79; incl. E. rectangulus P85]

Trochotoma Eudes-Deslongchamps 1842 BR05 [incl. Ditremaria d’Orbigny 1843 BR05, Rimulus d’Orbigny 1842 (n. n.) KC60]

Zygites delphinula (Laube 1868) TE01 [=Pleurotomaria delphinula TE01, Delphinula cancellata Klipstein 1843 non Gray 1828 KC60, BR17, *Zygites cancellatus BR17]

*Type species of generic name indicated

REFERENCES

[A88] Askew, T. M. 1988. A new species of pleurotomariid gastropod from the western Atlantic. Nautilus 102 (3): 89–91.

[B79] Batten, R. L. 1979. Gastropods from Perak, Malaysia. Part 2. The trochids, patellids and neritids. American Museum Novitates 2685: 1–26.

[BR05] Bouchet, P., & J.-P. Rocroi. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1–2): 1–397.

[BR17] Bouchet, P., J.-P. Rocroi, B. Hausdorf, A. Kaim, Y. Kano, A. Nützel, P. Parkhaev, M. Schrödl & E. E. Strong. 2017. Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia 61 (1–2): 1–526.

[C60] Cox, L. R. 1960. Gastropoda: general characteristics of Gastropoda. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt I. Mollusca 1: Mollusca—General Features, Scaphopoda, Amphineura, Monoplacophora, Gastropoda—General Features, Archaeogastropoda and some (mainly Paleozoic) Caenogastropoda and Opisthobranchia pp. I84–I169. Geological Society of America, and University of Kansas Press.

[D94] Dzik, J. 1994. Evolution of 'small shelly fossils' assemblages of the early Paleozoic. Acta Palaeontologica Polonica 39 (3): 247–313.

[G31] Grabau, A. W. 1931. The Permian of Mongolia: A report on the Permian fauna of the Jisu Honguer limestone of Mongolia and its relations to the Permian of other parts of the world. American Museum of Natural History: New York.

[HPA88] Harasewych, M. G., S. A. Pomponi & T. M. Askew. 1988. Spongivory in pleurotomariid gastropods. Nautilus 102 (3): 92–98.

[KC60] Knight, J. B., L. R. Cox, A. M. Keen, R. L. Batten, E. L. Yochelson & R. Robertson. 1960. Gastropoda: systematic descriptions. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt I. Mollusca 1: Mollusca—General Features, Scaphopoda, Amphineura, Monoplacophora, Gastropoda—General Features, Archaeogastropoda and some (mainly Paleozoic) Caenogastropoda and Opisthobranchia pp. I169–I331. Geological Society of America, and University of Kansas Press.

[PE02] Pan, H.-Z., & D. H. Erwin. 2002. Gastropods from the Permian of Guangxi and Yunnan Provinces, south China. Journal of Paleontology 76 (Memoir 56): 1–49.

[P85] Pan Y.-T. 1985. Fossil Gastropoda from the Permian of Sichuan, Guizhou, Yunnan and Guangxi. Acta Palaeontologica Sinica 24 (1): 29–37.

[P84] Peel, J. S. 1984. Autecology and systematics of a new Silurian anomphalid gastropod from western North Greenland. Rapp. Grønlands geol. Unders. 121: 77–87.

[S11] Simone, L. R. L. 2011. Phylogeny of the Caenogastropoda (Mollusca), based on comparative morphology. Arquivos de Zoologia 42 (4): 161–323.

[TE01] Tong, J., & D. H. Erwin. 2001. Triassic gastropods of the southern Qinling Mountains, China. Smithsonian Contributions to Paleobiology 92: 1–47.

[TTE93] Tracey, S., J. A. Todd & D. H. Erwin. 1993. Mollusca: Gastropoda. In: Benton, M. J. (ed.) The Fossil Record 2 pp. 131–167. Chapman & Hall: London.

[W63a] Waterhouse, J. B. 1963a. Permian gastropods of New Zealand. Part 1—Bellerophontacea and Euomphalacea. New Zealand Journal of Geology and Geophysics 6 (1): 88–112.

[W63b] Waterhouse, J. B. 1963b. Permian gastropods of New Zealand. Part 2—Pleurotomariacea (in part). New Zealand Journal of Geology and Geophysics 6 (2): 115–154.

[W63c] Waterhouse, J. B. 1963c. Permian gastropds of New Zealand. Part 3—Pleurotomariacea (concluded). New Zealand Journal of Geology and Geophysics 6 (4): 587–622.

Last updated: 19 June 2021.

Vetigastropoda

Temnotropis carinata, from Knight et al. (1960).

Belongs within: Gastropoda.
Contains: Seguenziidae, Chilodontaidae, Eucyclidae, Pleurotomarioidea, Porcellioidea, Pseudophoridae, Planitrochidae, Ptychomphalidae, Murchisoniidae, Plethospiridae, Gosseletinidae, Luciellidae, Eotomariinae, Neilsoniinae, Lepetelloidea, Fissurellidae, Lepetodriloidea, Scissurellidae, Anatomidae, Haliotis, Holopeidae, Proconulidae, Microdomatidae, Trochidae, Ataphridae, Angariidae, Arene, Conradiidae, Margaritidae, Skeneidae, Liotiidae, Nododelphinulidae, Anomphalinae, Pycnomphalinae, Turbininae, Calliostomatinae, Margarella, Solariella, Colloniinae, Crossostomatinae, Tegula (Chlorostoma), Tegula (Omphalius), Phasianella, Tricolia.

The Vetigastropoda are a major clade within the gastropods that contains the greater part of the paraphyletic 'Archaeogastropoda' of older classifications. Living members of the clade do not undergo a planktotrophic stage in the life cycle, and have a paucispiral protoconch (Bouchet et al. 2005). Many vetigastropods have a nacreous inner layer to the shell. Specialised epipodial sensory organs are present on the foot.

The shell of many vetigastropods bears an exhalant opening in the form of a slit or selenizone. The Pleurotomariida are primarily fossil forms, represented in the modern fauna only by members of the Pleurotomariidae. Members of the Lepetellida have few-whorled shells with a more or less expanded aperture, with the shell becoming entirely limpet-like in some lineages. The Scissurelloidea bear low shells with only a thin inner nacreous layer, a slit or hole on the outer lip, and a round, multispiral operculum (Knight et al. 1960).

