Ichneumonidae

Male Anomalon, copyright Rick Shackleton.


Belongs within: Vespina.
Contains: Orthocentrus, Atrophini, Glyptini, Euceros, Phygadeuontini, Hemigasterini, Cryptini, Scolobatini, Temelucha, Ophion, Enicospilus, Tersilochinae, Mesochorinae, Oxytorinae, Metopiinae, Labenini, Groteini, Brachycyrtini, Phytodietini, Gravenhorstiini, Campoplegini, Diplazontinae, Neoxoridini, Rhyssini, Ephialtini, Pimplini, Ichneumonini, Phaeogenini, Platylabini, Oedicephalini, Heresiarchini.

The Ichneumonidae, ichneumon wasps, are a diverse group of often large parasitoid wasps, often with a strongly exserted ovipositor in females. The majority of species attack larvae of holometabolous insects, though some species feed on spiders or their egg-sacs. Major subgroups of the Ichneumonidae include the Ichneumoninae, characterised by a relatively shortly projecting ovipositor (projecting by less than half the length of the hind tibia) protected by a rigid sheath (Gauld 1984).

Characters (from Gauld 1984): Hind wing with radio-medial crossvein arising from Rs distal to its junction with R1. Forewing with vein 2m-cu usually present; first discal cell and first submarginal cell conjoined, not separated by vein.

