Scolytinae

Cryphalus ruficollis, photographed by J. E. Mercado.


Belongs within: Phytophaga.
Contains: Pityophthorina, Corthylina, Scolytodes, Ips, Cyrtogenius, Xyleborus, Hypothenemus, Scolytogenes.

The Scolytinae are a subgroup of the bark beetles in which the bases of the elytra are simple and flat (Wood 1986).

Characters (from Wood 1986): Basal margins of elytra forming a straight transverse line across body, unarmed, rarely with a weakly elevated continuous line; scutellum usually large, flat (rarely absent or highly modified); pronotum weakly to strongly dclivous on anterior half and usually armed by many asperate crenulations, particularly on median half; head usually partly or entirely concealed from dorsal aspect, somewhat narrower; rarely with scales or deeply divided setae.

<==Scolytinae
    |--Corthylini [Amphicranidae, Araptidae, Corthyli, Pityophthoridae] W86
    |    |--Pityophthorina W86
    |    `--Corthylina W86
    |--Cactopinus Schwarz 1899 [incl. Cactopinorus Bright 1967; Cactopinini] W86
    |    |--*C. hubbardi W86
    |    |--C. cactophthorus [=*Cactopinorus cactophthorus] W86
    |    |--C. depressus W86
    |    `--C. desertus W86
    |--Scolytoplatypus Schaufuss 1893 (see below for synonymy) W86
    |    |--*S. permirus W86
    |    |--S. congonus W86
    |    |--S. mikado [=*Strophionocerus mikado, *Taeniocerus mikado] W86
    |    |--S. papuanus W86
    |    `--S. tycon [=*Spongocerus tycon] W86
    |--Carphodicticini W86
    |    |--Dendrodicticus Schedl 1958 W86
    |    |    `--*D. argentiniae W86
    |    |--Carphodicticus Wood 1971 W86
    |    |    `--*C. cristatus W86
    |    `--Craniodicticus Blandford 1895 W86
    |         `--*C. mucronatus W86
    |--Ctenophorini [Ctenophoridae, Hexacolidae, Problechilidae] W86
    |    |--Scolytodes W86
    |    |--Gymnochilus Eichhoff 1867 [=Problechilus Eichhoff 1878; incl. Meringopalpus Hagedorn 1905] W86
    |    |    `--*G. zonatus [=*Problechilus zonatus; incl. *Meringopalpus fallax] W86
    |    |--Pycnarthrum Eichhoff 1878 (see below for synonymy) W86
    |    |    `--P. hispidum (see below for synonymy) W86
    |    `--Microborus Blandford 1897 [incl. Pseudocrypturgus Eggers 1919] W86
    |         `--*M. boops [incl. *Pseudocrypturgus camerunus] W86
    |--Xyloterini [Xyloteroideae] W86
    |    |--Xyloterinus Swaine 1918 W86
    |    |    `--X. politus [=Bostrichus politus] W86
    |    |--Indocryphalus Eggers 1939 [incl. Dendrotrypum Schedl 1951] W86
    |    |    |-I. intermedius [=Xyloterus intermedius; incl. *Indocryphalus malaisei] W86
    |    |    `--I. aceris [=Xyloterus aceris, *Dendrotrypum aceris] W86
    |    `--Trypodendron Stephens 1830 [incl. Xyloterus Erichson 1836] W86
    |         |--*T. domesticus (Linnaeus 1758) [=Dermestes domesticus] W86
    |         |--T. betulae W86
    |         |--T. lineatum [=Bostrichus lineatus, *Xyloterus lineatus] W86
    |         `--T. rufitarsis W86
    |--Crypturgini W86
    |    |--Aphanarthrum Wollaston 1854 W86
    |    |    `--*A. euphorbiae W86
    |    |--Deropria Enderlein 1929 W86
    |    |    `--*D. elongata [=Aphanarthrum elongatum] W86
    |    |--Dolurgus Eichhoff 1868 W86
    |    |    `--*D. pumilus [=Hylastes pumilus] W86
    |    |--Coleobothrus Enderlein 1929 W86
    |    |    `--*C. jandiacus W86
    |    |--Cisurgus Reitter 1894 W86
    |    |    `--*C. filum W86
    |    `--Crypturgus Erichson 1836 W86
