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Ichneumonidae

Male Euceros, photographed by Erick Adams.


Belongs within: Apocrita.
Contains: Atrophini, Glyptini, Orthocentrus, Labenini, Groteini, Brachycyrtini, Gravenhorstiini, Campoplegini, Neoxoridini, Rhyssini, Ephialtini, Theronia, Pimplini, Scolobatini, Phygadeuontini, Hemigasterini, Cryptini, Temelucha, Ophion, Enicospilus, Phytodietini, Ichneumonini, Phaeogenini, Platylabini.

The Ichneumonidae, ichneumon wasps, are a diverse group of often large parasitoid wasps, often with a strongly exserted ovipositor in females. The majority of species attack larvae of holometabolous insects, though some species feed on spiders or their egg-sacs (Gauld 1984).

Characters (from Gauld 1984): Hind wing with radio-medial crossvein arising from Rs distal to its junction with R1. Forewing with vein 2m-cu usually present; first discal cell and first submarginal cell conjoined, not separated by vein.

<==Ichneumonidae
    |--Hybrizon [Paxylommatidae, Paxylommatinae] G84
    |--Orthopelma N88 [Orthopelmatinae G84]
    |--Stilbopinae G84
    |--Neorhacodinae G84
    |--Agriotypus A71 [Agriotypidae, Agriotypinae G84]
    |    `--A. agrotypus A71
    |--Adelognathus Holmgren 1855 [Adelognathinae] HP-W05
    |    |--A. cubiceps Roman 1924 HP-W05
    |    `--A. difformis Holmgren 1857 HP-W05
    |--Collyria E12 [Collyriinae G84]
    |    |--C. armata Lucas 1849 E12
    |    `--C. erythrogaster Lucas 1849 E12
    |--Lycorina Holmgren 1859 (see below for synonymy) G84
    |    |--*L. triangulifera G84
    |    |--‘Glypta’ acitula [=*Chlorolycorina acitula] G84
    |    |--L. apicalis [=*Toxophoroides apicalis] G84
    |    `--L. fici [=L. (*Gonioglyphus) fici] G84
    |--Xorides [Xoridinae] G84
    |    `--X. (Cyanoxorides Cameron 1903) [incl. Spiloxorides Cameron 1903] G84
    |         |--X. (*C.) brookei G84
    |         |--X. (C.) australiensis G84
    |         |--X. (C.) crudelis G84
    |         `--*Spiloxorides’ ruficeps G84
    |--Phrudinae G84
    |    |--Brachyscleroma G84
    |    |--Earobia G84
    |    |--Astrensis G84
    |    `--Phrudus Foerster 1869 [incl. Ktenostilpnus Strobl 1901, Vendolius Roman 1914] G84
    |         `--*P. monilicornis [incl. *Ktenostilpnus aequearticulatus, *Vendolius stilpnius] G84
    |--Acaenitinae G84
    |    |  i. s.: Arotes G84
    |    |--Coleocentrini G84
    |    `--Yezoceryx [Acaenitini] G84
    |         |--*Y. scutellaris G84
    |         `--Y. apicipennis G84
    |--Banchinae SNS07
    |    |--Atrophini SNS07
    |    |--Glyptini G84
    |    `--Banchini G84
    |         |--Exetastes G84
    |         `--Philogalleria Cameron 1912 G84
    |              `--*P. sextuberculata G84
    |--Orthocentrinae [Plectiscini] G84
    |    |--Orthocentrus G84
    |    |--Stenomacrus G84
    |    |--Neurateles Ratzeburg 1848 [=Neuratelus Foerster 1869] G84
    |    |    `--*N. papyraceus G84
    |    `--Plectiscus Gravenhorst 1829 [=Brephoctonus Foerster 1869] G84
    |         `--*P. impurator [=*Brephoctonus impurator] G84
    |--Euceros Gravenhorst 1829 (see below for synonymy) G84
    |    |--E. pruinosus (Gravenhorst 1829) HP-W05 (see below for synonymy)
    |    |--E. annulicornis G84
    |    |--E. croceus G84
    |    |--E. frigidus Cresson 1869 HP-W05
    |    |--E. incisurae G84
    |    |--E. maculicornis G84
    |    |--E. melleus G84
    |    |--E. ruficeps G84
    |    |--*Pseudasthenara’ rufocincta G84
    |    |--E. serricornis [incl. E. egregius, *Tautozelus egregius] G84
    |    `--E. signicornis G84
    |--Labeninae GE05
    |    |--Labenini G84
    |    |--Groteini G84
    |    |--Brachycyrtini G84
    |    `--Poecilocryptini G84
    |         |--Alaothyris Gauld 1984 G84
    |         |    `--*A. elongissimus Gauld 1984 G84
    |         |--Urancyla Gauld 1984 G84
    |         |    `--*U. fulva Gauld 1984 G84
    |         `--Poecilocryptus Cameron 1901 [=Poecilopimpla Morley 1914 non Cameron 1903] G84
    |              |--*P. nigromaculatus [=*Poecilopimpla nigromaculata] G84
    |              |--P. nigripectus G84
    |              `--P. stramineus G84
    |--Anomaloninae [Anomalinae] G84
    |    |  i. s.: Pseudanomalon G84
    |    |--Gravenhorstiini G84
    |    `--Anomalon Panzer 1804 (see below for synonymy) G84