Members of the Trochoidea lack any trace of an exhalant slit. They have a single ctenidium whereas Pleurotomariida and Lepetellida have paired ctenidia albeit with the right ctenidium variously reduced (Knight et al. 1960). The Trochoidea have historically been divided between superfamilies Trochoidea and Turbinoidea on the basis of whether the operculum is corneous or calcareous, respectively, but this division is not currently supported. The Seguenzioidea, small to medium-sized deep-sea vetigastropods, resemble Trochoidea in the presence of a single ctenidium but retain a number of plesiomorphic characters such as the presence of separate epipodial and sensory-organ bearing tentacles on the foot (in Trochoidea, the epipodial sensory organs are borne directly on the epipodial tentacles).

<==Vetigastropoda (see below for synonymy)
    |--Pleurotomariida BR17
    |    |--Pleurotomarioidea GO06
    |    |--Porcellioidea BR17
    |    |--Sinuspira Perner 1907 KC60 (see below for synonymy)
    |    |    `--*S. tenera Barrande 1907 BR17 [=Murchisonia (*Sinuspira) tenera KC60]
    |    |--Pseudophoroidea [Pseudophoracea] BR17
    |    |    |--Pseudophoridae BR17
    |    |    `--Planitrochidae BR17
    |    |--Ptychomphaloidea BR17
    |    |    |--Ptychomphalidae BR17
    |    |    `--Rhaphistomella Kittle 1891 KC60 [=Raphistomella Diener 1926 KC60; Rhaphistomellidae BR17]
    |    |         |--*R. radians (Wissmann 1841) [=Pleurotomaria radians] BR17
    |    |         `--R. subdecussata [=Pleurotomaria subdecussata] G31
    |    |--Schizogonioidea BR17
    |    |    |--Schizogonium Koken 1889 [Schizogoniidae] BR05
    |    |    |    `--*S. scalare (Münster 1841) [=Pleurotomaria scalaris] KC60
    |    |    `--Pseudowortheniella Bandel 2009 [Pseudowortheniellidae] BR17
    |    |         `--*P. rarissima (Kittl 1891) [=Worthenia rarissima] BR17
    |    |--Murchisonioidea [Murchisoniacea, Murchisoniina] BR17
    |    |    |--Murchisoniidae BR05
    |    |    |--Plethospiridae BR17
    |    |    |--Farewellia Frýda & Blodgett 2004 FB04 [Farewelliidae BR17]
    |    |    |    `--*F. heidelbergerae Frýda & Blodgett 2004 FB04
    |    |    `--Ptychocaulus Perner 1907 KC60 [Ptychocaulidae BR17]
    |    |         `--*P. verneuili (Barrande 1907) [=Murchisonia verneuili] BR17
    |    `--Eotomarioidea BR17
    |         |  i. s.: Devonorhineoderma orbignyana F12
    |         |         Zlichomphalina F12
    |         |--Gosseletinidae BR05
    |         |--Luciellidae BR05
    |         |--Pseudoschizogonium Kutassy 1937 KC60 [Pseudoschizogoniidae BR17]
    |         |    `--*P. turriculatum Kutassy 1937 KC60
    |         |--Wortheniella Schwardt 1992 B00 [Wortheniellidae BR17]
    |         |    `--*W. coralliophila (Kittl 1891) [=Worthenia coralliophila] BR17
    |         |--Phanerotrematidae BR05
    |         |    |--Brachytomaria Koken 1925 KC60
    |         |    |    `--*B. baltica (Verneuil 1845) KC60, TTE93 [=Pleurotomaria baltica KC60]
    |         |    `--Phanerotrema Fischer 1885 BR05
    |         |         |--*P. labrosum (Hall 1860) [=Pleurotomaria labrosa] KC60
    |         |         |--P. austrralis Etheridge 1891 F71
    |         |         `--P. ornatum Sayre 1930 TTE93
    |         `--Eotomariidae [Eotomacea] KC60
    |              |  i. s.: Zhonghuaspira Pan & Erwin 2002 PE02
    |              |           |--*Z. gibbicircella (Wang & Xi 1980) [=Guizhouspira gibbicircella] PE02
    |              |           `--Z. gibbicircelloides (Wang 1982) [=Guizhouspira gibbicircelloides] PE02
    |              |         Quadricarina Blodgett & Johnson 1992 FB04
    |              |           |--Q. (Quadricarina) FB04
    |              |           |    |--*Q. (Q.) glabrobasis Blodgett & Johnson 1992 FB04
    |              |           |    |--Q. (Q.) lenticularis (Goldfuss 1844) [=Pleurotomaria lenticularis] FB04
    |              |           |    `--Q. (Q.) noklebergi Frýda & Blodgett 2004 FB04
    |              |           `--Q. (Blodgettina) reticulata Frýda 2000 FB04
    |              |         Paraoehlertia Frýda 1998 FB04
    |              |           `--*P. parva Frýda 1998 FB04
    |              |         Balbinipleura Bandel & Frýda 1996 FB04
    |              |           |--*B. bohemica Bandel & Frýda 1996 FB04
    |              |           |--B. koninckii Bandel & Frýda 1996 FB04
    |              |           |--B. krawczynskii Frýda & Blodgett 2004 FB04
    |              |           `--B. sextalineata Heidelberger 2001 FB04
    |              |         Ulrichospira Donald 1905 FB04
    |              |           `--*U. similis Donald 1905 KC60
    |              |         Kersadiella Blodgett, Frýda & Racheboeuf 1999 FB04
    |              |         Lacunospira Batten 1958 KC60
    |              |           `--*L. alta Batten 1958 KC60
    |              |--Eotomariinae BR05
    |              `--Neilsoniinae BR05
    `--+--+--Bathymargarites symplector GO06
       |  `--Lepetellida BR17
       |       |  i. s.: Lepetelloidea BR17
       |       |--Fissurellidae GO06
       |       `--+--+--Lepetodriloidea GO06
       |          |  `--Scissurelloidea BR05
       |          |       |--Scissurellidae GO06
       |          |       |--Anatomidae BR05
       |          |       `--Larochea Finlay 1927 BR05 [Larocheidae BR17, Larocheinae]
       |          |            |--*L. miranda Finlay 1927 P61
       |          |            `--L. secunda Powell 1937 P61
       |          `--Haliotoidea [Haliotacea] BR05
       |               |--Haliotidae [Haliotidia, Haliotinae] BR05
       |               |    |--Haliotis GO06
       |               |    `--Chelinotus Swainson 1840 [Chelinoti] BR05
       |               `--Temnotropis Laube 1868 [Temnotropidae] BR05
       |                    |--*T. carinata (Münster 1841) [=Sigaretus carinatus] KC60
       |                    |--T. bicarinata Laube 1869 TTE93
       |                    |--T. costellata Gemmellaro 1890 W63a
       |                    `--T. transitoria Gemmellaro 1890 W63a
       `--Trochoidea [Calliostomatoidei, Ciliipedata, Microdomatacea, Microdomatoidea, Trochida, Turbinoidea] PL96
            |  i. s.: ‘Fossarus’ dixoni Wrigley 1942 TTE93
            |--Holopeidae PE02
            |--Proconulidae BR05
            |--Microdomatidae BR05
            |--Trochidae BR05
            |--Ataphridae MCS96
            |--Angariidae BR17
            |--Arene BR17
            |--Conradiidae BR17
            |--Margaritidae BR17
            |--Skeneidae BR17
            |--Liotiidae BR05
            |--Nododelphinulidae BR17
            |--Tychobrahea Horný 1992 [Tychobraheidae] BR05
            |    `--*T. aerumnans Horný 1992 BR17
            |--Eucochlis Knight 1933 [Eucochlidae] BR05
            |    `--*E. perminuta Knight 1933 KC60
            |--Velainella Vasseur 1880 BR05 [=Velainiella (l. c.) KC60; Velainellidae BR05, Velainellinae, Vellainelloidea]
            |    `--*V. columnaris Vasseur 1880 KC60
            |--Araeonema Knight 1933 KC60 (see below for synonymy)
            |    `--*A. virgatum Knight 1933 KC60
            |--Epulotrochus Cossmann 1918 KC60 [Epulotrochidae BR17]
            |    `--*E. epulus (d’Orbigny 1850) [=Trochus epulus] KC60
            |--Sclarotrarda Gründel, Keupp & Lang 2017 [Sclarotrardidae] BR17
            |    `--*S. coronilla (Brösamlen 1909) [=Liotia coronilla] BR17