    |         |--*C. pusillus [=Bostrichus pusillus] W86
    |         `--C. cinereus JKW03
    |--Xyloctonini W86
    |    |--Xyloctonus Eichhoff 1872 W86
    |    |    `--*X. scolytoides W86
    |    |--Ctonoxylon Hagedorn 1910 W86
    |    |    `--*C. auratum W86
    |    |--Cryphalomimus Eggers 1927 W86
    |    |    `--*C. striatus W86
    |    |--Scolytomimus Blandford 1895 [incl. Neoxyloctonus Eggers 1923, Scolytocleptes Schedl 1962] W86
    |    |    |--*S. dilutus W86
    |    |    |--S. maculatus [=*Scolytocleptes maculatus] W86
    |    |    `--S. philippinensis [=*Neoxyloctonus philippinensis] W86
    |    `--Glostatus Schedl 1939 (see below for synonymy) W86
    |         |--*G. declividepressus W86
    |         |--G. acaciae [=*Apoglostatus acaciae] W86
    |         |--G. nigrivestris [=Ctonocryphus nigrivestris, *Paraglostaus nigrivestris] W86
    |         |--G. seydeli [=*Rhopalocryphus seydeli] W86
    |         `--*Ctonocryphus’ xyloctonus W86
    |--Ipini W86
    |    |--Ips W86
    |    |--Dendrochilus Schedl 1957 W86
    |    |    `--*D. strombosiopsis W86
    |    |--Pityokteines Fuchs 1911 [incl. Orthotomides Wood 1951] W86
    |    |    |--*P. curvidens [=Ips curvidens] W86
    |    |    `--P. lasiocarpi [=Orthotomicus lasiocarpi, *Orthotomides lasiocarpi] W86
    |    |--Orthotomicus Ferrari 1867 [=Neotomicus Fuchs 1911, Onthotomicus] W86
    |    |    |--*O. laricis [=Bostrichus laricis, *Neotomicus laricis] W86
    |    |    `--O. erosus JKW03
    |    |--Pityogenes Bedel 1888 [incl. Eggersia Lebedev 1926, Pityoceragenes Balachowsky 1947] W86
    |    |    |--*P. chalcographus [=Dermestes chalcographus] W86
    |    |    |--P. bidentatus [=Bostrichus bidentatus, *Eggersia bidentata] W86
    |    |    `--P. quadridens [=Bostrichus quadridens, *Pityoceragenes quadridens] W86
    |    `--Acanthotomicus Blandford 1894 [incl. Isophthorus Schedl 1938, Mimips Eggers 1932] W86
    |         |--*A. spinosus W86
    |         |--A. mimicus W86
    |         |--A. pilosus [=Ips pilosus, *Mimips pilosus] W86
    |         `--A. quadrituberculatus [=*Isophthorus quadrituberculatus] W86
    |--Scolytini [Camptoceridae] W86
    |    |--Scolytopsis Blandford 1896 W86
    |    |    `--*S. puncticollis W86
    |    |--Camptocerus Latreille 1829 W86
    |    |    |--*C. aeneipennis [=Hylesinus aeneipennis] W86
    |    |    `--C. auricomus W86
    |    |--Cnemonyx Eichhoff 1868 (see below for synonymy) W86
    |    |    |--*C. galeritus W86
    |    |    |--C. barbatus [=*Minulus barbatus] W86
    |    |    |--C. flavicornis [=*Loganius flavicornis] W86
    |    |    |--C. hylurgoides [=*Coptodryas hylurgoides, *Coptosomus hylurgoides] W86
    |    |    |--C. jucundus [=*Ceratolepis jucundus] W86
    |    |    `--C. panamensis W86
    |    `--Scolytus Geoffroy 1762 (see below for synonymy) W86
    |         |--*S. scolytus (see below for synonymy) W86
    |         |--S. claviger [=*Archaeoscolytus claviger] W86
    |         |--S. confusus [=Eccoptogaster confusus, *Confusoscolytus confusus] W86
    |         |--S. intricatus [=Eccoptogaster intricatus, *Tubuloscolytus intricatus] W86
    |         |--S. marawitzi [=*Pinetoscolytus marawitzi] W86
    |         |--S. multistriatus (see below for synonymy) W86
    |         |--S. pygmaeus [=Bostrichus pygmaeus, *Pygmaeoscolytus pygmaeus] W86
    |         `--S. rugulosus [=Bostrichus rugulosus, *Ruguloscolytus rugulosus] W86