    |         |--‘Ophion’ foliator [=*Nototrychys foliator, *Trachynotus foliator; incl. *Anomalon cruentatus] G84
    |         |--A. affine Lucas 1849 E12
    |         |--*Microcremastus’ amseli G84
    |         |--A. australense G84
    |         |--*Ophiononeura’ flavomaculata G84
    |         |--‘Nototrachis’ kozlovi [incl. *Pseudonototrachys pallidus] G84
    |         |--A. morleyi G84
    |         |--*Trachyopterus’ primus G84
    |         `--*Stictophion’ rufipes [incl. *Erythrophion ferrugineus] G84
    |--Campopleginae [Campoplegini, Macrini, Porizontinae] HP-W05
    |    |  i. s.: Rhorus Förster 1868 HP-W05
    |    |         Hypamblys Förster 1868 HP-W05
    |    |           `--H. albopictus (Gravenhorst 1829) HP-W05
    |    |         Mesoleius Holmgren 1855 [Mesoleiini] HP-W05
    |    |           |--M. sternoxanthus (Gravenhorst 1829) HP-W05
    |    |           `--M. tenthredinis A71
    |    |         Bathyplectes G84
    |    |         Rhimphoctona G84
    |    |         Lathrostizus G84
    |    |         Spudasticta G84
    |    |--Nonnini G84
    |    |--Helwigiini G84
    |    `--Campoplegini G84
    |--Diplazontinae A71
    |    |--Syrphophilus tricinctorius G84
    |    |--Enizemum G84
    |    |--Diplazon Viereck 1914 G84
    |    |    |--*D. laetatorius [=Ichneumon laetatorius; incl. D. generosus, D. varipes] G84
    |    |    `--D. fissorius A71
    |    |--Woldstedtius Carlson 1979 G84
    |    |    |--*W. biguttatus [=Bassus biguttatus] G84
    |    |    |--W. rubellus G84
    |    |    `--W. subditicius G84
    |    `--Syrphoctonus Foerster 1869 (see below for synonymy) G84
    |         |--*S. exsultans [=Bassus exsultans] G84
    |         |--‘Bassus’ elegans [=*Homocidus elegans, *Homotropus elegans] G84
    |         |--S. nigreoauratilis G84
    |         `--S. rubeoauratilis G84
    |--Pimplinae [Ephialtinae] IR05
    |    |  i. s.: Pseudorhyssa alpestris G84, A71
    |    |         Dolichomitus G84
    |    |--Neoxoridini IK05
    |    |--Rhyssini IK05
    |    |--Ephialtini IR05
    |    |--Theronia G84
    |    |--Pimplini G84
    |    `--Polysphinctini G84
    |         |--Hymenoepimecis argyraphaga G84, GE05
    |         |--Dreisbachia Townes 1962 [=Laufeia Tosquinet 1903 non Simon 1889] G84
    |         |    `--*D. mira [=*Laufeia mira] G84
    |         `--Eriostethus Morley 1914 [incl. Millironia Baltazar 1964] G84
    |              |--*E. pulcherrimus G84
    |              |--E. carinatus G84
    |              |--E. perkinsi G84
    |              `--E. trifasciatus [=*Millironia trifasciata] G84
    |--Ctenopelmatinae [Megaceriini, Scolobatinae] HP-W05
    |    |  i. s.: Lathrolestes G84
    |    |         Opheltes G84
    |    |--Scolobatini G84
    |    |--Ctenopelma A71 [Ctenopelmatini G84]
    |    |--Olethrodontini G84
    |    |--Perilissus A71 [Perilissini G84]
    |    |--Pionini G84
    |    |    |--Hodostates G84
    |    |    `--Austropion Gauld 1984 G84
    |    |         `--*A. mandibularis Gauld 1984 G84
    |    `--Euryproctini G84
    |         |--Hadrodactylus G84
    |         |--Euryproctus G84
    |         |--Denticeria Gauld 1984 G84
    |         |    `--*D. cardaleae Gauld 1984 G84
    |         `--Megaceria Szépligeti 1908 G84
    |              |--*M. opheltes G84
    |              |--M. pagana G84
    |              `--M. rufiventris G84
    |--Cryptinae (see below for synonymy) D02
    |    |  i. s.: Itamoplex A71
    |    |         Dotocryptus bellicosus [incl. D. macrocercus] A71
    |    |         Sphecophaga D02
    |    |           |--S. orientalis Donovan 2002 [=S. vesparum israelensis Harris & Read 1999 (n. n.)] D02
    |    |           `--S. vesparum D02
    |    |                |--S. v. vesparum D02
    |    |                `--S. v. burra D02
    |    |         Nemapodius G84
    |    |         Helcostizus G84
    |    |         Oecotelma G84
    |    |         Pleurogyrus G84
    |    |         Ethelurgus G84
    |    |         Rhembobius G84
    |    |         Aritranis G84
    |    |         Trychosis G84
    |    |         Theroscopus hemipterus G84
    |    |--Phygadeuontini G84
    |    |--Hemigasterini G84
    |    `--Cryptini G84
    |--Cremastinae G84
    |    |--Temelucha G84
    |    |--Creagrura G84
    |    |--Belesica G84
