Pleurotomarioidea

Holotype of Perotrochus charlestonensis, from Askew (1988).


Belongs within: Vetigastropoda.
Contains: Platyschismatinae, Turbonellininae.

The Pleurotomarioidea, slit shells, are a group of marine gastropods represented in the modern fauna only by the family Pleurotomariidae. Members of this group have a shell with a labral slit, followed by a corresponding selenizone (Bouchet et al. 2005).

<==Pleurotomarioidea
    |  i. s.: Mellarium Waterhouse 1960 W63c
    |           `--M. mutchi W63a
    |--Catantostoma Sandberger 1842 [Catantostomatidae, Catantostomatinae] BR05
    |    `--*C. clathratum Sandberger 1842 KC60
    |--Kittlidiscus Haas 1953 BR05 [=Schizodiscus Kittl 1891 non Hall & Clarke 1888 KC60; Kittlidiscidae]
    |    `--*K. bronni (Klipstein 1845) (see below for synonymy) KC60
    |--Rhaphischisma Knight 1936 BR05 [=Rotellina de Koninck 1881 non Agassiz 1846 KC60; Rhaphischismatidae]
    |    `--*R. planorbiformis (deKoninck 1881) [=*Rotellina planorbiformis] KC60
    |--Polytremariide [Polytremariinae] BR05
    |    |--Polytremaria d’Orbigny 1850 BR05
    |    |    `--*P. catenata (deKoninck 1843) [=Pleurotomaria catenata] KC60
    |    `--Plocostoma Gemmellaro 1889 KC60
    |         `--*P. neumayri (Gemmellaro 1889) [=Pleurotomaria (*Plocostoma) neumayri] KC60
    |--Zygitidae BR05
    |    |--Eirlysia Batten 1956 W63, TE01
    |    |    |--*E. exquisita Batten 1956 KC60
    |    |    `--E. reticulata Batten 1958 W63b
    |    `--Zygites Kittl 1891 BR05
    |         |--*Z. delphinula (Laube 1868) TE01 (see below for synonymy)
    |         `--Z. laevigatus Tong & Erwin 2001 TE01
    |--Trochotomidae [Ditremariinae] BR05
    |    |--Valfinia Cox 1958 [=Didymodon Fischer 1885 nec Hedw. 1801 (ICBN) nec Blake 1863] KC60
    |    |    |--*V. quinquecincta (Zieten 1832) [=Trochus quinquecinctus, Didymodon quinquecinctus] KC60
    |    |    `--V. thurmanni KC60
    |    `--Trochotoma Eudes-Deslongchamps 1842 BR05 (see below for synonymy)
    |         |--T. (Trochotoma) KC60
    |         |    |--*T. (T.) conuloides Eudes-Deslongchamps 1843 KC60
    |         |    `--T. (T.) bicarinata KC60
    |         `--T. (Discotoma Haber 1934) TE01
    |              |--T. (*D.) amata (d’Orbigny 1850) [=Ditremaria amata] TE01
    |              |--T. (D.) gansuensis Tong & Erwin 2001 TE01
    |              `--‘Discotoma’ planoconvexa TE01
    |--Portlockiellidae BR05
    |    |--Portlockiella Knight 1945 BR05 [incl. Portlockia deKoninck 1881 non M’Coy 1846 KC60]
    |    |    `--*P. kentuckyensis Knight 1945 KC60
    |    |--Agniesella Cossmann 1909 [=Pleuroderma Perner 1907 non Tschudi 1837] KC60
    |    |    `--*A. aratula (Perner 1907) [=Pleurotomaria (*Pleuroderma) aratula] KC60
    |    |--Latischisma Thomas 1940 W63c, KC60
    |    |    `--L. globosa Thomas 1940 W63c
    |    |--Tapinotomaria Batten 1956 KC60