    |--Micracini [Hylocuridae, Micracides] W86
    |    |--Pseudomicracis Eggers 1920 W86
    |    |    `--*P. elsae Eggers 1920 W86
    |    |--Micracisella Blackman 1928 [=Pseudomicracis Blackman 1920 non Eggers 1920] W86
    |    |    `--*M. opacicollis [=Micracis opacicollis, *Pseudomicracis opacicollis] W86
    |    |--Stenoclyptus Blackman 1943 W86
    |    |    `--S. sulcatus [=Pseudothysanoes sulcatus; incl. *S. rhois] W86
    |    |--Phloeocleptus Wood 1956 W86
    |    |    `--*P. caudatus W86
    |    |--Thysanoes LeConte 1876 W86
    |    |    `--*T. fimbricornis W86
    |    |--Traglostus Schedl 1938 W86
    |    |    `--*T. exornatus W86
    |    |--Saurotocis Wood 1984 W86
    |    |    `--*S. tomicoides [=Micracidendron tomicoides] W86
    |    |--Phloeocurus Wood 1984 W86
    |    |    `--*P. africanus [=Hylocurus africanus] W86
    |    |--Micracis LeConte 1868 W86
    |    |    |--*M. suturalis W86
    |    |    `--M. lignator [incl. M. truncatus] W86
    |    |--Miocryphalus Schedl 1939 [incl. Afromicracis Schedl 1959] W86
    |    |    |--*M. natalensis [=Stephanoderes natalensis] W86
    |    |    `--M. kenyaensis [=*Afromicracis kenyaensis] W86
    |    |--Lanurgus Eggers 1920 (see below for synonymy) W86
    |    |    |--*L. barbatus W86
    |    |    |--L. catenatus [=*Micraciops catenatus] W86
    |    |    |--L. elongatus [=*Landolphianus elongatus] W86
    |    |    `--L. podocarpi [incl. *Pseudohylocurus caplandicus] W86
    |    |--Hylocurus Eichhoff 1872 [incl. Micracisoides Blackman 1920] W86
    |    |    |--*H. elegans W86
    |    |    |--H. effeminatus W86
    |    |    |--H. equidens W86
    |    |    |--H. inaequalis W86
    |    |    |--H. medius W86
    |    |    `--H. rudis [=Micracis rudis, *Micracisoides rudis] W86
    |    `--Pseudothysanoes Blackman 1920 (see below for synonymy) W86
    |         |--P. rigidus [=Cryphalus rigidus; incl. *P. drakei] W86
    |         |--P. coniferae [=*Aphanocleptus coniferae] W86
    |         |--P. dislocatus [=*Cryptocleptes dislocatus, *Cryptulocleptus dislocatus] W86
    |         |--P. mongolica [=*Gretschkinia mongolica] W86
    |         |--P. mucronatus W86
    |         |--P. securigerus [=*Chalcohyus securigerus] W86
    |         |--P. spinatus [=*Bostrichips spinatus] W86
    |         `--P. squamosus [=*Neoglostatus squamosus] W86
    |--Dryocoetini [=Dryocoetoideae, Taphrorychini, Thamnurginae] W86
    |    |--Cyrtogenius W86
    |    |--Toxophthorus Wood 1986 [=Toxophorus Eggers 1920 (preoc.)] W86
    |    |    `--*T. africanus (Eggers 1920) [=*Toxophorus africanus] W86
    |    |--Taphramites Schedl 1947 W86
    |    |    `--*T. gnathotrichus Schedl 1947 W86
    |    |--Thamnurgus Eichhoff 1864 W86
    |    |    `--*T. euphorbiae [=Bostrichus euphorbiae] W86
    |    |--Chiloxylon Schedl 1959 W86
    |    |    `--*C. rufulus W86
    |    |--Dactylotrypes Eggers 1927 W86
    |    |    `--D. longicollis [=Xyloterus longicollis; incl. *D. uyttenboogaarti] W86
    |    |--Dendrocranulus Schedl 1937 W86
    |    |    `--*D. tardus W86
    |    |--Lymantor Lovendal 1889 W86
    |    |    `--‘Tomicus’ coryli [incl. *Lymantor sepicola] W86
    |    |--Taphronurgus Reitter 1913 W86
    |    |    `--*T. exul [=Thamnurgus exul] W86
    |    |--Dolurgocleptes Schedl 1965 W86
    |    |    `--*D. malgassicus W86
    |    |--Dendrographus Schedl 1964 [incl. Protopityophthorus Schedl 1973] W86