    |    |--Gahus Gauld 1984 G84
    |    |    `--*G. siccus Gauld 1984 G84
    |    |--Dimophora Foerster 1869 [=Dimophorus Thomson 1889; incl. Oligotmema Cushman 1920] G84
    |    |    |--*D. robusta G84
    |    |    `--*Oligotmema’ prima G84
    |    |--Trathala Cameron 1899 (see below for synonymy) G84
    |    |    |--*T. striata G84
    |    |    |--T. agnina M83
    |    |    |--*Epicremastus’ concolor G84
    |    |    |--T. flavoorbitalis G84
    |    |    |--*Paurolexis’ flavus G84
    |    |    `--*Haristaeus’ nigrifrons G84
    |    `--Pristomerus Curtis 1836 (see below for synonymy) G84
    |         |--*P. vulnerator [=Ichneumon vulnerator] G84
    |         |--*Pristocelus’ atriceps G84
    |         |--P. atrifemur Girault 1925 G25b
    |         |--P. austrinus [incl. Porizon agilis, *Pristomeridia agilis] G84
    |         |--P. bicinctus G84
    |         |--*Nesanomalon’ dimidiatum G84
    |         |--P. giraulti G84 [incl. P. atriceps Girault 1925 non Pristocelus atriceps G84, G25a]
    |         `--‘Ichneumon’ spinator [incl. Pristomerus appalachianus, *Neopristomerus appalachianus] G84
    |--Mesochorinae G84
    |    |--Lepidura G84
    |    |--Latilumbus G84
    |    |--Oncocotta G84
    |    |--Piestetron G84
    |    |--Rhaibaspis G84
    |    |--Plectochorus Uchida 1933 G84
    |    |    `--*P. iwatensis [=Mesochorus iwatensis] G84
    |    |--Stictopisthus Thomson 1886 G84
    |    |    |--*S. bilineatus [=Mesochorus bilineatus] G84
    |    |    `--S. australiensis G84
    |    |--Mesochorus Gravenhorst 1829 (see below for synonymy) G84
    |    |    |--*M. splendidulus G84
    |    |    |--*Cryptochorus’ obliterator G84
    |    |    |--*Zamesochorus’ orientalis G84
    |    |    |--*Edrisa’ pilicornis G84
    |    |    `--M. pinarae Girault 1932 G32
    |    |--Astiphromma Foerster 1869 (see below for synonymy) G84
    |    |    |--*A. scutellata [=Mesochorus scutellatus] G84
    |    |    |--‘Mesochorus (*Dolichochorus)’ longiceps G84
    |    |    |--*Demophorellus’ mimulus G84
    |    |    |--‘Mesochorus’ nigriceps [=*Mesochorella nigriceps] G84
    |    |    `--*Pseudacoenitus’ transylvannicus G84
    |    `--Cidaphus Foerster 1869 (see below for synonymy) G84
    |         |--*C. thuringiacus G84
    |         |--‘Mesochorus’ alarius [=*Mater alarius, *Ophthalmochorus alarius, *Plesiophthalmus alarius] G84
    |         |--*Tetragonalys’ barbarica G84
    |         |--C. glabrosus G84
    |         `--*Plesiophthalmidea’ paniscoides G84
    |--Ophioninae [Enicospilini, Ophionini] G84
    |    |--Ophion G84
    |    |--Enicospilus G84
    |    |--Pamophion Gauld 1977 G84
    |    |    `--*P. sorus Gauld 1977 G84
    |    |--Xylophion Gauld 1979 G84
    |    |    |--*X. xylus [=Ophion xylus] G84
    |    |    `--X. ketus G84
    |    |--Riekophion Gauld 1977 G84
    |    |    |--*R. emandibulator [=Allocamptus emandibulator] G84
    |    |    |--R. bolus G84
    |    |    `--R. conspicuus G84
    |    |--Stauropoctonus Brauns 1889 [=Stauropodoctonus Morley 1913; incl. Nipponophion Uchida 1928] G84
    |    |    |--*S. bombycivorus [=Ophion bombycivorus; incl. *Nipponophion variegatus] G84
    |    |    |--‘Nipponophion’ chopini Girault 1932 G32
    |    |    `--S. torresi G84
    |    |--Dicamptus Szépligeti 1905 G84
    |    |    |--*D. giganteus G84
    |    |    |--D. collessi G84
    |    |    |--D. fuscicornis [incl. D. triangularis] G84
    |    |    |--D. indicus G84
    |    |    `--D. uptoni G84
    |    `--Leptophion Cameron 1901 [incl. Coiloneura Szépligeti 1905, Spilophion Cameron 1905] G84
    |         |--*L. longiventris [incl. L. chopini, *Coiloneura melanostigma] G84
    |         |--L. anici G84
    |         |--L. antennatus G84
    |         |--L. iochus G84
    |         |--*Spilophion’ maculipennis G84
    |         |--L. tetus G84
    |         |--L. unicalcaratus G84
    |         `--L. yampus G84
    |--Tryphoninae A71
    |    |  i. s.: Cteniscus A71
    |    |         Anisoctenion alacer A71
    |    |         Erromenus Holmgren 1855 HP-W05
    |    |         Ctenochira Förster 1855 HP-W05
    |    |         Eridolius Förster 1868 HP-W05
    |    |         Polyblastus Hartig 1837 HP-W05
    |    |           `--P. cothurnatus A71
    |    |--Phytodietini G84