            |--Metriomphalus Cossmann 1918 [Metriomphalidae] BR17
            |    |--*M. davoustii (d’Orbigny 1850) [=Turbo davoustii] BR17
            |    |--M. segregatus (Hébert & Eudes-Deslongchamps 1860) GK02
            |    `--M. textorius (Broili 1907) TTE93
            |--Elasmonematidae BR05
            |    |--Elasmonema Fischer 1885 BR05 [=Callonema Hall 1879 non Conrad 1875 KC60]
            |    |    `--*E. bellatulum (Hall 1861) [=Loxonema bellatulum, *Callonema bellatulum] KC60
            |    |--Anematina Knight 1933 B79
            |    |    `--*A. proutana (Hall 1858) B79 [=Holopea proutana KC60]
            |    |--Streptotrochus Perner 1907 KC60
            |    |    `--*S. rugulosus Perner 1907 KC60
            |    `--Discordichilus Cossmann 1918 KC60
            |         `--*D. mollis (Lindström 1884) [=Trochus mollis] KC60
            |--Anomphalidae [Anomphalacea, Anomphaloidea] P84
            |    |  i. s.: Cycloscena Fletcher 1958 W63b
            |    |           |--*C. anomphala Fletcher 1958 W63b
            |    |           |--C. costata Waterhouse 1963 W63b
            |    |           |--C. glabra Waterhouse 1963 W63b
            |    |           `--‘Pleurotomaria’ rotunda Etheridge 1872 [=Platyschisma rotunda] W63b
            |    |         Eiselia Dietz 1911 P84
            |    |           `--*E. dyadica Dietz 1911 KC60
            |    |--Anomphalinae P84
            |    `--Pycnomphalinae P84
            |--Turbinidae [Astylophthalma] BR17
            |    |  i. s.: Natiria de Koninck 1881 TE01 [incl. Fritschia Picard 1904 KC60]
            |    |           |--*N. lirata (Phillips 1836) [=Natica lirata] TE01
            |    |           `--N. costata TE01
            |    |         Pterocheilos Moore 1867 [=Mooria Cossmann 1899, Tinochilus Fischer 1885] KC60
            |    |           `--*P. primus Moore 1867 [=*Mooria prima, *Tinochilus primus] KC60
            |    |--Turbininae BR05
            |    |--Moreanellus Fischer & Weber 1997 [Moreanellinae] BR05
            |    |    `--*M. moreanus (d’Orbigny 1850) [=Trochus moreanus] BR17
            |    `--Prisogaster Mörch 1850 BR05 [=Amyxa Troschel 1852 KC60; Prisogasterinae, Prisogastrinae BR05]
            |         `--*P. niger (Wood 1828) [=Turbo niger, *Amyxa nigra] KC60
            |--Calliostomatidae BR17
            |    |--Calliostomatinae BR17
            |    |--Margarella BR17
            |    |--Callotrochus Kutassy 1938 BR17 [=Mesotrochus Kutassy 1927 non Wasmann 1890 KC60; Callotrochinae BR17]
            |    |    `--*C. triadicus (Kutassy 1927) [=Trochus triadicus, *Mesotrochus triadicus] BR17
            |    |--Fautrix Marshall 1995 BR05 [Fautricinae BR17, Fautricini]
            |    |    `--*F. candida Marshall 1995 BR17
            |    |--Thysanodonta Marshall 1988 BR05 [Thysanodontidae, Thysanodontinae BR17]
            |    |    `--*T. aucklandica Marshall 1988 BR17
            |    `--Xeniostoma McLean 2012 [Xeniostomatinae] BR17
            |         `--*X. inexpectans McLean 2012 BR17
            |--Solariellidae [Solariellinae] BR05
            |    |--Solariella BR05
            |    |--Cidarina Dall 1909 KC60
            |    |    `--*C. cidaris (Carpenter 1864) KC60 [=Margarita cidaris KC60, Margarites cidaris O27]
            |    `--Minolia Adams 1860 H92 [=Minosia Dunker 1882 KC60; incl. Minolops Iredale 1929 KC60; Minoliinae]
            |         |--*M. punctata Adams 1860 H92
            |         |--M. aspecta C64
            |         |--M. casta (Nevill & Nevill 1874) H92
            |         |--M. gradata Sowerby 1895 H92
            |         |--M. holdsworthana Nevill & Nevill 1871 H92
            |         |--M. nedyma Melvill 1897 BD86
            |         |--M. singaporensis (Pilsbry 1889) [=Monilea (Minolia) singaporensis] BC01
            |         |--M. strigata Tenison-Woods 1878 F71
            |         `--M. vitiliginea (Menke 1843) H92
            |--Colloniidae BR17
            |    |--Colloniinae BR05
            |    |--Crossostomatinae BR17
            |    |--Lewisiella Stoliczka 1868 KC60 [Lewisiellinae BR17]
            |    |    `--*L. conica (d’Orbigny 1853) [=Pitonnillus conicus] BR17
            |    |--Liotipoma McLean & Kiel 2007 [Liotipomatinae] BR17
            |    |    `--*L. wallisensis McLean & Kiel 2007 BR17
            |    |--Petropoma Gabb 1877 BR05 [Petropomatinae BR17, Petropominae]
            |    |    `--*P. peruanum Gabb 1877 KC60
            |    `--Moelleria Jeffreys 1865 non Scop. 1777 (ICBN) BR05 [Moelleriinae BR17]
            |         |--*M. costulata (Möller 1842) KC60 [=Margarita costulata KC60, Cyclostrema costulatum J64]
            |         |--M. drusiana Dall 1919 O27
            |         |--M. laevigata Jeffreys in Friele 1875 N79
            |         `--M. quadrae Dall 1897 O27
            |--Tegula Lesson 1832 BR05 [Tegulidae BR17, Tegulinae, Tegulini]
            |    |  i. s.: T. excavata (Lamarck 1822) HW78
            |    |         T. fasciata WE63
            |    |         T. ‘lugubris’ (Philippi 1844) [=Trochus lugubris non Gmelin 1791] F27
            |    |--T. (Tegula) BR17
            |    |    |--*T. (T.) elegans Lesson 1832 BR17
            |    |    `--T. (T.) pellisserpentis (Wood 1828) KC60 [=Trochus pellisserpentis KC60; incl. T. strigilatus C64]
            |    |--T. (Agathistoma Olsson & Harbison 1953) KC60
            |    |    |--T. (*A.) viridula (Gmelin 1791) [=Trochus viridulus] KC60
            |    |    |--T. (A.) corteziana McLean 1970 BC01
            |    |    |--T. (A.) felipensis McLean 1970 BC01
            |    |    |--T. (A.) picta McLean 1970 BC01
            |    |    `--T. (A.) verrucosa McLean 1970 BC01
            |    |--T. (Chlorostoma) KC60
            |    |--T. (Omphalius) KC60
            |    `--T. (Promartynia Dall 1909) KC60
            |         `--T. (*P.) pulligo (Gmelin 1791) [=Trochus pulligo] KC60
            |              |--T. p. pulligo O27
            |              `--T. p. taylori Oldroyd 1922 O27
            `--Phasianellidae [Phasianelloidea] BR05
                 |  i. s.: Pseudophasianus Cossmann 1918 KC60
                 |           `--*P. elatus (Fuchs 1870) [=Turbo elatus] KC60
                 |--Phasianella BR05
                 |--Gabrieloninae BR05
                 |    |--Eugabrielona BR05
                 |    `--Gabrielona Iredale 1917 BR05
                 |         `--*G. nepeanensis (Gatliff & Gabriel 1908) [=Phasianella nepeanensis] KC60
                 `--Tricoliinae [Tricoliidae] BR05
                      |--Tricolia BR05
                      |--Pellax Finlay 1926 KC60
                      |    |--*P. huttoni (Pilsbry 1888) P61 [=Phasianella huttoni P61, Tricolia (*Pellax) huttoni KC60]
                      |    |--P. rosea [=Tricolia rosea] F26
                      |    `--P. virgo [=Tricolia huttoni] F26
                      `--Eulithidium Pilsbry 1898 KC60
                           |--E. dianthum (McLean 1970) [=Tricolia diantha] BC01
                           |--E. luridum (Dall 1897) [=Eucosmia lurida, Phasianella (Eulithidium) lurida] O27
                           |--E. pterocladicum (Robertson 1958) [=Tricolia affinis pterocladica] BC01
                           |--E. substriatum (Carpenter 1864) (see below for synonymy) O27
                           `--E. typicum (Dall 1908) (see below for synonymy) O27