    |    |    `--*D. pygmaeus [=Pelicerus pygmaeus; incl. *Protopityophthorus durus] W86
    |    |--Peridryocoetes Wood 1984 W86
    |    |    `--*P. nitens [=Ozodendron nitens] W86
    |    |--Ozopemon Hagedorn 1908 W86
    |    |    `--*O. regius W86
    |    |--Dryocoetiops Schedl 1957 W86
    |    |    |--*D. laevis [=Ozopemon laevis] W86
    |    |    `--D. coffeae W86
    |    |--Triotemnus Wollaston 1864 [incl. Cladoctoproctus Schedl 1975] W86
    |    |    |--*T. subretusus W86
    |    |    `--T. scrafa [=*Cladoctoproctus scrafa] W86
    |    |--Dryocoetes Eichhoff 1864 [=Anodius Motschulsky 1860 (nom. rej.); incl. Dryocoetinus Balachowsky 1949] W86
    |    |    |--*D. autographus [=Bostrichus autographus, *Anodius autographus] W86
    |    |    |--D. confusus W86
    |    |    `--D. villosus [=Bostrichus villosus, *Dryocoetinus villosus] W86
    |    |--Tiarophorus Schreiner 1882 (see below for synonymy) W86
    |    |    |--*T. elongatus W86
    |    |    |--T. camerunus [=*Hypaspistes camerunus, *Orthaspistes camerunus] W86
    |    |    `--T. scrutator [=Thamnurgus scrutator, *Pseudothamnurgus scrutator] W86
    |    |--Xylocleptes Ferrari 1867 [incl. Hylonius Nunberg 1973, Xestips Hagedorn 1912] W86
    |    |    |--*X. bispinus [=Bostrichus bispinus] W86
    |    |    |--X. brunneus [=*Hylonius brunneus] W86
    |    |    `--X. marginatus [=*Xestips marginatus] W86
    |    |--Taphrorychus Eichhoff 1878 (see below for synonymy) W86
    |    |    |--*T. bicolor [=Bostrichus bicolor] W86
    |    |    |--T. machnovskii [=Hypothenemus machnovskii, *Saliciphilus machnovskii] W86
    |    |    |--T. mikuniyamensis [=*Pseudopoecilips mikuniyamensis] W86
    |    |    `--T. primitus [=*Taphroterus primitus] W86
    |    `--Coccotrypes Eichhoff 1878 (see below for synonymy) W86
    |         |--*C. dactyliperda [=Bostrichus dactyliperda] W86
    |         |--C. advena [incl. *Thamnurgides persicae] W86
    |         |--C. carpophagus (see below for synonymy) W86
    |         |--C. rhizophorae [=*Spermatoplex rhizophorae, *Dendrurgus rhizophorae] W86
    |         `--C. sannio [=*Poecilips sannio] W86
    |--Xyleborini [Webbinae, Xylebori] W86
    |    |--Australoplatypus incompertus GE05
    |    |--Xyleborus W86
    |    |--Xyleborites Wickham 1913 W86
    |    |    `--*X. longipennis Wickham 1913 W86
    |    |--Sampsonius Eggers 1935 W86
    |    |    `--*S. sexdentatus W86
    |    |--Xyleborinus Reitter 1913 W86
    |    |    `--*X. saxeseni [=Bostrichus saxeseni] W86
    |    |--Schedlia Browne 1950 W86
    |    |    `--*S. sumatrana [=Xyleborus sumatranus] W86
    |    |--Dryocoetoides Hopkins 1915 W86
    |    |    `--D. capucinus [=Xyleborus capucinus; incl. *D. guatemalensis] W86
    |    |--Leptoxyleborus Wood 1980 W86
    |    |    `--*L. sordicauda [=Phloeotrogus sordicauda] W86
    |    |--Theoborus Hopkins 1915 W86
    |    |    `--*T. theobromae W86
    |    |--Euwallacea Hopkins 1915 W86
    |    |    `--*E. wallacei [=Xyleborus wallacei] W86
    |    |--Taurodemus Wood 1980 W86
    |    |    `--*T. sharpi [=Xyleborus sharpi] W86
    |    |--Mesoscolytus Broun 1904 W86
    |    |    `--*M. inurbanus W86
    |    |--Hadrodemius Wood 1980 W86
    |    |    `--*H. globus [=Xyleborus globus] W86
    |    |--Taphrodasus Wood 1980 W86
    |    |    `--*T. percorthylus [=Xyleborus percorthylus] W86
