Homalopoma sanguineum, copyright H. Zell.

Belongs within: Vetigastropoda.

The Colloniinae are a group of small gastropods characterised by a periumbilical cord (Monari et al. 1996).

Characters (from Monari et al. 1996): Phaneromphalous base with periumbilical cord, carina or ridge formed by small enlarged part of otherwise simple or slightly thickened peristome. Angulation on protoconch whorl and/or opisthocline lineation on teleoconch rarely appear. Operculum (where known) calcareous.

<==Colloniinae [Colloniini, Homalopomatinae, Homalopomatini] BR05
    |--Bakonyia Szabó 1981 MCS96
    |--Coanollonia Woodring 1928 KC60
    |    `--*C. ambla Woodring 1928 KC60
    |--Cyniscella Cossmann 1888 KC60
    |    `--*C. cornupastoris (Lamarck 1804) [=Cyclostoma cornupastoris] KC60
    |--Gelasinostoma Gardner 1947 KC60
    |    `--*G. elegantulum (Dall 1892) [=Collonia elegantula] KC60
    |--Otomphalus Cossmann 1902 KC60
    |    `--*O. dumasi Cossmann 1902 KC60
    |--Anadema Adams & Adams 1854 KC60
    |    `--*A. caelata (Adams 1854) [=Omphalius caelata] KC60
    |--Leptocollonia Powell 1951 [incl. Cynisca Adams & Adams 1854 non Gray 1844] KC60
    |    `--*L. thielei Powell 1951 KC60
    |--Rangimata Marwick 1928 KC60
    |    `--*R. pervia Marwick 1928 KC60
    |--Tipua Marwick 1943 KC60
    |    `--*T. tricincta (Marshall 1919) [=Submargarita tricincta] KC60
    |--Vexinia Cossmann 1918 KC60
    |    `--*V. crassa (Baudon 1853) [=Delphinula crassa] KC60
    |--Otollonia Woodring 1928 KC60
    |    |--*O. siderea (Guppy 1896) [=Liotia siderea] KC60
    |    `--O. fricki (Crosse 1865) [=Arene (Otollonia) fricki] PP78
    |--‘Bothropoma’ Thiele 1924 non Wagner 1908 [Bothropomatinae] BR05
    |    |--*B. isseli Thiele 1924 BR17
    |    `--B. rubrostriatum Rolán, Rubio & Fernández-Garcés 1997 BC01
    |--Cantrainea Jeffreys 1883 [=Cantraineia Fischer 1885] KC60
    |    |--*C. peloritanum (Cantraine 1835) [=Turbo peloritanus, Homalopoma (*Cantrainea) peloritanum] KC60
    |    `--C. panamense (Dall 1908) H84
    |--Cirsochilus Cossmann 1888 KC60
    |    |--*C. (Cirsochilus) striatum (Lamarck 1804) [=Delphinula striata] KC60
    |    `--C. (Claibornia Palmer 1937) KC60
    |         `--C. (*C.) lineatum (Lea 1933) [=Turbo lineata] KC60
    |--Charisma Hedley 1915 KC60
    |    |--*C. (Charisma) compacta Hedley 1915 KC60
    |    |--C. (Cavostella Laseron 1954) KC60
    |    |    `--C. (*C.) radians Laseron 1954 KC60
    |    `--C. (Cavotera Laseron 1954) KC60
    |         `--C. (*C.) simplex Laseron 1954 KC60
    |--Argalista Iredale 1915 KC60
    |    |--*A. fluctuata (Hutton 1883) P61 (see below for synonymy)
    |    |--A. crassicostata (Murdoch 1905) P61
    |    |--A. micans Powell 1931 P61
    |    |--A. nana Finlay 1930 P61
    |    |--A. rotella Powell 1937 P61
    |    |--A. umbilicata Powell 1926 P61
    |    `--A. variecostata Powell 1937 P61
    |--Collonia Gray 1850 BR05
    |    |--*C. (Collonia) marginata (Lamarck 1804) KC60 [=Delphinula marginata KC60, Turbo marginatus C64]
    |    |--C. (Bonnetella Cossmann 1908) [incl. Bonnetia Cossmann 1907 non Robineau-Desvoidy 1830] KC60
    |    |    `--C. (*B.) planispira (Cossmann 1907) [=Bonnetia planispira] KC60
    |    |--C. (Circulopsis Cossmann 1902) KC60
    |    |    `--C. (*C.) megalomphalus Cossmann 1902 KC60
    |    |--C. (Heniastoma Cossmann 1918) KC60
    |    |    `--C. (*H.) flammulata Cossmann 1888 KC60
    |    `--C. (Parvirota Cossmann 1902) KC60
    |         `--C. (*P.) rotatoria (Deshayes 1863) [=Turbo rotatorius] KC60
    |--Leptothyra Pease 1869 KC60
    |    |--*L. costata Pease 1869 KC60
    |    |--L. bacula Carpenter 1864 O27
    |    |--L. carpenteri Pilsbry 1888 O27
    |    |--L. grippi Dall 1911 O27
    |    |--L. juanensis Dall 1919 O27
    |    |--L. laeta (Montrouzier 1863) [=Turbo laetus] H09
    |    |--L. lurida Dall 1885 O27
    |    |--L. morini Viader 1951 L96
    |    |--L. nanina (Souverbie 1864) [=Turbo naninus] H09
    |    `--L. paucicostata Dall 1871 O27
    |         |--L. p. paucicostata O27
    |         `--L. p. fenestrata Bartsch 1919 O27
    `--Homalopoma Carpenter 1864 BR05 [incl. Leptonyx Carpenter 1864 non Swainson 1833 KC60]
         |  i. s.: ‘Leptonyx’ bacula Carpenter 1864 C64
         |         H. carpenteri (Pilsbry 1888) H84
         |--*H. (Homalopoma) sanguineum (Linnaeus 1758) KC60 (see below for synonymy)
         |--H. (Boutillieria Cossmann 1888) [incl. Otaulax Cossmann 1888] KC60
         |    `--H. (*B.) eugeni (Deshayes 1863) [=Turbo eugeni] KC60
         |--H. (Collonista Iredale 1918) KC60
         |    `--H. (*C.) pictum (Pease 1868) KC60 [=Collonia picta KC60, *Collonista picta F27]
         |--H. (Eutinochilus Cossmann 1918) [=Homalochilus Cossmann 1892 non Fischer 1856] KC60
         |    `--H. (*E.) miliaris (Cossmann 1892) [=Collonia miliaris] KC60
         |--H. (Leptothyropsis Woodring 1928) KC60
         |    `--H. (*L.) philipianum (Dall 1889) [=Leptothyra philipiana] KC60
         `--H. (Phanerolepida Dall 1907) KC60
              `--H. (*P.) transenna (Watson 1879) [=Turbo transenna] KC60