Vetigastropoda incertae sedis:
  Micromphalus Knight 1945 KC60 [Micromphalidae BR17]
    `--*M. turris Knight 1945 KC60
  Seguenzioidea [Seguenziida, Seguenziiformes, Seguenziina, Seguenzinina] PL96
    |--Seguenziidae BR05
    |--Chilodontaidae BR05
    |--Eucyclidae BR17
    |--Laubella Kittl 1891 [Laubellidae] BR05
    |    `--*L. delicata (Laube 1868) [=Pleurotomaria delicata] KC60
    |--Eudaronia Cotton 1945 [Eudaroniidae, Eudaroniinae] BR17
    |    `--*E. jaffaensis (Verco 1909) [=Cyclostrema jaffaensis] BR17
    |--Lanascala Bandel 1992 BR05 [Lanascalidae BR17]
    |    `--*L. cassiana Bandel 1992 BR17
    |--Pseudoturcica Bandel 2010 [Pseudoturcicidae] BR17
    |    `--*P. wareni (Kaim 2004) [=Turcica wareni] BR17
    |--Sabrinella Bandel 1993 [Sabrinellidae] BR17
    |    `--*S. doris (Laube 1869) [=Delphinula doris] BR17
    |--Cataegis McLean & Quinn 1987 BR05 [Cataegidae BR17, Cataeginae]
    |    |--*C. toreuta McLean & Quinn 1987 BR17
    |    `--C. meroglypta McLean & Quinn 1987 BR05
    |--Choristellidae [Choristellinae] BR17
    |    |--Bichoristes BR05
    |    `--Choristella Bush 1897 BR05
    |         `--*C. leptalea Bush 1897 BR17
    |--Eucycloscala Cossmann 1895 PE02 (see below for synonymy)
    |    |--*E. binodosa (Münster 1841) [=Trochus binodosus] KC60
    |    |--E. asiatica Batten 1985 PE02
    |    `--E. medionodosus Batten 1985 PE02
    |--Eunemopsis Kittl 1891 TE01 [Eunemopsidae BR17]
    |    |--*E. epaphus (Laube 1869) [=Turbo epaphus] TE01
    |    |--E. dolomitica Kittl 1891 TE01
    |    `--E. praecurrens TE01
    |--Pendromidae BR17
    |    |--Rugulina C96
    |    `--Pendroma Dall 1927 BR05
    |         `--*P. perplexa Dall 1927 BR17
    `--Trochaclididae [Trochaclidoidea] BR17
         |--Trochaclis Thiele 1912 [Trochaclidinae] BR05
         |    `--*T. antarctica Thiele 1912 BR17
         `--Acremodonta Marshall 1983 [Acremodontinae] BR05
              `--*A. crassicosta (Powell 1937) [=Thoristella crassicosta] BR17
  Spirotomaria Koken 1925 KC60
    `--*S. rudissima (Koken 1897) [=Pleurotomaria rudissima] KC60
  Ptychozone Perner 1907 KC60
    `--*P. aberrans (Perner 1903) [=Worthenia aberrans] KC60
  Platyconus Perner 1907 KC60
    `--*P. incumbens (Perner 1907) [=Pleurotomaria (*Platyconus) incumbens] KC60
  Gyroma Oehlert 1888 KC60
    `--*G. baconnierensis (Oehlert 1888) [=Pleurotomaria baconnierensis] KC60
  Trachybembix Böhm 1895 [=Trachybembyx Diener 1916] KC60
    `--*T. junonis (Kittl 1894) [=Pleurotomaria junonis] KC60
  Transylvanella Kutassy 1937 KC60
    `--*T. acmaeiformis Kutassy 1937 KC60