    |    |--Cryptoxyleborus Schedl 1937 W86
    |    |    `--*C. naevus W86
    |    |--Premnobius Eichhoff 1878 [incl. Premnophilus Browne 1962] W86
    |    |    |--*P. cavipennis W86
    |    |    `--P. joveri [=Xyleborus joveri, *Premnophilus joveri] W86
    |    |--Cnestus Sampson 1911 [incl. Tosaxyleborus Murayama 1950] W86
    |    |    |--*C. magnus W86
    |    |    `--C. murayamai [incl. *Tosaxyleborus pallidipennis] W86
    |    |--Coptoborus Hopkins 1915 [incl. Streptocranus Schedl 1939] W86
    |    |    |--C. vespatorius [=Xyleborus vespatorius; incl. *C. emarginatus] W86
    |    |    `--C. mirabilis [=*Streptocranus mirabilis] W86
    |    |--Arixyleborus Hopkins 1915 [incl. Xyleboricus Eggers 1923] W86
    |    |    |--*A. rugosipes W86
    |    |    `--A. canaliculatus [=*Xyleboricus canaliculatus] W86
    |    |--Terminalinus Hopkins 1915 [incl. Kelantanius Nunberg 1961] W86
    |    |    |--*T. terminaliae W86
    |    |    `--T. punctatopilosus [=Xyleborus punctatopilosus, *Kelantanius punctatopilosus] W86
    |    |--Eccoptopterus Motschulsky 1863 [incl. Eurydactylus Hagedorn 1909, Platydactylus Eichhoff 1886] W86
    |    |    |--E. spinosus [=Scolytus spinosus; incl. *Eccoptopterus sexspinosus] W86
    |    |    `--E. gracilipes [=*Platydactylus gracilipes, *Eurydactylus gracilipes] W86
    |    |--Ambrosiodmus Hopkins 1915 [incl. Browneia Nunberg 1963, Phloeotrogus Motschulsky 1863 (nom. rej.)] W86
    |    |    |--*A. tachygraphus [=Xyleborus tachygraphus] W86
    |    |    |--A. obliquecaudata [=*Phloeotrogus oliquecaudata] W86
    |    |    `--A. obliquus [=Pityophthorus obliquus; incl. Xyleborus illepidus, *Browneia illepida] W86
    |    |--Amasa Lea 1894 [incl. Anaxyleborus Wood 1980, Pseudoxyleborus Eggers 1930] W86
    |    |    |--*A. thoracicus W86
    |    |    |--*Pseudoxyleborus’ beesoni W86
    |    |    `--A. truncatus [=Tomicus truncatus, *Anaxyleborus truncatus] W86
    |    |--Coptodryas Hopkins 1915 [incl. Adryocoetes Eggers in Schedl 1952 (n. n.), Microperus Wood 1980] W86
    |    |    |--*C. confusa W86
    |    |    |--C. pullus [=Xyleborus pullus; incl. *Adryocoetes nitidus (n. n.)] W86
    |    |    `--C. theae [=Xyleborus theae, *Microperus theae] W86
    |    |--Webbia Hopkins 1915 (see below for synonymy) W86
    |    |    |--*W. dipterocarpi W86
    |    |    |--W. pabo W86
    |    |    |--W. subuculae [=*Prowebbia subuculae] W86
    |    |    `--W. trepanicauda [=Xyleborus trepanicauda, *Pseudowebbia trepanicauda] W86
    |    `--Xylosandrus Reitter 1913 [incl. Apoxyleborus Wood 1980] W86
    |         |--*X. morigerus [=Xyleborus morigerus] W86
    |         |--X. compactus MHG04
    |         |--X. mancus [=Xyleborus mancus, *Apoxyleborus mancus] W86
    |         `--X. retusus W86
    `--Cryphalini [Cryphaloidea, Eidophelinae, Ernoporinae, Trypophloeinae] W86
         |--Hypothenemus W86
         |--Scolytogenes W86
         |--Cryphalities Cockerell 1917 W86
         |    `--*C. rugosissimus Cockerell 1917 W86
         |--Periocryphalus Wood 1971 W86
         |    `--*P. pullus W86
         |--Trischidias Hopkins 1915 W86
         |    `--*T. georgiae W86
         |--Neocryphus Nunberg 1956 W86
         |    `--*N. argentinensis W86
         |--Stegomerus Wood 1967 W86
         |    `--*S. vulgaris W86
         |--Allernoporus Kurentzov 1941 W86
         |    `--*A. euonymi W86
         |--Coriacephilus Schedl 1939 W86