<==Ichneumonidae
    |--+--Tryphoninae G84
    |  |    |  i. s.: Cteniscus A71
    |  |    |         Anisoctenion alacer A71
    |  |    |         Erromenus Holmgren 1855 HP-W05
    |  |    |           `--E. zonator MS01
    |  |    |         Ctenochira Förster 1855 HP-W05
    |  |    |         Eridolius Förster 1868 HP-W05
    |  |    |         Polyblastus Hartig 1837 HP-W05
    |  |    |           |--P. cothurnatus A71
    |  |    |           `--P. elegans Szépligeti in Mocsáry & Szépligeti 1901 MS01
    |  |    |--Phytodietini PK17
    |  |    |--Sphinctus [Sphinctini] G84
    |  |    |--Idiogrammatini G84
    |  |    |--Ankylophon Gauld 1984 [Ankylophonini] G84
    |  |    |    `--*A. obligatus Gauld 1984 G84
    |  |    |--Exenterus A71 [Exenterini G84]
    |  |    |    |--E. abruptorius A71
    |  |    |    |--E. adspersus A71
    |  |    |    `--E. amictorius A71
    |  |    |--Tryphonini G84
    |  |    |    |--Grypocentrus G84
    |  |    |    `--Tryphon MS01
    |  |    |         |--T. confinis MS01
    |  |    |         |--T. consobrinus MS01
    |  |    |         |--T. incestus A71
    |  |    |         |--T. semirufus A71
    |  |    |         |--T. trochanteratus MS01
    |  |    |         `--T. vulgaris MS01
    |  |    `--Oedemopsini [Eclytini, Thymaridini] G84
    |  |         |--Debophanes Gauld 1984 G84
    |  |         |    `--*D. areolatus Gauld 1984 G84
    |  |         |--Thymaris Foerster 1869 [=Thymarus Thomson 1883] G84
    |  |         |    `--‘Mesoleptus’ tener [incl. *Thymaris pulchricornis] G84
    |  |         `--Oedemopsis Tschek 1869 (see below for synonymy) G84
    |  |              |--‘Tryphon’ scabriculus [=*Hybophanes scabriculus; incl. *Oedemopsis rogenhoferi] G84
    |  |              |--O. davisi [incl. Tryphon nasutus, *Campothreptus nasutus, *Zarhynchus nasutus] G84
    |  |              `--O. hobartensis G84
    |  `--+--Anomaloninae [Anomalinae] G84
    |     |    |  i. s.: Pseudanomalon G84
    |     |    |--Gravenhorstiini PK17
    |     |    `--Anomalon Panzer 1804 (see below for synonymy) G84
    |     |         |--‘Ophion’ foliator (see below for synonymy) G84
    |     |         |--A. affine Lucas 1849 E12
    |     |         |--*Microcremastus’ amseli G84
    |     |         |--A. australense G84
    |     |         |--*Ophiononeura’ flavomaculata G84
    |     |         |--‘Nototrachis’ kozlovi [incl. *Pseudonototrachys pallidus] G84
    |     |         |--A. morleyi G84
    |     |         |--*Trachyopterus’ primus G84
    |     |         `--*Stictophion’ rufipes [incl. *Erythrophion ferrugineus] G84
    |     `--Campopleginae [Porizontinae] G84
    |          |  i. s.: Rhorus Förster 1868 HP-W05
    |          |         Hypamblys Förster 1868 HP-W05
    |          |           `--H. albopictus (Gravenhorst 1829) HP-W05
    |          |         Bathyplectes G84
    |          |         Rhimphoctona G84
    |          |         Lathrostizus G84
    |          |         Spudasticta G84
    |          |--Nonnini G84
    |          |--Helwigiini G84
    |          `--Campoplegini PK17
    `--+--+--Diplazontinae PK17
       |  `--Pimplinae [Ephialtinae] IR05
       |       |  i. s.: Pseudorhyssa alpestris G84, A71
       |       |         Dolichomitus G84
       |       |--Neoxoridini IK05
       |       |--Rhyssini IK05
       |       |--Ephialtini IR05
       |       |--Pimplini PK17
       |       `--Polysphinctini G84
       |            |--Hymenoepimecis argyraphaga G84, GE05
       |            |--Dreisbachia Townes 1962 [=Laufeia Tosquinet 1903 non Simon 1889] G84
       |            |    `--*D. mira [=*Laufeia mira] G84
       |            `--Eriostethus Morley 1914 [incl. Millironia Baltazar 1964] G84
       |                 |--*E. pulcherrimus G84
       |                 |--E. carinatus G84
       |                 |--E. perkinsi G84
       |                 `--E. trifasciatus [=*Millironia trifasciata] G84
       `--+--Buathra laborator PK17
          `--Ichneumoninae NI11
               |  i. s.: Amblyteles A71
               |           |--A. culpatorius MS01
               |           |--A. glaucatorius MS01
               |           |--A. infractorius MS01
               |           `--A. subfuscus A71
               |         Barichneumon A71
               |--Ichneumonini PK17
               |--Phaeogenini G84
               |--Platylabini NI11
               |--Trogini G84
               |--Ceratojoppini G84
               |--Eurylabus [Eurylabini] G84
               |--Heterischnus [Heterischnini] G84
               |--Zimmeriini G84
               |--Oedicephalini NI11
               |--Heresiarchini NI11
               |--Magwengiella Heinrich 1938 [Ctenocalini] G84
               |    `--*M. obtusa G84
               |--Ischnojoppa Kriechbaumer 1898 G84 [incl. Bodargus Cameron 1902 G84; Ishnojoppini NI11]
               |    `--I. luteator (Fabricius 1798) NI11 (see below for synonymy)
               |--Compsophorus [Compsophorini] NI11
               |    `--C. crassispina (Cameron 1901) (see below for synonymy) NI11
               |--Goedartiini NI11
               |    |--Goedartia alboguttata P93
               |    `--Maraces flavobalteata Cameron 1902 (see below for synonymy) NI11
               |--Joppocryptini NI11
               |    |--Allonotus rufus Cameron 1907 NI11
               |    `--Eccoptosage NI11
               |         |--E. cincticornis (Cameron 1902) (see below for synonymy) NI11
               |         `--E. quadridentata (Cameron 1905) [=Druscia quadridentata, Druseia quadridentata] NI11
               |--Protichneumonini G84
               |    |--Catadelphus arrogator G84, P93
               |    |--Yeppoona Gauld 1984 G84
               |    |    `--*Y. grandis Gauld 1984 G84
               |    `--Protichneumon P93
               |         |--P. fusorius P93
               |         `--P. pisorius P93
               |--Listrodromini NI11
               |    |--Neotypus G84
               |    |--Anisobas kankoensis (Uchida 1932) [=Amblyteles kankoensis, Validentia kankoensis] NI11
               |    `--Listrodromus Wesmael 1845 [incl. Sogna Cameron 1907] G84
               |         |--*L. nycthemerus [=Ichneumon nycthemerus] G84
               |         `--*Sogna’ crassipes G84
               `--Alomyini NI11
                    |--Alomya G84
                    |--Megalomya G84
                    |--Lusiuxta floreni Diller 2006 NI11
                    |--Diadromus Wesmael 1845 NI11, G84 [incl. Thyraeella Holmgren 1890 G84]
                    |    |--*D. troglodytes [=Ichneumon troglodytes] G84
                    |    `--D. collaris (Gravenhorst 1829) NI11 [=Ishnus collaris NI11, *Thyraeella collaris G84]
                    `--Tycherus NI11
                         |--T. cacumencultor Diller 2006 NI11
                         `--T. sabahensis Diller 2006 NI11

