Cryptini

Goryphus basilaris, from here.

Belongs within: Ichneumonidae.
Contains: Gotra.

The Cryptini are a group of mostly medium-sized or large ichneumon wasps with a single bulla in forewing vein 2m-cu and without complete propodeal areolation (Gauld 1984).

<==Cryptini [Mesostenini] G84
    |--Ceratomansa Cushman 1922 [Baryceratina] G84
    |    |--*C. prima G84
    |    `--C. spinifrons (Brullé 1846) [=Mesostenus spinifrons; incl. C. curvilineata] G84
    |--Gabuniina G84
    |    |--Eurycryptus Cameron 1901 [incl. Alriada Cameron 1911, Didiaspis Seyrig 1952, Neotorbda Uchida 1932] G84
    |    |    |--*E. laticeps G84
    |    |    |--‘*Didiaspis’ fondamentalis G84
    |    |    |--‘Torbda (*Neotorbda)’ sakaguchii G84
    |    |    `--‘*Alriada’ spilocephala G84
    |    `--Xanthocryptus Cameron 1901 [incl. Lorentzia Cameron 1911 non Cossman 1908] G84
    |         |--*X. robustus G84
    |         |--‘*Lorentzia’ flavomaculata G84
    |         |--X. lugubris G84
    |         `--X. novozealandicus [=Arhytis novozealandicus; incl. X. albopictus] G84
    |--Mesostenina G84
    |    |--Mesostenus Gravenhorst 1829 (see below for synonymy) G84
    |    |    |--*M. transfuga [=*Stenaraeus transfuga] G84
    |    |    |--M. longicaudis [incl. Coleocentrus texanus, *Derocentrus texanus] G84
    |    |    |--‘*Umlima’ penetralis G84
    |    |    |--M. physoscelus G84
    |    |    `--M. zarathae G17
    |    |--Junctivena Gauld 1984 G84
    |    |    `--*J. gallowayi Gauld 1984 G84
    |    |--Diloa Cheesman 1936 G84
    |    |    |--*D. pudens [=Mesostenus pudens] G84
    |    |    `--D. antipodialis G84
    |    |--Paranacis Gauld 1984 G84
    |    |    `--*P. brunnea Gauld 1984 G84
    |    `--Tomagotra Gauld 1984 G84
    |         `--*T. roddi Gauld 1984 G84
    |--Ischnina G84
    |    |--Ischnus Gravenhorst 1892 (see below for synonymy) G84
    |    |    |--‘Ichneumon’ inquisitorius (see below for synonymy) G84
    |    |    |--‘Habrocryptus’ assertorius MS01
    |    |    |--‘*Aglaocryptus’ curvimaculatus G84
    |    |    |--I. migrator P93
    |    |    `--I. nigricollis MS01
    |    |--Nebostenus Gauld 1984 G84
    |    |    |--*N. crypticus Gauld 1984 G84
    |    |    `--N. terebratus Gauld 1984 G84
    |    `--Xylostenus Gauld 1984 G84
    |         `--*X. curtus Gauld 1984 G84
    |--Osprynchotina G84
    |    |--Osprynchotus BM76
    |    |--Stenarella Szépligeti 1916 [incl. Orientostenaraeus Uchida 1930, Parasilsila Cheesman 1936] G84
    |    |    |--*S. gladiator [=Ichneumon gladiator] G84
    |    |    |--‘Mesostenus’ insidiator [incl. *Orientostenaraeus chinensis] G84
    |    |    `--S. victoriae [=Mesostenus victoriae; incl. *Parasilsila trilineata] G84
    |    `--Iaria Cheesman 1936 G84
    |         |--*I. palata [=Mesostenus palatus] G84
    |         `--I. papiliomaculata [incl. I. veitchi] G84
    |--Goryphina G84
    |    |--Goryphus Holmgren 1868 (see below for synonymy) G84
    |    |    |--*G. basilaris G84
    |    |    |--‘*Psacus’ areolaris G84
    |    |    |--‘*Brachycoryphus’ calabaricus G84
    |    |    |--‘*Miramilia’ communis G84
    |    |    |--‘*Scenopathus’ ferrugineus G84
    |    |    |--G. flavocinctus G84
    |    |    |--‘*Cratocryptus’ maculiceps (see below for synonymy) G84
    |    |    |--‘*Filistina’ maculipennis G84
    |    |    `--G. turneri G84
    |    `--Syntrips Gauld 1984 G84
    |         `--*S. maculatus Gauld 1984 G84
    `--Lorio Cheesman 1936 [Ceratocryptina] G84
         `--*L. austerus [=Mesostenus austerus] G84