Nomen nudum: Callonema elevatum TTE93

Araeonema Knight 1933 KC60 [incl. Palaeoturbina Wenz 1938 KC60, Turbina deKoninck 1881 non Brown in Herrmannsen 1847 KC60; Araeonematidae BR17]

Eucycloscala Cossmann 1895 PE02 [incl. Trochoscala Koken 1897 KC60, Urceolabrum Wade 1917 KC60; Eucycloscalidae, Eucycloscalinae]

Eulithidium substriatum (Carpenter 1864) [=Eucosmia substriata, Phasianella (Eulithidium) substriata] O27

Eulithidium typicum (Dall 1908) [=Phasianela (Eulithidium) typica, Eucosmia variegata Carpenter 1864 non P. variegata Lamarck 1822] O27

Sinuspira Perner 1907 KC60 [incl. Morania Horny 1953 non Walcott 1919 KC60; Sinuspiridae, Sinuspirina, Sinuspiroidea BR17]

Vetigastropoda [Amberleyacea, Amberleyata, Amberleyoidea, Astylacea, Auriformes, Aristerobranchia, Choristelloidei, Ctenidiobranchia, Fissibranchiata, Fissurelloidei, Haliotoideae, Haliotoidei, Pleurotomariacea, Pleurotomariina, Zeugobranchia, Zygobranchia]

*Type species of generic name indicated

REFERENCES

[B00] Bandel, K. 2000. The new family Cortinellidae (Gastropoda, Mollusca) connected to a review of the evolutionary history of the subclass Neritimorpha. Neues Jahrbuch für Geologie und Paläontologie—Abhandlungen 217 (1): 111–129.

[BD86] Barash, A., & Z. Danin. 1986. Further additions to the knowledge of Indo-Pacific Mollusca in the Mediterranean Sea (Lessepsian migrants). Spixiana 9 (2): 117–141.

[B79] Batten, R. L. 1979. Gastropods from Perak, Malaysia. Part 2. The trochids, patellids and neritids. American Museum Novitates 2685: 1–26.

[BR05] Bouchet, P., & J.-P. Rocroi. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1–2): 1–397.

[BR17] Bouchet, P., J.-P. Rocroi, B. Hausdorf, A. Kaim, Y. Kano, A. Nützel, P. Parkhaev, M. Schrödl & E. E. Strong. 2017. Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia 61 (1–2): 1–526.

[BC01] Boyko, C. B., & J. R. Cordeiro. 2001. Catalog of Recent type specimens in the Division of Invertebrate Zoology, American Museum of Natural History. V. Mollusca, part 2 (class Gastropoda [exclusive of Opisthobranchia and Pulmonata], with supplements to Gastropoda [Opisthobranchia], and Bivalvia). Bulletin of the American Museum of Natural History 262: 1–170.

[C64] Carpenter, P. P. 1864. Supplementary report on the present state of our knowledge with regard to the Mollusca of the west coast of North America. Report of the British Association for the Advancement of Science 33: 517–686.

[C96] Crame, J. A. 1996. Evolution of high-latitude molluscan faunas. In: Taylor, J. D. (ed.) Origin and Evolutionary Radiation of the Mollusca pp. 119–131. Oxford University Press: Oxford.

[F26] Finlay, H. J. 1926. A further commentary on New Zealand molluscan systematics. Transactions and Proceedings of the New Zealand Institute 57: 320–485.

[F27] Finlay, H. J. 1927. New specific names for austral Mollusca. Transactions and Proceedings of the New Zealand Institute 57: 488–533.

[F71] Fletcher, H. O. 1971. Catalogue of type specimens of fossils in the Australian Museum, Sydney. Australian Museum Memoir 13: 1–167.

[F12] Frýda, J. 2012. Phylogeny of Palaeozoic gastropods inferred from their ontogeny. In: Talent, J. A. (ed.) Earth and Life: Global biodiversity, extinction intervals and biogeographic perturbations through time pp. 395–435. Springer.

[FB04] Frýda, J., & R. B. Blodgett. 2004. New Emsian (Late Early Devonian) gastropods from Limestone Mountain, Medfra B-4 quadrangle, west-central Alaska (Farewell Terrane), and their paleobiogeographic affinities and evolutionary significance. Journal of Paleontology 78 (1): 111–132.

[GO06] Giribet, G., A. Okusu, A. R. Lindgren, S. W. Huff, M. Schrödl & M. K. Nishiguchi. 2006. Evidence for a clade composed of molluscs with serially repeated structures: monoplacophorans are related to chitons. Proceedings of the National Academy of Sciences of the USA 103 (20): 7723–7728.

[G31] Grabau, A. W. 1931. The Permian of Mongolia: A report on the Permian fauna of the Jisu Honguer limestone of Mongolia and its relations to the Permian of other parts of the world. American Museum of Natural History: New York.