*Argalista fluctuata (Hutton 1883) P61 [=Cyclostrema fluctuatum P61, Homalopoma (*Argalista) fluctuatum KC60; incl. Argalista fluctuata var. immaculata (Suter 1908) F27]

*Homalopoma (Homalopoma) sanguineum (Linnaeus 1758) KC60 [=Turbo sanguineus KC60, Leptonyx sanguineus C64; incl. T. coccineus C64]

*Type species of generic name indicated


[BR05] Bouchet, P., & J.-P. Rocroi. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1–2): 1–397.

[BR17] Bouchet, P., J.-P. Rocroi, B. Hausdorf, A. Kaim, Y. Kano, A. Nützel, P. Parkhaev, M. Schrödl & E. E. Strong. 2017. Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia 61 (1–2): 1–526.

[BC01] Boyko, C. B., & J. R. Cordeiro. 2001. Catalog of Recent type specimens in the Division of Invertebrate Zoology, American Museum of Natural History. V. Mollusca, part 2 (class Gastropoda [exclusive of Opisthobranchia and Pulmonata], with supplements to Gastropoda [Opisthobranchia], and Bivalvia). Bulletin of the American Museum of Natural History 262: 1–170.

[C64] Carpenter, P. P. 1864. Supplementary report on the present state of our knowledge with regard to the Mollusca of the west coast of North America. Report of the British Association for the Advancement of Science 33: 517–686.

[F27] Finlay, H. J. 1927. A further commentary on New Zealand molluscan systematics. Transactions and Proceedings of the New Zealand Institute 57: 320–485.

[H09] Hedley, C. 1909. The Marine Fauna of Queensland: Address by the President of Section D. Australasian Association for the Advancement of Science: Brisbane.

[H84] Hickman, C. S. 1984. Implications of radular tooth-row functional integration for archaeogastropod systematics. Malacologia 25 (1): 143–160.

[KC60] Knight, J. B., L. R. Cox, A. M. Keen, R. L. Batten, E. L. Yochelson & R. Robertson. 1960. Gastropoda: systematic descriptions. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt I. Mollusca 1: Mollusca—General Features, Scaphopoda, Amphineura, Monoplacophora, Gastropoda—General Features, Archaeogastropoda and some (mainly Paleozoic) Caenogastropoda and Opisthobranchia pp. I169–I331. Geological Society of America, and University of Kansas Press.

[L96] Loch, I. 1996. Notice: R. Viader collection. Molluscan Research 17: 110.

[MCS96] Monari, S., M. A. Conti & J. Szabó. 1996. Evolutionary systematics of Jurassic Trochoidea: the family Colloniidae and the subfamily Proconulidae. In: Taylor, J. D. (ed.) Origin and Evolutionary Radiation of the Mollusca pp. 199–204. Oxford University Press: Oxford.

[O27] Oldroyd, I. S. 1927. The Marine Shells of the West Coast of North America vol. 2 pt 3. Stanford University Press: Stanford University (California).

[PP78] Poorman, F. L., & L. H. Poorman. 1978. Additional molluscan records from Bahía de Los Angeles, Baja California Norte. Veliger 20 (4): 369–374.

[P61] Powell, A. W. B. 1961. Shells of New Zealand: An illustrated handbook 4th ed. Whitcombe and Tombs Limited: Christchurch.

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