    |    |--Sphinctus [Sphinctini] G84
    |    |--Idiogrammatini G84
    |    |--Ankylophon Gauld 1984 [Ankylophonini] G84
    |    |    `--*A. obligatus Gauld 1984 G84
    |    |--Exenterus A71 [Exenterini G84]
    |    |    |--E. abruptorius A71
    |    |    |--E. adspersus A71
    |    |    `--E. amictorius A71
    |    |--Tryphonini G84
    |    |    |--Grypocentrus G84
    |    |    `--Tryphon A71
    |    |         |--T. incestus A71
    |    |         `--T. semirufus A71
    |    `--Oedemopsini [Eclytini, Thymaridini] G84
    |         |--Debophanes Gauld 1984 G84
    |         |    `--*D. areolatus Gauld 1984 G84
    |         |--Thymaris Foerster 1869 [=Thymarus Thomson 1883] G84
    |         |    `--‘Mesoleptus’ tener [incl. *Thymaris pulchricornis] G84
    |         `--Oedemopsis Tschek 1869 (see below for synonymy) G84
    |              |--‘Tryphon’ scabriculus [=*Hybophanes scabriculus; incl. *Oedemopsis rogenhoferi] G84
    |              |--O. davisi [incl. Tryphon nasutus, *Campothreptus nasutus, *Zarhynchus nasutus] G84
    |              `--O. hobartensis G84
    |--Tersilochinae G84
    |    |--Barycnemis G84
    |    |--Heterocola G84
    |    |--Tersilochus conotracheli G84
    |    |--Allophroides G84
    |    |--Allophrys Foerster 1869 G84
    |    |    `--*A. oculatus [=Thersilochus oculatus] G84
    |    |--Areyonga Gauld 1984 G84
    |    |    `--*A. eremica Gauld 1984 G84
    |    |--Horstmannolochus Gauld 1984 G84
    |    |    `--*H. pulchripennis [=Thersilochus pulchripennis] G84
    |    |--Petilochus Gauld 1984 G84
    |    |    `--*P. naumanni Gauld 1984 G84
    |    |--Probles Foerster 1869 G84
    |    |    `--‘Porizon’ erythrostomus [incl. *Probles melanarius] G84
    |    |--Sathropterus Foerster 1869 G84
    |    |    `--*S. pumilus [=Thersilochus pumilus] G84
    |    |--Phradis Foerster 1869 [incl. Eutomus Foerster 1869 non Hope 1838, Isurgus Foerster 1869] G84
    |    |    |--*P. brevis [=Thersilochus (*Phradis) brevis] G84
    |    |    `--‘Thersilochus’ morionellus [incl. *Isurgus lanceolatus, *Eutomus lanceolatus] G84
    |    |--Stethantyx Townes 1971 G84
    |    |    |--S. argentinensis G84
    |    |    |--S. nearctica G84
    |    |    `--S. parkeri G84
    |    `--Diaparsis Foerster 1869 (see below for synonymy) G84
    |         |--‘Porizon’ truncatus [incl. P. nutritor, *Diaparsis nutritor] G84
    |         |--*Luchatema’ baldufi G84
    |         |--D. francisci Girault 1933 G33
    |         |--D. nigroguttipennis Girault 1933 G33
    |         |--‘Thersilochus (*Ischnobatis)’ stramineus G84
    |         `--D. temporalis G84
    |--Oxytorinae [Microleptinae] G84
    |    |--Cylloceria [incl. Asphragis] G84
    |    |--Microleptes G84
    |    |--Oxtorus G84
    |    |--Tatogaster G84
    |    |--Hyperacmus G84
    |    |--Plectiscidea G84
    |    |--Symplecis G84
    |    |--Sphingozona G84
    |    |--Aperileptus Foerster 1869 G84
    |    |    `--*A. albipalpus [=Plectiscus albipalpus] G84
    |    |--Helictes Haliday 1837 [=Myriarthrus Foerster 1869; incl. Paipila Cameron 1905] G84
    |    |    |--*H. erythrostoma [=Ichneumon erythrostoma, *Myriarthrus erythrostoma] G84
    |    |    `--*Paipila’ longipes G84
    |    |--Laepserus Foerster 1869 [incl. Acroblapticus Schmiedeknecht 1911, Gnathochorisis Foerster 1869] G84
    |    |    |--*L. crassulus [=Blapticus crassulus] G84
    |    |    |--‘Blapticus’ dentifer [=*Acroblapticus dentifer] G84
    |    |    `--*Gnathochorisis’ flavipes G84
    |    |--Proclitus Foerster 1869 (see below for synonymy) G84
    |    |    |--*P. grandis G84
    |    |    |--*Clepticus’ praetor [=*Mischoxorides praetor] G84
    |    |    `--*Aclastoneura’ tricolor G84
    |    |--Megastylus Schiødte 1838 (see below for synonymy) G84
    |    |    |--*M. cruentator [incl. M. mediator, *Idioxenus mediator] G84
    |    |    |--‘Myriarthrus’ cingulator [=*Myriarthridea cingulator] G84
    |    |    |--*Dicolus’ insectator G84
    |    |    |--*Letosha’ longicoxis G84
    |    |    `--*Miomeroides’ transsylvanicus G84
    |    `--Eusterinx Foerster 1869 (see below for synonymy) G84
    |         |--*E. oligomera G84
    |         |--*Dallatorrea’ armata G84
    |         |--‘Catomicrus’ bispinosa (see below for synonymy) G84