Ichneumonidae incertae sedis:
  Tanychora petiolata R02
  Tanychorella R02
  Rhyssella A71
    |--R. approximator I92
    `--R. curvipes A71
  Oocenteter A71
  Exeristes G84
    |--E. comstockii A71
    `--E. ruficollis G84
  Glabrodorsum stokesii H91
  Sinicorussus luzhongensis GE05
  Ariostonia Girault 1933 G33
    `--*A. fusciapex Girault 1933 G33
  Photocryptus apicalis BM76
  Acroricnus BM76
  Perithous BM76
    |--P. divinator BM76
    `--P. mediator BM76
         |--P. m. mediator BM76
         `--P. m. neomexicanus BM76
  Hoplocryptus BM76
  Caenocryptus BM76
  Lochetica BM76
  Neorhachodes BM76 [Neorhacodinae G84]
    `--N. enslini BM76
  Erigorgus perspicillator MS01
  Nepiesta nigra Szépligeti in Mocsáry & Szépligeti 1901 MS01
  Apilasta nigrifemur Szépligeti in Mocsáry & Szépligeti 1901 MS01
  Trachyderma scabra MS01
  Promethes MS01
    |--P. pulchellus MS01
    `--P. sulcator MS01
  Liopsis festivus MS01
  Zootrephes MS01
    |--Z. cingulatus MS01
    `--Z. lateralis MS01
  Aniarophron MS01
    |--A. niger Szépligeti in Mocsáry & Szépligeti 1901 MS01
    |--A. nigricornis MS01
    `--A. ornatus MS01
  Phthorima rossica Szépligeti in Mocsáry & Szépligeti 1901 MS01
  Microcryptus MS01
    |--M. basizonius MS01
    `--M. xanthostigma Szépligeti in Mocsáry & Szépligeti 1901 MS01
  Acanthocryptus MS01
    |--A. flagitator [incl. A. pumilio] MS01
    `--A. quadrispinosus MS01
  Gnathocryptus vagabundus MS01
  Stenareus transfuga MS01
  Spilocryptus MS01
  Atractodes MS01
    |--A. insignis MS01
    `--A. proprius MS01
  Exyston cinctellus MS01
  Diaborus crassiceps Szépligeti in Mocsáry & Szépligeti 1901 MS01
  Mesoleptus MS01
    |--M. flavovariegatus Lucas 1849 E12
    |--M. melanocephalus MS01
    `--M. (Zemiotes) vulnerator MS01
  Cephaloglypta laricis W87
  Alcima orbitalis P93
  Amblyjoppa P93
    |--A. fuscipennis P93
    `--A. proteus P93
  Banchus P93
    |--B. falcatorius P93
    |--B. palpalis P93
    `--B. pictus P93 [=Ichneumon pictus L02]
  Callajoppa P93
    |--C. cirrogaster P93
    `--C. exaltatoria P93
  Diphyus P93
    |--D. longigena P93
    |--D. monitorius P93
    `--D. palliatorius P93
  Lymantrichneumon disparis P93
  Hybrizon [Paxylommatidae, Paxylommatinae] G84
  Orthopelma N88 [Orthopelmatinae G84]
  Stilbopinae G84
  Agriotypus A71 [Agriotypidae, Agriotypinae G84]
    `--A. agrotypus A71
  Adelognathus Holmgren 1855 HP-W05 [Adelognathinae G84]
    |--A. cubiceps Roman 1924 HP-W05
    `--A. difformis Holmgren 1857 HP-W05
  Collyria E12 [Collyriinae G84]
    |--C. armata Lucas 1849 E12
    `--C. erythrogaster Lucas 1849 E12
  Lycorina Holmgren 1859 (see below for synonymy) G84
    |--*L. triangulifera G84
    |--‘Glypta’ acitula [=*Chlorolycorina acitula] G84
    |--L. apicalis [=*Toxophoroides apicalis] G84
    `--L. fici [=L. (*Gonioglyphus) fici] G84
  Xorides [Xoridinae] G84
    `--X. (Cyanoxorides Cameron 1903) [incl. Spiloxorides Cameron 1903] G84
         |--X. (*C.) brookei G84
         |--X. (C.) australiensis G84
         |--X. (C.) crudelis G84
         `--*Spiloxorides’ ruficeps G84
  Phrudinae G84
    |--Brachyscleroma G84