    |    |    |--*T. rugosa Batten 1956 KC60
    |    |    `--T. mirabilis Batten 1958 PE02
    |    `--Shansiella Yin 1932 W63c
    |         |--*S. altispiralis Yin 1932 W63c
    |         |--S. carbonaria (Norwood & Pratten 1855) B79
    |         |--S. conica Batten 1958 W63c
    |         `--S. tabulata Batten 1958 W63c
    |--Sinuopeidae W63b
    |    |  i. s.: Lunulazona Sadlick & Nielsen 1963 PE02, KB04
    |    |         Naticasinus Pan & Erwin 2002 PE02, KB04
    |    |           `--*N. sinus Pan & Erwin 2002 KB04
    |    |         Pandospira Waterhouse 1963 W63b
    |    |           |--*P. fimbriae Waterhouse 1963 W63b
    |    |           `--P. nuda Waterhouse 1963 W63b
    |    |         Keeneeia Etheridge 1902 W63b
    |    |           |--*K. platyschismoides Etheridge 1902 W63b
    |    |           |--K. ocula (Sowerby 1838) [=Trochus oculus, Euomphalus oculus, Platyschisma oculus] W63b
    |    |           `--K. trochiforme Fletcher 1958 F71
    |    |--Platyschismatinae BR05
    |    |--Turbonellininae BR05
    |    `--Sinuopeinae BR05
    |         |--Sinuopea Ulrich 1911 BR05
    |         |    `--*S. sweeti (Whitfield 1880) [=Holopea sweeti] KC60
    |         |--Horiostomella Perner 1903 KC60
    |         |    `--*H. otiosa Perner 1903 KC60
    |         `--Sellinema Perner 1903 KC60
    |              `--*S. dive Perner 1903 KC60
    |--Phymatopleuridae W63c
    |    |--Paragoniozona Nelson 1947 KC60
    |    |    `--*P. nodolirata Nelson 1947 KC60
    |    |--Discotropis Yochelson 1956 B79
    |    |    `--D. publicus Yochelson 1956 KC60
    |    |--Borestus Thomas 1940 [incl. Platypleurotomaria Wanner 1942] KC60
    |    |    `--*B. wrighti Thomas 1940 KC60
    |    |--Callitomaria Batten 1956 KC60
    |    |    `--*C. stanislavi Batten 1956 KC60
    |    |--Eymarella Cossmann 1897 [=Echetus Koken 1896 non Krøyer 1864] KC60
    |    |    `--*E. subscalariformis [=Pleurotomaria subscalariformis] KC60
    |    |--Lamellospira Batten 1958 KC60, B79
    |    |    `--*L. conica Batten 1958 KC60
    |    |--Phymatopleura Girty 1939 BR05 [=Orestes Girty 1912 nec Blackinstone & Fryer 1880 nec Redtenbacher 1906 KC60]
    |    |    |--*P. nodosus (Girty 1912) [=*Orestes nodosus] KC60
    |    |    `--P. brazoensis W63c
    |    |--Spirovallum Waterhouse 1963 [Spirovallini] W63c
    |    |    |--*S. liratum Waterhouse 1963 W63c
    |    |    `--S. fasciatum Waterhouse 1963 W63c
    |    |--Euryalox Cossmann 1896 TE01 [=Sagana Koken 1896 non Walker 1855 KC60]
    |    |    |--*E. juvavica (Koken 1894) TE01 [=Pleurotomaria juvavica KC60, *Sagana juvavica KC60]
    |    |    `--E. subcancellata [=Pleurotomaria subcancellata d’Orbigny non Morris 1845] TE01
    |    |--Codinella Kittl 1899 TE01
    |    |    |--*C. generelli (Stoppani 1858) [=Trochus generelli] TE01
    |    |    |--C. leda (Kittl 1894) TE01
    |    |    `--C. mammiformis TE01