    |         |--*Cymodusoides’ gracilis G84
    |         |--‘Tryphon’ pusillus [incl. *Catomicrus trichops] G84
    |         |--*Holomeristus’ tenuicinctus G84
    |         `--‘Catastenus’ trifasciatus [incl. *Trestis tricincta] G84
    |--Metopiinae G84
    |    |--Chorinaeus funebris G84
    |    |--Carria Schmiedeknecht 1924 G84
    |    |    `--*C. paradoxa G84
    |    |--Sciron Fitton in Gauld 1984 G84
    |    |    `--*S. fundator Fitton in Gauld 1984 G84
    |    |--Trieces Townes 1946 G84
    |    |    `--*T. texanus [=Exochus texanus] G84
    |    |--Hypsicera Latreille 1829 (see below for synonymy) G84
    |    |    |--*H. femoralis (see below for synonymy) G84
    |    |    `--H. curvator G84
    |    |--Triclistus Foerster 1869 G84
    |    |    |--*T. podagricus [=Exochus podagricus] G84
    |    |    `--T. pygmaeus G84
    |    |--Drepanoctonus Pfankuch 1911 [incl. Zonopius Benoit 1961] G84
    |    |    |--*D. tibialis G84
    |    |    |--D. bifasciatus G84
    |    |    `--*Zonopius’ brevicinctus G84
    |    |--Seticornuta Morley 1913 [incl. Megatrema Cameron 1907 non Sowerby 1823] G84
    |    |    |--*Megatrema’ albopilosa [incl. *Seticornuta albicalcar] G84
    |    |    |--S. apicalis G84
    |    |    `--S. terminalis G84
    |    |--Exochus Gravenhorst 1829 [incl. Amesolytus Foerster 1869, Mima Davis 1897, Xanthexochus Morley 1913] G84
    |    |    |--*E. gravipes [=Ichneumon gravipes] G84
    |    |    |--*Amesolytus’ ferrugineus G84
    |    |    |--*Xanthexochus’ scutellatus G84
    |    |    `--*Mima’ washingtonensis G84
    |    |--Metopius Panzer 1806 (Metopius) [=Peltopius Clément 1927; incl. Peltastes Illiger 1807] G84
    |    |    |--*M. (M.) vespoides (see below for synonymy) G84
    |    |    |--M. (M.) crassicornis G84
    |    |    |--M. (M.) michaelseni G84
    |    |    `--M. (M.) unifenestratus [=M. (Ceratopius) unifenestratus] G84
    |    `--Colpotrochia Holmgren 1856 (see below for synonymy) G84
    |         |--C. cincta [=Sphex cincta; incl. Ichneumon elegantulus, *Colpotrochia elegantula] G84
    |         |--*Aithris’ coenutus G84
    |         |--*Sychnoleteroides’ flavus G84
    |         |--*Exochoides’ mexicana G84
    |         |--*Colpotrochioides’ orientalis G84
    |         |--C. pilosa [=*Inoresa pilosa] G84
    |         |--‘Exochoides’ texana [=*Ischyrocnemopsis texana] G84
    |         |--‘Tryphon’ trifasciatus [=*Alcocerus trifasciatus] G84
    |         `--*Scallama’ trilineata G84
    `--Ichneumoninae A71
         |  i. s.: Amblyteles subfuscus A71
         |         Barichneumon A71
         |--Ichneumonini G84
         |--Phaeogenini G84
         |--Platylabini G84
         |--Trogini G84
         |--Ceratojoppini G84
         |--Eurylabus [Eurylabini] G84
         |--Goedartia [Goedartiini] G84
         |--Heterischnus [Heterischnini] G84
         |--Zimmeriini G84
         |--Magwengiella Heinrich 1938 [Ctenocalini] G84
         |    `--*M. obtusa G84
         |--Ischnojoppa Kriechbaumer 1898 [incl. Bodargus Cameron 1902; Ischnojoppini] G84
         |    `--I. luteator (see below for synonymy) G84
         |--Alomyini G84
         |    |--Alomya G84
         |    `--Megalomya G84
         |--Listrodromini G84
         |    |--Validentia G84
         |    |--Anisobas G84
         |    |--Neotypus G84
         |    `--Listrodromus Wesmael 1845 [incl. Sogna Cameron 1907] G84
         |         |--*L. nycthemerus [=Ichneumon nycthemerus] G84
         |         `--*Sogna’ crassipes G84
         |--Protichneumonini G84
         |    |--Catadelphus G84
         |    |--Yeppoona Gauld 1984 G84
         |    |    `--*Y. grandis Gauld 1984 G84
         |    `--Coelichneumon G84
         |         |--‘Ichneumon’ comitator G84
         |         `--C. iridipennis [=Ichnneumon iridipennis] G84
         `--Oedicephalini [Notosemini] G84
              |--Notosemus G84
              |--Oedicephalus G84
              |--Aulojoppa G84
              |--Bambuscopus G84
              `--Imeria Cameron 1903 (see below for synonymy) G84
                   |--*I. albomaculata G84
                   |--*Elasmognathus’ cephalotes [=*Elasmognathias cephalotes] G84
                   |--*Caenojoppa’ longitarsis G84
                   `--‘Caenojoppa (*Imeriella)’ seyrigi G84