    |--Earobia G84
    |--Astrenis G84
    `--Phrudus Foerster 1869 [incl. Ktenostilpnus Strobl 1901, Vendolius Roman 1914] G84
         `--*P. monilicornis [incl. *Ktenostilpnus aequearticulatus, *Vendolius stilpnius] G84
  Acaenitinae G84
    |  i. s.: Arotes G84
    |--Coleocentrini G84
    `--Yezoceryx [Acaenitini] G84
         |--*Y. scutellaris G84
         `--Y. apicipennis G84
  Orthocentrinae [Plectiscini] G84
    |--Orthocentrus G84
    |--Stenomacrus G84
    |--Neurateles Ratzeburg 1848 [=Neuratelus Foerster 1869] G84
    |    `--*N. papyraceus G84
    `--Plectiscus Gravenhorst 1829 [=Brephoctonus Foerster 1869] G84
         |--*P. impurator [=*Brephoctonus impurator] G84
         `--P. subtilis MS01
  Banchinae SNS07
    |--Atrophini SNS07
    |--Glyptini G84
    `--Banchini G84
         |--Philogalleria Cameron 1912 G84
         |    `--*P. sextuberculata G84
         `--Exetastes G84
              |--E. csikii Szépligeti in Mocsáry & Szépligeti 1901 MS01
              |--E. fornicator MS01
              |--E. medianus MS01
              |--E. minor Szépligeti in Mocsáry & Szépligeti 1901 MS01
              `--E. notatus MS01
  Euceros G84
  Cryptinae (see below for synonymy) D02
    |  i. s.: Itamoplex A71
    |         Dotocryptus bellicosus [incl. D. macrocercus] A71
    |         Sphecophaga D02
    |           |--S. orientalis Donovan 2002 [=S. vesparum israelensis Harris & Read 1999 (n. n.)] D02
    |           `--S. vesparum D02
    |                |--S. v. vesparum D02
    |                `--S. v. burra D02
    |         Nemapodius G84
    |         Oecotelma G84
    |         Pleurogyrus G84
    |         Ethelurgus G84
    |         Rhembobius G84
    |         Aritranis fugitivus G84, P93
    |         Trychosis sibirica Szépligeti in Mocsáry & Szépligeti 1901 G84, MS01
    |         Theroscopus hemipterus G84
    |--Phygadeuontini G84
    |--Hemigasterini G84
    `--Cryptini G84
  Ctenopelmatinae [Megaceriini, Scolobatinae] G84
    |  i. s.: Lathrolestes G84
    |         Opheltes G84
    |--Scolobatini G84
    |--Ctenopelma A71 [Ctenopelmatini G84]
    |--Olethrodontini G84
    |--Perilissus (Spanotectus) MS01 [Perilissini G84]
    |    |--P. (S.) filicornis MS01
    |    `--P. (S.) vernalis MS01
    |--Pionini G84
    |    |--Hodostates G84
    |    `--Austropion Gauld 1984 G84
    |         `--*A. mandibularis Gauld 1984 G84
    |--Euryproctini G84
    |    |--Hadrodactylus barbatus Szépligeti in Mocsáry & Szépligeti 1901 G84, MS01 (see below for synonymy)
    |    |--Euryproctus G84
    |    |--Denticeria Gauld 1984 G84
    |    |    `--*D. cardaleae Gauld 1984 G84
    |    `--Megaceria Szépligeti 1908 G84
    |         |--*M. opheltes G84
    |         |--M. pagana G84
    |         `--M. rufiventris G84
    `--Mesoleius Holmgren 1855 HP-W05 [Mesoleiini G84]
         |  i. s.: M. sternoxanthus (Gravenhorst 1829) HP-W05
         |         M. tenthredinis G84
         `--M. (Scopesis) MS01
              |--M. (S.) bipunctatus Szépligeti in Mocsáry & Szépligeti 1901 MS01
              `--M. (S.) rufolabris MS01
  Cremastinae G84
