Ananias alticonica, from Pan & Erwin (2002).

Belongs within: Vetigastropoda.

The Eotomariinae are a group of gastropods known from the Middle Ordovician to Permian.

Characters (from Knight et al. 1960): Labral slit moderate to deep, umbilicus narrow or absent; ornament collabral and spiral elements, collabral dominant.

    |--Deseretospira Gordon & Yochelson 1987 [Deseretospirides, Deseretospirini] BR05
    |--Ptychomphalina Bayle 1885 [Ptychomphalininae, Ptychomphalinini] BR05
    |--Glabrocingulum Thomas 1940 [Glabrocingulides, Glabrocingulini] BR05
    |    `--*G. beggi Thomas 1940 KC60
    `--Eotomariini [Eotomariides, Liospirinae] BR05
         |--Eotomaria Ulrich & Scofield 1897 BR05 [incl. Spiroraphe Perner 1907 KC60]
         |    `--*E. canalifera Ulrich in Ulrich & Scofield 1897 KC60
         |--Pseudobaylea Dickins 1963 W63
         |    `--*P. freneyensis Dickins 1963 [=Mourlonia (Pseudobaylea) freneyensis] W63
         |--Arastra Stoyanow 1948 KC60
         |    `--*A. torquata Stoyanow 1948 KC60
         |--Trepospira Ulrich & Scofield 1897 [incl. Kansana Tasch 1953] KC60
         |    `--*T. sphaerulata (Conrad 1842) [=Pleurotomaria sphaerulata] KC60
         |--Bembexia Oehlert & Oehlert 1888 FB04
         |    `--*B. larteti (Munier-Chalmas 1876) [=Pleurotomaria larteti] KC60
         |--Liospira Ulrich & Scofield 1897 BR05 [incl. Eocryptaenia Koken 1925 KC60]
         |    |--*L. micula (Hall 1862) [=Pleurotomaria micula] KC60
         |    `--L. angustata Ulrich & Scofield 1897 YN83
         |--Clathrospira Ulrich & Scofield 1897 KC60
         |    |--*C. subconica (Hall 1847) [=Pleurotomaria subconica] KC60
         |    |--C. angulata Yu & Ning 1983 YN83
         |    `--C. conica YN83
         `--Ananias Knight 1945 W63, PE02
              |--*A. welleri (Newell 1935) PE02 [=Phanerotrema welleri PE02, Glabrocingulum (*Ananias) welleri KC60]
              |--A. alticonica (Wang in Wang & Xi 1980) [=Guizhouspira alticonica] PE02
              |--A. campbelli (Waterhouse 1963) [=Glabrocingulum (Ananias) campbelli] W63
              |--A. gibber Chronic 1952 W63
              |--A. guojiashanensis Tong & Erwin 2001 TE01
              |--A. johannisaustriae (Klipstein 1843) (see below for synonymy) TE01
              `--A. wannense (Newell 1935) W63

Ananias johannisaustriae (Klipstein 1843) [=Pleurotomaria johannisaustriae, A. joannisaustriae (l. c.); incl. Worthenia apunctata Kittl 1894] TE01

*Type species of generic name indicated


[BR05] Bouchet, P., J.-P. Rocroi, J. Frýda, B. Hausdorf, W. Ponder, Á. Valdés & A. Warén. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1-2): 1-397.

[FB04] Frýda, J., & R. B. Blodgett. 2004. New Emsian (Late Early Devonian) gastropods from Limestone Mountain, Medfra B-4 quadrangle, west-central Alaska (Farewell Terrane), and their paleobiogeographic affinities and evolutionary significance. Journal of Paleontology 78 (1): 111-132.

[KC60] Knight, J. B., L. R. Cox, A. M. Keen, R. L. Batten, E. L. Yochelson & R. Robertson. 1960. Gastropoda: systematic descriptions. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt I. Mollusca 1: Mollusca—General Features, Scaphopoda, Amphineura, Monoplacophora, Gastropoda—General Features, Archaeogastropoda and some (mainly Paleozoic) Caenogastropoda and Opisthobranchia, pp. I169-I331. Geological Society of America, and University of Kansas Press.

[PE02] Pan, H.-Z., & D. H. Erwin. 2002. Gastropods from the Permian of Guangxi and Yunnan Provinces, south China. Journal of Paleontology 76 (Memoir 56): 1-49.

[TE01] Tong, J., & D. H. Erwin. 2001. Triassic gastropods of the southern Qinling Mountains, China. Smithsonian Contributions to Paleobiology 92: 1-47.

[W63] Waterhouse, J. B. 1963. Permian gastropds of New Zealand. Part 3—Pleurotomariacea (concluded). New Zealand Journal of Geology and Geophysics 6 (4): 587-622.

[YN83] Yu W. & Ning H. 1983. Middle Ordovician gastropods from Ruoqiang, Xinjiang. Acta Palaeontologica Sinica 22 (2): 199-203.

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