Ichneumonidae incertae sedis:
  Tanychora petiolata R02
  Tanychorella R02
  Rhyssella A71
    |--R. approximator I92
    `--R. curvipes A71
  Oocenteter A71
  Exeristes G84
    |--E. comstockii A71
    `--E. ruficollis G84
  Glabrodorsum stokesii H91
  Sinicorussus luzhongensis GE05
  Ariostonia Girault 1933 G33
    `--*A. fusciapex Girault 1933 G33

Anomalon Panzer 1804 [=Anomalum Schulz 1906, incl. Erythrophion Cameron 1906, Microcremastus Hedwig 1961, Nototrychys Marshall 1872, Ophiononeura Cameron 1904, Pseudonototrachys Meyer 1930, Stictophion Cameron 1906, Trachynotus Gravenhorst 1829 non Latreille 1829, Trachyopterus Morley 1912; Anomalonini] G84

Astiphromma Foerster 1869 [=Astiphrommus Thomson 1886; incl. Demophorellus Hedwig 1955, Dolichochorus Strobl 1904, Mesochorella Szépligeti 1911, Pseudacoenitus Kiss 1924] G84

‘Catomicrus’ bispinosa [incl. C. alpigenus, *Acanthostroblia alpigena, *Ischyracis alpigenus, *Stroblia alpigena] G84