    |--Temelucha G84
    |--Creagrura G84
    |--Belesica G84
    |--Gahus Gauld 1984 G84
    |    `--*G. siccus Gauld 1984 G84
    |--Dimophora Foerster 1869 [=Dimophorus Thomson 1889; incl. Oligotmema Cushman 1920] G84
    |    |--*D. robusta G84
    |    `--*Oligotmema’ prima G84
    |--Cremastus G84
    |    |--C. geminus MS01
    |    |--C. guttifer MS01
    |    |--C. melanarius Szépligeti in Mocsáry & Szépligeti 1901 MS01
    |    `--C. pungens MS01
    |--Trathala Cameron 1899 (see below for synonymy) G84
    |    |--*T. striata G84
    |    |--T. agnina M83
    |    |--*Epicremastus’ concolor G84
    |    |--T. flavoorbitalis G84
    |    |--*Paurolexis’ flavus G84
    |    `--*Haristaeus’ nigrifrons G84
    `--Pristomerus Curtis 1836 (see below for synonymy) G84
         |--*P. vulnerator [=Ichneumon vulnerator] G84
         |--*Pristocelus’ atriceps Szépligeti 1905 G84
         |--P. atrifemur Girault 1925 G25b
         |--P. austrinus [incl. Porizon agilis, *Pristomeridia agilis] G84
         |--P. bicinctus G84
         |--*Nesanomalon’ dimidiatum G84
         |--P. giraulti G84 (see below for synonymy)
         `--‘Ichneumon’ spinator [incl. Pristomerus appalachianus, *Neopristomerus appalachianus] G84
  Ophioninae [Enicospilini, Ophionini] G84
    |--Ophion G84
    |--Enicospilus G84
    |--Pamophion Gauld 1977 G84
    |    `--*P. sorus Gauld 1977 G84
    |--Xylophion Gauld 1979 G84
    |    |--*X. xylus [=Ophion xylus] G84
    |    `--X. ketus G84
    |--Riekophion Gauld 1977 G84
    |    |--*R. emandibulator [=Allocamptus emandibulator] G84
    |    |--R. bolus G84
    |    `--R. conspicuus G84
    |--Stauropoctonus Brauns 1889 [=Stauropodoctonus Morley 1913; incl. Nipponophion Uchida 1928] G84
    |    |--*S. bombycivorus [=Ophion bombycivorus; incl. *Nipponophion variegatus] G84
    |    |--‘Nipponophion’ chopini Girault 1932 G32
    |    `--S. torresi G84
    |--Dicamptus Szépligeti 1905 G84
    |    |--*D. giganteus G84
    |    |--D. collessi G84
    |    |--D. fuscicornis [incl. D. triangularis] G84
    |    |--D. indicus G84
    |    `--D. uptoni G84
    `--Leptophion Cameron 1901 [incl. Coiloneura Szépligeti 1905, Spilophion Cameron 1905] G84
         |--*L. longiventris [incl. L. chopini, *Coiloneura melanostigma] G84
         |--L. anici G84
         |--L. antennatus G84
         |--L. iochus G84
         |--*Spilophion’ maculipennis G84
         |--L. tetus G84
         |--L. unicalcaratus G84
         `--L. yampus G84
  Tersilochinae G84
  Mesochorinae G84
  Oxytorinae G84
  Metopiinae G84
  Zagryphus nasutus HR11
  Lymeon orbus HR11
  Labeninae G84
    |--Labenini HR11
    |--Groteini G84
    |--Brachycyrtini G84
    `--Poecilocryptini G84
         |--Alaothyris Gauld 1984 G84
         |    `--*A. elongissimus Gauld 1984 G84
         |--Urancyla Gauld 1984 G84
         |    `--*U. fulva Gauld 1984 G84
         `--Poecilocryptus Cameron 1901 [=Poecilopimpla Morley 1914 non Cameron 1903] G84
              |--*P. nigromaculatus [=*Poecilopimpla nigromaculata] G84
              |--P. nigripectus G84
              `--P. stramineus G84