[GK02] Gründel, J., & T. Kowalke. 2002. Palaeorissoinidae, a new family of marine and brackish water Rissooidea (Gastropoda, Littorinimorpha). Neues Jahrbuch für Geologie und Paläontologie Abhandlungen 226 (1): 43–57.

[H92] Herbert, D. G. 1992. Revision of the Umboniinae in southern Africa and Mozambique (Mollusca: Prosobranchia: Trochidae). Annals of the Natal Museum 33 (2): 379–459.

[HW78] Hoffman, D. L., & P. J. Weldon. 1978. Flight responses of two intertidal gastropods (Prosobranchia: Trochidae) to sympatric predatory gastropods from Barbados. Veliger 20 (4): 361–366.

[J64] Jeffreys, J. G. 1864. The Upper Tertiary fossils at Uddevalla, in Sweden. Report of the British Association for the Advancement of Science 33 (Notices and Abstracts): 73–79.

[KC60] Knight, J. B., L. R. Cox, A. M. Keen, R. L. Batten, E. L. Yochelson & R. Robertson. 1960. Gastropoda: systematic descriptions. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt I. Mollusca 1: Mollusca—General Features, Scaphopoda, Amphineura, Monoplacophora, Gastropoda—General Features, Archaeogastropoda and some (mainly Paleozoic) Caenogastropoda and Opisthobranchia pp. I169–I331. Geological Society of America, and University of Kansas Press.

[MCS96] Monari, S., M. A. Conti & J. Szabó. 1996. Evolutionary systematics of Jurassic Trochoidea: the family Colloniidae and the subfamily Proconulidae. In: Taylor, J. D. (ed.) Origin and Evolutionary Radiation of the Mollusca pp. 199–204. Oxford University Press: Oxford.

[N79] Norman, A. M. 1879. The Mollusca of the fiords near Bergen, Norway. Journal of Conchology 2: 8–77.

[O27] Oldroyd, I. S. 1927. The Marine Shells of the West Coast of North America vol. 2 pt 3. Stanford University Press: Stanford University (California).

[PE02] Pan, H.-Z., & D. H. Erwin. 2002. Gastropods from the Permian of Guangxi and Yunnan Provinces, south China. Journal of Paleontology 76 (Memoir 56): 1–49.

[P84] Peel, J. S. 1984. Autecology and systematics of a new Silurian anomphalid gastropod from western North Greenland. Rapp. Grønlands geol. Unders. 121: 77–87.

[PL96] Ponder, W. F., & D. R. Lindberg. 1996. Gastropod phylogeny—challenges for the 90s. In: Taylor, J. D. (ed.) Origin and Evolutionary Radiation of the Mollusca pp. 135–154. Oxford University Press: Oxford.

[P61] Powell, A. W. B. 1961. Shells of New Zealand: An illustrated handbook 4^th ed. Whitcombe and Tombs Limited: Christchurch.

[TE01] Tong, J., & D. H. Erwin. 2001. Triassic gastropods of the southern Qinling Mountains, China. Smithsonian Contributions to Paleobiology 92: 1–47.

[TTE93] Tracey, S., J. A. Todd & D. H. Erwin. 1993. Mollusca: Gastropoda. In: Benton, M. J. (ed.) The Fossil Record 2 pp. 131–167. Chapman & Hall: London.

[WE63] Warmke, G. L., & D. S. Erdman. 1963. Records of marine mollusks eaten by bonefish in Puerto Rican waters. Nautilus 76 (4): 115–120.

[W63a] Waterhouse, J. B. 1963a. Permian gastropods of New Zealand. Part 2—Pleurotomariacea (in part). New Zealand Journal of Geology and Geophysics 6 (2): 115–154.

[W63b] Waterhouse, J. B. 1963b. Permian gastropods of New Zealand. Part 4—Platyceratacea, Anomphalacea, Neritacea, and correlations. New Zealand Journal of Geology and Geophysics 6 (5): 817–842.

Last updated: 26 February 2022.

Gosseletinidae

Baylea heterocarinata, from Pan (1985).

Belongs within: Vetigastropoda.

The Gosseletinidae are a group of gastropods known from the Middle Ordovician to the Triassic (Knight et al. 1960).

Characters (from Knight et al. 1960): Labral slit short, selenizone flat and generally without bordering threads.

<==Gosseletinidae
    |--Triangularia Frech 1894 [Triangulariinae] BR05
    |    `--*T. paradoxa Frech 1894 KC60
    |--Gosseletininae BR05
    |    |--Pseudocryptaenia Koken 1925 KC60
    |    |    `--*P. lahuseni (Koken 1897) [=Pleurotomaria lahuseni] KC60
    |    |--Stenoloron Oehlert 1888 KC60
    |    |    `--*S. viennayi (Oehlert 1888) [=Pleurotomaria (*Stenoloron) viennayi] KC60
    |    |--Platyloron Oehlert 1888 KC60
    |    |    `--*P. bishofi (Goldfuss 1844) [=Pleurotomaria bishofi] KC60
    |    |--Rufilla Koken 1896 KC60
    |    |    `--*R. densecincta Koken 1896 KC60
    |    |--Callistadia Knight 1945 P85
    |    |    |--*C. bella Knight 1945 KC60
    |    |    `--C. qinglongensis Pan 1985 P85
    |    |--Gosseletina Bayle in Fischer 1885 BR05 [=Gosseletia de Koninck 1883 non Barrois 1882 KC60]
    |    |    |--*G. callosa (de Koninck 1843) TE01 [=Pleurotomaria callosa TE01, *Gosseletia callosa KC60]
    |    |    |--G. calypso TE01
    |    |    |--G. dangchangensis Tong & Erwin 2001 TE01
    |    |    `--G. fasciolata (Münster 1841) TE01
    |    `--Baylea deKoninck 1883 P85 [=Yvania Fischer 1885 KC60]
    |         |--*B. yvanii (Léveillé 1835) [=Trochus yvanii, *Yvania yvanii] KC60
    |         |--B. heterocarinata Pan 1985 P85
    |         |--B. konincki [=Yvania konincki] F71
    |         |--B. multicarinata P85
    |         |--‘Yvania’ reticulata W63
    |         `--‘Yvania’ salmonensis W63
    `--Coelozoninae BR05
         |  i. s.: Umbotropis Perner 1903 FB04
         |           |--*U. albicans Perner 1903 KC60
         |           `--U. rihai Frýda & Bandel 1997 FB04
         |         Humboldtiella Blodgett & Frýda 1999 FB04
         |           `--*H. undulata (Roemer 1843) FB04
         |         Arctozone Frýda & Blodgett 2004 FB04
         |           `--*A. cooki Frýda & Blodgett 2004 FB04
         |--Planozonini [Planozonides] KC60
         |    |--Planozone Perner 1907 BR05
         |    |    `--*P. ramificans Perner 1907 KC60
         |    |--Pleurorima Perner 1907 KC60
         |    |    `--P. pragensis (Koken 1889) (see below for synonymy) KC60
         |    `--Cataschisma Branson 1909 [incl. Globispira Koken 1925] KC60
         |         |--*C. typa Branson 1909 KC60
         |         `--C. exquisita KC60
         `--Coelozonini [Coelozonides, Euryzoninae] BR05
              |--Coelozone Perner 1907 BR05
              |    `--*C. verna (Perner 1907) [=Pleurotomaria (*Coelozone) verna] KC60
              |--Euryzone Koken 1896 BR05
              |    `--*E. delphinuloides (Schlotheim 1820) [=Helicites delphinuloides] KC60
              |--Latitaenia Koken 1925 KC60
              |    `--*L. rotelloidea (Koken 1896) [=Pleurotomaria rotelloidea] KC60
              `--Conotoma Perner 1907 KC60
                   `--*C. eximia (Perner 1907) [=Pleurotomaria (Clathrospira) eximia] KC60