Cidaphus Foerster 1869 [incl. Mater Schulz 1911, Ophthalmochorus Roman 1925, Plesiophthalmidea Viereck 1914, Plesiophthalmus Foerster 1869 non Motschoulsky 1858, Tetragonalys Morley 1913] G84

Colpotrochia Holmgren 1856 [incl. Aethris Schulz 1906, Aithris Cameron 1900, Alcocerus Foerster 1869, Exochoides Cresson 1868, Inoresa Cameron 1909, Ischyrocnemopsis Ashmead 1900, Scallama Cameron 1899] G84

Cryptinae [Gelinae, Hemiteloidae, Pezomachoidae, Pezomachoides, Phygadeuonides, Phygadeuontinae, Phygadeuontoidae, Stilpnoidae] D02

Diaparsis Foerster 1869 [=Diaparsus Thomson 1889; incl. Ischnobatis Foerster 1869, Luchatema Walkley 1956] G84

Euceros Gravenhorst 1829 [=Eumesius Westwood 1840, Omaloceros Giraud 1857; incl. Pseudasthenara Uchida 1930, Tautozelus Foerster 1869; Eucerotinae] G84

Euceros pruinosus (Gravenhorst 1829) HP-W05 [=Tryphon pruinosus G84; incl. *E. crassicornis G84, *Eumesius crassicornis G84, *Omaloceros crassicornis G84]

Eusterinx Foerster 1869 [incl. Acanthostroblia Roman 1925, Catomicrus Thomson 1888, Cymodusoides Viereck 1925, Dallatorrea Ashmead 1902, Holomeristus Foerster 1869, Ischyracis Foerster 1869, Stroblia Schmiedeknecht 1911 non Pokorny 1893, Trestis Foerster 1869] G84

Hypsicera Latreille 1829 [=Metacoelus Foerster 1869, Polyclistus Foerster 1869; incl. Plesioexochus Cameron 1905] G84

*Hypsicera femoralis [=Ichneumon femoralis, *Metacoelus femoralis, *Polyclistus femoralis; incl. *Plesioexochus rufipes] G84

Imeria Cameron 1903 [incl. Caenojoppa Cameron 1905, Elasmognathias Ashmead 1906, Elasmognathus Ashmead 1905, Imeriella Heinrich 1938] G84

Ischnojoppa luteator [=Ichneumon luteator; incl. Is. adspersor, Is. agraensis, Is. akonis, Joppa lutea, *Is. lutea, *Bodargus rufus, Is. rufa] G84

Lycorina Holmgren 1859 [incl. Chlorolycorina Cushman 1920, Gonioglyphus Seyrig 1932, Toxophoroides Cresson 1873; Lycorininae] G84

Megastylus Schiødte 1838 [=Megalostylus Schulz 1906; incl. Dicolus Foerster 1869, Idioxenus Foerster 1869, Letosha Cameron 1909, Miomeroides Kiss 1924, Myriarthridea Viereck 1914] G84

Mesochorus Gravenhorst 1829 [incl. Cryptochorus Aubert 1965, Edrisa Cameron 1907, Zamesochorus Viereck 1912] G84

*Metopius (Metopius) vespoides [=Sphex vespoides, *Peltopius vespoides; incl. Ichneumon necatorius, *Peltastes necatorius] G84