Anomalon Panzer 1804 [=Anomalum Schulz 1906, incl. Erythrophion Cameron 1906, Microcremastus Hedwig 1961, Nototrychys Marshall 1872, Ophiononeura Cameron 1904, Pseudonototrachys Meyer 1930, Stictophion Cameron 1906, Trachynotus Gravenhorst 1829 non Latreille 1829, Trachyopterus Morley 1912; Anomalonini] G84

Compsophorus crassispina (Cameron 1901) [=Magrettia crassispina, Charitojoppa crassispina, Xenojoppa crassispina] NI11

Cryptinae [Gelinae, Hemiteloidae, Pezomachoidae, Pezomachoides, Phygadeuonides, Phygadeuontinae, Phygadeuontoidae, Stilpnoidae] D02

Eccoptosage cincticornis (Cameron 1902) [=Anisobas cincticornis, Acanthojoppa cincticornis; incl. Ac. nigrinerva Cameron 1909, Eccoptosage minor Szepligeti 1910] NI11

Hadrodactylus barbatus Szépligeti in Mocsáry & Szépligeti 1901 G84, MS01 [=Mesoleptus (Hadrodactylus) barbatus MS01]

Ischnojoppa luteator (Fabricius 1798) NI11 [=Ichneumon luteator NI11, Is. luteatrix NI11, Is. luteolator (l. c.) NI11, Joppa luteator NI11, Ic. adspersor Thunberg 1822 NI11, J. lutea Fabricius 1804 NI11, Ic. lutea NI11, *Ischnojoppa lutea G84; incl. Ic. agraensis Cameron 1897 NI11, Exephanes akonis Matsumura 1912 NI11, Atanyjoppa akonis NI11, *Bodargus rufus Cameron 1902 G84, Joppa rufa Brulle 1846 NI11]

Lycorina Holmgren 1859 [incl. Chlorolycorina Cushman 1920, Gonioglyphus Seyrig 1932, Toxophoroides Cresson 1873; Lycorininae] G84

Maraces flavobalteata Cameron 1902 [=M. flavorbalteata, Eradha flavobalteata, M. flavobalteatus] NI11

Oedemopsis Tschek 1869 [=Oedematopsis Morley 1908; incl. Campothreptus Foerster 1869, Hybophanes Foerster 1869, Zarhynchus Ashmead 1900 non Oberholser 1899] G84

‘Ophion’ foliator [=*Nototrychys foliator, *Trachynotus foliator; incl. *Anomalon cruentatus] G84

Pristomerus Curtis 1836 [incl. Neopristomerus Viereck 1912, Nesanomalon Morley 1913, Pristocelus Szépligeti 1905, Pristomeridia Ashmead 1900] G84

Pristomerus giraulti G84 [incl. P. atriceps Girault 1925 non Pristocelus atriceps Szépligeti 1905 G84, G25a]

Trathala Cameron 1899 [incl. Epicremastus Szépligeti 1905, Haristaeus Cameron 1910, Paurolexis Cameron 1907] G84

*Type species of generic name indicated

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Last updated: 23 September 2017.

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