    |    |--Glyptotomaria Knight 1945 KC60
    |    |    |--*G. (Glyptotomaria) apiarium Knight 1945 KC60
    |    |    `--G. (Dictyotomaria Knight 1945) KC60
    |    |         `--G. (*D.) scitula (Meek & Worthen 1861) [=Pleurotomaria scitula] KC60
    |    `--Discotomaria Batten 1956 P85
    |         |--*D. basisulcata Batten 1956 KC60
    |         |--D. klobukowskii (Mansuy 1912) P85, B79 (see below for synonymy)
    |         |--D. turritus (Delpey 1942) P85, B79 [=Euomphalus turritus P85, Discotropis turritus B79]
    |         `--D. xingwenensis Pan 1985 P85
    `--Pleurotomariidae [Pleurotomariae, Pleurotomariinae] BR05
         |--Angyomphalus Cossmann 1916 P84
         |    `--*A. radians (deKoninck 1843) [=Euomphalus radians, Trepospira (*Angyomphalus) radians] KC60
         |--Mikadotrochus Lindholm 1927 HPA88
         |    `--*M. beyrichi (Hilger 1877) [=Pleurotomaria beyrichi] KC60
         |--Conotomaria Cox 1959 HPA88
         |    `--*C. mailleana (d’Orbigny 1843) [=Pleurotomaria mailleana] KC60
         |--Entemnotrochus Fischer 1885 HPA88
         |    `--*E. adansonianus (Crosse & Fischer 1861) [=Pleurotomaria adansoniana] KC60
         |--Bathrotomaria Cox 1956 KC60
         |    `--*B. reticulata (Sowerby 1821) [=Trochus reticulatus] KC60
         |--Leptomaria Eudes-Deslongchamps 1864 KC60
         |    `--*L. amoena (Eudes-Deslongchamps 1849) [=Pleurotomaria amoena] KC60
         |--Stuorella Kittl 1891 [Stuorellidae] KC60
         |    `--*S. subconcava (Münster 1841) [=Trochus subconcavus] KC60
         |--Obornella Cox 1959 KC60
         |    `--*O. plicopunctata (Eudes-Deslongchamps 1849) [=Pleurotomaria plicopunctata] KC60
         |--Chelotia Bayle in Fischer 1885 KC60
         |    `--*C. concava (Deshayes 1832) [=Pleurotomaria concava] KC60
         |--Pyrgotrochus Fischer 1885 KC60
         |    |--*P. bitorquatus (Eudes-Deslongchamps 1849) [=Pleurotomaria bitorquata] KC60
         |    `--P. luciensis C60
         |--Pleurotomaria Defrance 1826 (nom. cons.) BR05 [=Pleurotomarium de Blainville 1825 (nom. rej.) KC60]
         |    |--*P. anglica (Sowerby 1818) [=Trochus anglicus] KC60
         |    |--P. elora LYR78
         |    |--P. hokonuiensis RW49
         |    |--P. planicostata Girty 1937 W63c
         |    `--P. timorensis Wanner 1922 W63b
         `--Perotrochus Fischer 1885 A88
              |--*P. quoyana (Fischer & Bernardi 1856) [=Pleurotomaria quoyana] KC60
              |--P. africanus (Tomlin 1948) A88
              |--P. amabilis (Bayer 1963) A88
              |--P. charlestonensis Askew 1988 A88
              |--P. lucaya SW11
              |--P. maureri Harasewych & Askew 1993 BC01
              |--P. midas Bayer 1966 A88
              |--P. pyramus Bayer 1967 A88
              |--P. tangaroana Bouchet & Metivier 1982 A88
              `--P. teremachii (Kuroda 1955) A88