Oedemopsis Tschek 1869 [=Oedematopsis Morley 1908; incl. Campothreptus Foerster 1869, Hybophanes Foerster 1869, Zarhynchus Ashmead 1900 non Oberholser 1899] G84

Pristomerus Curtis 1836 [incl. Neopristomerus Viereck 1912, Nesanomalon Morley 1913, Pristocelus Szépligeti 1905, Pristomeridia Ashmead 1900] G84

Proclitus Foerster 1869 [incl. Aclastoneura Kriechbaumer 1896, Clepticus Haliday 1839 non Cuvier 1829, Mischoxorides Ashmead 1900] G84

Syrphoctonus Foerster 1869 [incl. Homocidus Morley 1911, Homoporus Thomson 1890 non Thomson 1878, Homotropus Foerster 1869] G84

Trathala Cameron 1899 [incl. Epicremastus Szépligeti 1905, Haristaeus Cameron 1910, Paurolexis Cameron 1907] G84

*Type species of generic name indicated

REFERENCES

[A71] Askew, R. R. 1971. Parasitic Insects. Heinemann Educational Books: London.

[D02] Donovan, B. J. 2002. Description of Sphecophaga orientalis sp. n. (Hymenoptera: Ichneumonidae: Cryptinae), a potential parasitoid of Vespula spp. (Hymenoptera: Vespidae: Vespinae) in New Zealand. New Zealand Entomologist 25: 3-15.

[E12] Evenhuis, N. L. 2012. Publication and dating of the Exploration Scientifique de l'Algérie: Histoire Naturelle des Animaux Articulés (1846–1849) by Pierre Hippolyte Lucas. Zootaxa 3448: 1–61.

[G84] Gauld, I. D. 1984. An Introduction to the Ichneumonidae of Australia. British Museum (Natural History).

[G25a] Girault, A. A. 1925a. Indications (in new insects) of ruling power and laws in nature. Privately published (reprinted Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 184-186).

[G25b] Girault, A. A. 1925b. An essay on when a fly is lovable, the ceremony of baptizing some and unlovely hate. Privately published (reprinted Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 187-190).

[G32] Girault, A. A. 1932. New lower Hymenoptera from Australia and India. Privately published (reprinted Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 293-298).

[G33] Girault, A. A. 1933. Some beauties inhabitant not of commercial boudoirs but of Nature’s bosom, notably new insects. Privately published (reprinted Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 299-303).

[GE05] Grimaldi, D., & M. S. Engel. 2005. Evolution of the Insects. Cambridge University Press: New York.

[HP-W05] Heitland, W., & H. Pschorn-Walcher. 2005. Biology and parasitoids of the peculiar alder sawfly, Platycampus luridiventris (Fallen) (Insecta, Hymenoptera, Tenthredinidae). Senckenbergiana Biologica 85 (2): 215-231.

[H91] Howarth, F. G. 1991. Environmental impacts of classical biological control. Annual Review of Entomology 36: 485-509.

[IK05] Idris, A. B., & L. Khairiza. 2005. A catalogue of Neoxoridini and Rhyssini (Ichneumonidae: Pimplinae) from Malaysia. Serangga 10 (1-2): 93-110.

[IR05] Idris, A. B., & A. Rizki. 2005. Notes on the tribe Ephialtini (Hymenoptera: Ichneumonidae: Pimplinae) of Malaysia. Serangga 10 (1-2): 111-126.

[I92] Imes, R. 1992. The Practical Entomologist. Aurum Press: London.

[M83] Martin, N. A. 1983. Miscellaneous observations on a pasture fauna: An annotated species list. DSIR Entomology Division Report 3: 1-98.

[N88] Noonan, G. R. 1988. Biogeography of North American and Mexican insects, and a critique of vicariance biogeography. Systematic Zoology 37 (4): 366-384.

[R02] Rasnitsyn, A. P. 2002. Superorder Vespidea Laicharting, 1781. Order Hymenoptera Linné, 1758 (=Vespida Laicharting, 1781). In History of Insects (A. P. Rasnitsyn & D. L. J. Quicke, eds) pp. 242-254. Kluwer Academic Publishers: Dordrecht.

SNS07] Sudheer, K., T. C. Narendran & A. Saji. 2007. Taxonomy of a new ichneumonid genus from the Middle East (Insecta, Hymenoptera, Ichneumonidae). Senckenbergiana Biologica 87 (2): 185-188.

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