Pleurorima pragensis (Koken 1889) [=Pleurotomaria pragensis; incl. Pleurotomaria (*Pleurorima) migrans Perner 1907] KC60

*Type species of generic name indicated

REFERENCES

[BR05] Bouchet, P., J.-P. Rocroi, J. Frýda, B. Hausdorf, W. Ponder, Á. Valdés & A. Warén. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1-2): 1-397.

[F71] Fletcher, H. O. 1971. Catalogue of type specimens of fossils in the Australian Museum, Sydney. Australian Museum Memoir 13: 1-167.

[FB04] Frýda, J., & R. B. Blodgett. 2004. New Emsian (Late Early Devonian) gastropods from Limestone Mountain, Medfra B-4 quadrangle, west-central Alaska (Farewell Terrane), and their paleobiogeographic affinities and evolutionary significance. Journal of Paleontology 78 (1): 111-132.

[KC60] Knight, J. B., L. R. Cox, A. M. Keen, R. L. Batten, E. L. Yochelson & R. Robertson. 1960. Gastropoda: systematic descriptions. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt I. Mollusca 1: Mollusca—General Features, Scaphopoda, Amphineura, Monoplacophora, Gastropoda—General Features, Archaeogastropoda and some (mainly Paleozoic) Caenogastropoda and Opisthobranchia, pp. I169-I331. Geological Society of America, and University of Kansas Press.

[P85] Pan Y.-T. 1985. Fossil Gastropoda from the Permian of Sichuan, Guizhou, Yunnan and Guangxi. Acta Palaeontologica Sinica 24 (1): 29-37.

[TE01] Tong, J., & D. H. Erwin. 2001. Triassic gastropods of the southern Qinling Mountains, China. Smithsonian Contributions to Paleobiology 92: 1-47.

[W63] Waterhouse, J. B. 1963. Permian gastropds of New Zealand. Part 3—Pleurotomariacea (concluded). New Zealand Journal of Geology and Geophysics 6 (4): 587-622.

Neilsoniinae

Peruvispira robusta, from Waterhouse (1963).

Belongs within: Vetigastropoda.

The Neilsoniinae are a group of relatively high-spired gastropods with a labral slit, known from the Lower Carboniferous to Upper Permian (Knight et al. 1960).

<==Neilsoniinae [Neilsoniini]
    |--Neilsonia Thomas 1940 BR05
    |    `--*N. roscobiensis Thomas 1940 KC60
    |--Apachella Winters 1956 W63
    |    `--*A. translirata Winters 1956 KC60
    |--Carinella Waterhouse 1963 W63
    |    `--*C. bicarinata Waterhouse 1963 W63
    |--‘Pagodina’ Wanner 1942 nec van Beneden 1853 nec Stabile 1864 KC60
    |    `--*P. typus Wanner 1942 KC60
    |--Pareuryalox Haas 1953 KC60
    |    `--*P. perornata Haas 1953 KC60
    |--Planikeeneia Fletcher 1958 F71, KC60
    |    |--*P. minor Etheridge 1898 KC60, F71
    |    |--P. depressum (Dana 1849) [=Platyschisma depressum; incl. Plat. allandalensis] F71
    |    |--P. insculpta Fletcher 1958 F71
    |    `--P. occasa Fletcher 1958 [incl. Platyschisma rotundatum var. farleyensis Etheridge 1919] F71
    `--Peruvispira Chronic 1949 W63, PE02 [incl. Pleurocinctosa Fletcher 1958 W63]
         |  i. s.: *P. delicata Chronic 1949 PE02
         |         P. allandalensis (Fletcher 1958) [=Pleurocinctosa allandalensis] F71
         |         P. alticonica Wang & Xi 1980 PE02
         |         P. incerta [=Pagodina incerta] W63
         |         P. jucunda Pan & Erwin 2002 PE02
         |         P. kempseyensis W63
         |         P. lingshuiensis Pan 1985 PE02
         |         P. nuda (Dana 1849) [=Pleurocinctosa nuda] F71
         |         P. turrita Yu in Wang & Xi 1980 PE02
         |         P. umariensis (Reed 1928) W63
         |--+--P. allandalensis (Fletcher 1958) W63
         |  `--P. trifilata Dana 1847 W63 (see below for synonymy)
         `--+--P. elegans (Fletcher 1958) [=Pleurocinctosa elegans] W63
            `--+--P. robusta Waterhouse 1963 W63
               `--+--P. imbricata Waterhouse 1963 W63
                  |    |--P. i. imbricata W63
                  |    `--P. i. alta Waterhouse 1963 W63
                  |--P. modesta Waterhouse 1963 W63
                  `--P. volupta Waterhouse 1963 W63

Peruvispira trifilata Dana 1847 W63 [=Pleurotomaria trifilata KC60, *Pleurocinctosa trifilata KC60; incl. Pleurotomaria morrisiana W63]

*Type species of generic name indicated

REFERENCES

[BR05] Bouchet, P., J.-P. Rocroi, J. Frýda, B. Hausdorf, W. Ponder, Á. Valdés & A. Warén. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1-2): 1-397.