Cryptini incertae sedis:
  ‘Poecilocryptus’ Kriechbaumer 1901 non Cameron 1901 G84
  Biconus G84
  Isotima G84
  *Buodias ruficoxis G84
  Melcha G84
  Aprix Townes 1961 G84
    `--*A. nutatoria (Fabricius 1775) [=Ichneumon nutatorius] G84
  Euchalinus Townes 1961 G84
    `--*E. balteatus [=Skeatia balteata] G84
  Glabridorsum Townes 1970 G84
    `--*G. stokesii [=Gambrus stokesii] G84
  Hackerocryptus Gauld 1984 G84
    `--*H. dentatus Gauld 1984 G84
  Myrmeleonostenus Uchida 1936 G84
    `--*M. babai G84
  Neaprix Gauld 1984 G84
    `--*N. insolens Gauld 1984 G84
  Takastenus Uchida 1931 G84
    `--*T. longidentatus G84
  Arhytis Townes 1970 G84
    `--*A. maculiscutis [=Echthrus maculiscutis] G84
  Allophatnus Cameron 1905 [incl. Phaedraspis Cameron 1906, Stictocryptus Cameron 1907] G84
    |--*A. fulvipes G84
    `--A. fulvitergus [=Cryptus fulvitergus; incl. *Phaedraspis rufobalteata, *Stictocryptus testaceus] G84
  Gambroides Betrem 1941 [incl. Vadonina Seyrig 1952] G84
    |--*G. javensis [=Eripternimorpha javensis] G84
    `--‘*Vadonina’ nimbipennis G84
  Irabatha Cameron 1906 G84
    |--I. albispina G84
    `--I. cairnsensis G84
  Listrognathus (Suvalta Cameron 1903) [incl. Stivadens Townes 1961] G84
    |--L. (*S.) rugifrons G84
    `--L. (S.) annulipes [=L. (*Stivadens) annulipes] G84
  Lophoglutus Gauld 1984 G84
    |--*L. bouceki Gauld 1984 G84
    `--L. monticolus G84
  Necolio Cheesman 1936 [incl. Afrocryptus Seyrig 1952] G84
    |--*N. jugosus G84
    `--‘*Afrocryptus’ imperialis G84
  Thelodon Townes 1961 G84
    |--*T. spilonotus [=Silsila spilonota] G84
    `--T. elongatus [incl. T. papuensis] G84
  Cremnocryptus Cushman 1945 G84
    |--*C. spiniferus [=Polyaenus spiniferus] G84
    `--C. arrogans G84
  Arthula Cameron 1900 [incl. Kuniocryptus Sonan 1937, Orientocryptus Uchida 1931] G84
    |--*A. brunneocornis G84
    |--‘*Orientocryptus’ flavofasciatus [=*Kuniocryptus flavofasciatus] G84
    `--‘*Orientocryptus’ formosanus G84
  Stiromesostenus Cameron 1911 [incl. Erythromesostenus Cameron 1911] G84
    |--*S. xanthostomus G84
    |--S. albiorbitalis G84
    `--S. rufus [=*Erythromesostenus rufus] G84
  Anacis Porter 1967 [incl. Baeocryptus Porter 1967, Dochmidium Porter 1967] G84
    |--*A. festiva G84
    |--‘*Dochmidium’ camponotus G84
    |--A. exul G84
    `--‘Ichneumon (Cryptus)’ rubripes [=*Baeocryptus rubripes] G84
  Ateleute Foerster 1869 (see below for synonymy) G84
    |--*A. linearis G84
    |--‘*Ptychostenus’ minusculae G84
    |--‘*Talorga’ spinipes G84
    `--‘*Tsirirella’ tsiriria G84
  Cryptus Fabricius 1804 non Jurine 1801 (nom. rej.) non Panzer 1804 (n. o.) G84
    |--C. annulicornis Lucas 1849 E12
    |--C. armatus Lucas 1849 E12
    |--C. bicolor Lucas 1849 E12
    |--C. incubitor MS01
    |--C. levaillantii Lucas 1849 E12
    |--C. murorum MS01
    `--C. viduatorius MS01
  Gotra G84

Ateleute Foerster 1869 [=Ateleuta Schulz 1906; incl. Psychostenus Uchida 1955, Talorga Cameron 1911, Tsirirella Seyrig 1952] G84

‘*Cratocryptus’ maculiceps [=*Cratocryptodes maculiceps, *Cratocryptoides maculiceps, *Loiada maculiceps] G84

Goryphus Holmgren 1868 [incl. Brachycoryphus Kriechbaumer 1894, Cratocryptodes Schulz 1906, Cratocryptoides Schmiedeknecht 1908, Cratocryptus Cameron 1905 non Thomson 1873, Filistina Cameron 1902, Loiada Cameron 1905, Miramilia Seyrig 1952, Psacus Holmgren 1868, Scenopathus Enderlein 1918] G84

‘Ichneumon’ inquisitorius [incl. Ic. porrectorius, *Habrocryptus porrectorius, *Ischnus porrectorius, *Erythrocryptus rufus] G84

Ischnus Gravenhorst 1892 [=Habrocryptus Thomson 1873; incl. Aglaocryptus Cameron 1903, Erythrocryptus Cameron 1905] G84

Mesostenus Gravenhorst 1829 [=Stenaraeus Thomson 1896; incl. Derocentrus Cushman 1919, Umlina Cameron 1902] G84

*Type species of generic name indicated

REFERENCES

[BM76] Bohart, R. M., & A. S. Menke. 1976. Sphecid Wasps of the World. University of California Press: Berkeley.

[E12] Evenhuis, N. L. 2012. Publication and dating of the Exploration Scientifique de l'Algérie: Histoire Naturelle des Animaux Articulés (1846–1849) by Pierre Hippolyte Lucas. Zootaxa 3448: 1–61.

[G84] Gauld, I. D. 1984. An Introduction to the Ichneumonidae of Australia. British Museum (Natural History).

[G17] Girault, A. A. 1917. New Javanese Hymenoptera. Privately published (reprinted Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 59–71).

[MS01] Mocsáry, A., & V. Szépligeti. 1901. Hymenopterák [Hymenopteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 121–169. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[P93] Pittaway, A. R. 1993. The Hawkmoths of the Western Palaearctic. Harley Books: Colchester.