Discotomaria klobukowskii (Mansuy 1912) P85, B79 [=Euomphalus klobukowskii P85, Discotropis klobukowskii B79; incl. E. rectangulus P85]

*Kittlidiscus bronni (Klipstein 1845) [=Pleurotomaria bronni, *Schizodiscus bronni; incl. P. plana Klipstein 1845 non Münster 1844] KC60

Trochotoma Eudes-Deslongchamps 1842 BR05 [incl. Ditremaria d’Orbigny 1843 BR05, Rimulus d’Orbigny 1842 (n. n.) KC60]

*Zygites delphinula (Laube 1868) TE01 [=Pleurotomaria delphinula TE01, Delphinula cancellata Klipstein 1845 non Gray 1828 KC60]

*Type species of generic name indicated

REFERENCES

[A88] Askew, T. M. 1988. A new species of pleurotomariid gastropod from the western Atlantic. Nautilus 102 (3): 89-91.

[B79] Batten, R. L. 1979. Gastropods from Perak, Malaysia. Part 2. The trochids, patellids and neritids. American Museum Novitates 2685: 1-26.

[BR05] Bouchet, P., J.-P. Rocroi, J. Frýda, B. Hausdorf, W. Ponder, Á. Valdés & A. Warén. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1-2): 1-397.

[BC01] Boyko, C. B. & J. R. Cordeiro. 2001. Catalog of Recent type specimens in the Division of Invertebrate Zoology, American Museum of Natural History. V. Mollusca, part 2 (class Gastropoda [exclusive of Opisthobranchia and Pulmonata], with supplements to Gastropoda [Opisthobranchia], and Bivalvia). Bulletin of the American Museum of Natural History 262: 1-170.

[C60] Cox, L. R. 1960. Gastropoda: general characteristics of Gastropoda. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt I. Mollusca 1: Mollusca—General Features, Scaphopoda, Amphineura, Monoplacophora, Gastropoda—General Features, Archaeogastropoda and some (mainly Paleozoic) Caenogastropoda and Opisthobranchia, pp. I84-I169. Geological Society of America, and University of Kansas Press.

[F71] Fletcher, H. O. 1971. Catalogue of type specimens of fossils in the Australian Museum, Sydney. Australian Museum Memoir 13: 1-167.

[HPA88] Harasewych, M. G., S. A. Pomponi & T. M. Askew. 1988. Spongivory in pleurotomariid gastropods. Nautilus 102 (3): 92-98.

[KC60] Knight, J. B., L. R. Cox, A. M. Keen, R. L. Batten, E. L. Yochelson & R. Robertson. 1960. Gastropoda: systematic descriptions. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt I. Mollusca 1: Mollusca—General Features, Scaphopoda, Amphineura, Monoplacophora, Gastropoda—General Features, Archaeogastropoda and some (mainly Paleozoic) Caenogastropoda and Opisthobranchia, pp. I169-I331. Geological Society of America, and University of Kansas Press.

[KB04] Kues, B. S., R. L. Batten, D. H. Erwin & Pan H.-Z. 2004. A Late Permian Chinese gastropod species, possibly larval, in the Middle Pennsylvanian of New Mexico. Journal of Paleontology 78 (2): 420-423.

[LYR78] Linsley, R. M., E. L. Yochelson & D. M. Rohr. 1978. A reinterpretation of the mode of life of some Paleozoic frilled gastropods. Lethaia 11: 105-112.

[PE02] Pan, H.-Z., & D. H. Erwin. 2002. Gastropods from the Permian of Guangxi and Yunnan Provinces, south China. Journal of Paleontology 76 (Memoir 56): 1-49.

[P85] Pan Y.-T. 1985. Fossil Gastropoda from the Permian of Sichuan, Guizhou, Yunnan and Guangxi. Acta Palaeontologica Sinica 24 (1): 29-37.

[P84] Peel, J. S. 1984. Autecology and systematics of a new Silurian anomphalid gastropod from western North Greenland. Rapp. Grønlands geol. Unders. 121: 77-87.

[RW49] Rout, M. V., & R. W. Willett. 1949. The geology of the Wairaki survey district, Southland. Transactions and Proceedings of the Royal Society of New Zealand 77 (2): 291-305.

[SW11] Smith, S. A., N. G. Wilson, F. E. Goetz, C. Feehery, S. C. S. Andrade, G. W. Rouse, G. Giribet & C. W. Dunn. 2011. Resolving the evolutionary relationships of molluscs with phylogenomic tools. Nature 480: 364-367.

[TE01] Tong, J., & D. H. Erwin. 2001. Triassic gastropods of the southern Qinling Mountains, China. Smithsonian Contributions to Paleobiology 92: 1-47.

[W63a] Waterhouse, J. B. 1963a. Permian gastropods of New Zealand. Part 1—Bellerophontacea and Euomphalacea. New Zealand Journal of Geology and Geophysics 6 (1): 88-112.

[W63b] Waterhouse, J. B. 1963b. Permian gastropods of New Zealand. Part 2—Pleurotomariacea (in part). New Zealand Journal of Geology and Geophysics 6 (2): 115-154.

[W63c] Waterhouse, J. B. 1963c. Permian gastropds of New Zealand. Part 3—Pleurotomariacea (concluded). New Zealand Journal of Geology and Geophysics 6 (4): 587-622.

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