[F71] Fletcher, H. O. 1971. Catalogue of type specimens of fossils in the Australian Museum, Sydney. Australian Museum Memoir 13: 1-167.

[KC60] Knight, J. B., L. R. Cox, A. M. Keen, R. L. Batten, E. L. Yochelson & R. Robertson. 1960. Gastropoda: systematic descriptions. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt I. Mollusca 1: Mollusca—General Features, Scaphopoda, Amphineura, Monoplacophora, Gastropoda—General Features, Archaeogastropoda and some (mainly Paleozoic) Caenogastropoda and Opisthobranchia, pp. I169-I331. Geological Society of America, and University of Kansas Press.

[PE02] Pan, H.-Z., & D. H. Erwin. 2002. Gastropods from the Permian of Guangxi and Yunnan Provinces, south China. Journal of Paleontology 76 (Memoir 56): 1-49.

[W63] Waterhouse, J. B. 1963. Permian gastropds of New Zealand. Part 3—Pleurotomariacea (concluded). New Zealand Journal of Geology and Geophysics 6 (4): 587-622.

Eotomariinae

Ananias alticonica, from Pan & Erwin (2002).

Belongs within: Vetigastropoda.

The Eotomariinae are a group of gastropods known from the Middle Ordovician to Permian.

Characters (from Knight et al. 1960): Labral slit moderate to deep, umbilicus narrow or absent; ornament collabral and spiral elements, collabral dominant.

<==Eotomariinae
    |--Deseretospira Gordon & Yochelson 1987 [Deseretospirides, Deseretospirini] BR05
    |--Ptychomphalina Bayle 1885 [Ptychomphalininae, Ptychomphalinini] BR05
    |--Glabrocingulum Thomas 1940 [Glabrocingulides, Glabrocingulini] BR05
    |    `--*G. beggi Thomas 1940 KC60
    `--Eotomariini [Eotomariides, Liospirinae] BR05
         |--Eotomaria Ulrich & Scofield 1897 BR05 [incl. Spiroraphe Perner 1907 KC60]
         |    `--*E. canalifera Ulrich in Ulrich & Scofield 1897 KC60
         |--Pseudobaylea Dickins 1963 W63
         |    `--*P. freneyensis Dickins 1963 [=Mourlonia (Pseudobaylea) freneyensis] W63
         |--Arastra Stoyanow 1948 KC60
         |    `--*A. torquata Stoyanow 1948 KC60
         |--Trepospira Ulrich & Scofield 1897 [incl. Kansana Tasch 1953] KC60
         |    `--*T. sphaerulata (Conrad 1842) [=Pleurotomaria sphaerulata] KC60
         |--Bembexia Oehlert & Oehlert 1888 FB04
         |    `--*B. larteti (Munier-Chalmas 1876) [=Pleurotomaria larteti] KC60
         |--Liospira Ulrich & Scofield 1897 BR05 [incl. Eocryptaenia Koken 1925 KC60]
         |    |--*L. micula (Hall 1862) [=Pleurotomaria micula] KC60
         |    `--L. angustata Ulrich & Scofield 1897 YN83
         |--Clathrospira Ulrich & Scofield 1897 KC60
         |    |--*C. subconica (Hall 1847) [=Pleurotomaria subconica] KC60
         |    |--C. angulata Yu & Ning 1983 YN83
         |    `--C. conica YN83
         `--Ananias Knight 1945 W63, PE02
              |--*A. welleri (Newell 1935) PE02 [=Phanerotrema welleri PE02, Glabrocingulum (*Ananias) welleri KC60]
              |--A. alticonica (Wang in Wang & Xi 1980) [=Guizhouspira alticonica] PE02
              |--A. campbelli (Waterhouse 1963) [=Glabrocingulum (Ananias) campbelli] W63
              |--A. gibber Chronic 1952 W63
              |--A. guojiashanensis Tong & Erwin 2001 TE01
              |--A. johannisaustriae (Klipstein 1843) (see below for synonymy) TE01
              `--A. wannense (Newell 1935) W63

Ananias johannisaustriae (Klipstein 1843) [=Pleurotomaria johannisaustriae, A. joannisaustriae (l. c.); incl. Worthenia apunctata Kittl 1894] TE01

*Type species of generic name indicated

REFERENCES

[BR05] Bouchet, P., J.-P. Rocroi, J. Frýda, B. Hausdorf, W. Ponder, Á. Valdés & A. Warén. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1-2): 1-397.

[FB04] Frýda, J., & R. B. Blodgett. 2004. New Emsian (Late Early Devonian) gastropods from Limestone Mountain, Medfra B-4 quadrangle, west-central Alaska (Farewell Terrane), and their paleobiogeographic affinities and evolutionary significance. Journal of Paleontology 78 (1): 111-132.

[KC60] Knight, J. B., L. R. Cox, A. M. Keen, R. L. Batten, E. L. Yochelson & R. Robertson. 1960. Gastropoda: systematic descriptions. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt I. Mollusca 1: Mollusca—General Features, Scaphopoda, Amphineura, Monoplacophora, Gastropoda—General Features, Archaeogastropoda and some (mainly Paleozoic) Caenogastropoda and Opisthobranchia, pp. I169-I331. Geological Society of America, and University of Kansas Press.

[PE02] Pan, H.-Z., & D. H. Erwin. 2002. Gastropods from the Permian of Guangxi and Yunnan Provinces, south China. Journal of Paleontology 76 (Memoir 56): 1-49.

[TE01] Tong, J., & D. H. Erwin. 2001. Triassic gastropods of the southern Qinling Mountains, China. Smithsonian Contributions to Paleobiology 92: 1-47.

[W63] Waterhouse, J. B. 1963. Permian gastropds of New Zealand. Part 3—Pleurotomariacea (concluded). New Zealand Journal of Geology and Geophysics 6 (4): 587-622.

[YN83] Yu W. & Ning H. 1983. Middle Ordovician gastropods from Ruoqiang, Xinjiang. Acta Palaeontologica Sinica 22 (2): 199-203.