         |    `--*C. coriaceus [=Stephanoderes coriaceus] W86
         |--Ernocladius Wood 1980 W86
         |    `--*E. corpulentus [=Cryphalus corpulentus] W86
         |--Cryphalogenes Wood 1980 W86
         |    `--*C. euphorbiae W86
         |--Hemicryphalus Schedl 1963 W86
         |    `--*H. argutus [=Eidophelus argutus] W86
         |--Margadillius Hopkins 1915 W86
         |    `--*M. margadilaonis W86
         |--Ptilopodius Hopkins 1915 W86
         |    `--*P. stephegynis W86
         |--Trypophloeus Fairmaire 1868 [=Glyptoderus Eichhoff 1878] W86
         |    |--*T. binodulus [=Bostrichus binodulus, *Glyptoderus binodulus] W86
         |    `--T. populi [=Cryphalus populi] W86
         |--Eidophelus Eichhoff 1875 [incl. Phellodendrophagus Krivolutskaya 1958] W86
         |    |--*E. imitans W86
         |    `--E. elegans [=*Phellodendrophagus elegans] W86
         |--Procryphalus Hopkins 1915 W86
         |    |--P. mucronatus [=Cryphalus mucronatus; incl. *P. populi] W86
         |    `--P. utahensis W86
         |--Cryptocarenus Eggers 1937 [incl. Tachyderes Blackman 1943] W86
         |    |--*C. diadematus W86
         |    `--C. seriatus [incl. *Tachyderes floridensis] W86
         |--Acorthylus Brèthes 1922 [incl. Phacrylus Schedl 1938] W86
         |    |--*A. asperatus W86
         |    `--A. bosqui [=*Phacrylus bosqui] W86
         |--Stephanopodius Schedl 1941 (see below for synonymy) W86
         |    |--*S. dispar [=Stephanoderes dispar] W86
         |    `--S. ghanaensis [=Hypocryphalus ghanaensis, *Cryphalomimetes ghanaensis, *Cryphalomimus ghanaensis] W86
         |--Ernoporus Thomson 1859 (see below for synonymy) W86
         |    |--*E. tiliae [=Bostrichus tiliae; incl. Cryphalus lederi, *Cryphalops lederi] W86
         |    |--E. concentralis [=*Euptilius concentralis] w86
         |    `--E. melodori [=*Stephanorhopalus melodori] W86
         |--Ernoporicus Berger 1917 [incl. Eocryphalus Kurentzov 1941, Ernopocerus Balachowsky 1949] W86
         |    |--*E. spessivtzevi W86
         |    |--E. caucasicus [=Ernoporus caucasicus, *Ernopocerus caucasicus] W86
         |    `--E. semenovi [=*Eocryphalus semenovi] W86
         |--Cosmoderes Eichhoff 1878 (see below for synonymy) W86
         |    |--*C. monilicollis [incl. *Vitaderes luffa (n. n.)] W86
         |    |--C. granulicollis [=*Dendriops granulicollis] W86
         |    `--C. setistriatus [=Cryphalus setistriatus, *Erioschidias setistriatus] W86
         |--Hypocryphalus Hopkins 1915 [incl. Dacryphalus Hopkins 1915] W86
         |    |--*H. rotundus W86
         |    |--H. mangiferae W86
         |    `--H. obesus [=*Dacryphalus obesus] W86
         `--Cryphalus Erichson 1836 (see below for synonymy) W86
              |--*C. asperatus [=Bostrichus asperatus; incl. B. abietus, *Taenioglyptes abietus] W86
              |--C. birosimensis [=*Ernocryphalus birosimensis] W86
              |--C. kolbei (Hagedorn 1912) [=*Allarthrum kolbei] W86
              |--C. lipingensis [=*Acryphalus lipingensis] W86
              |--C. piceus [=*Jugocryphalus piceus] W86
              |--C. ruficollis W86
              |--C. sidneyanus [=Bostrichus sidneyanus, *Pseudocryphalus sidneyanus] W86
              |--C. sylvicola [=Hypothenemus sylvicola; incl. *Ericryphalus henshawi, *Piperius pini] W86
              `--C. walkeri [=*Cryptarthrum walkeri] W86

Cnemonyx Eichhoff 1868 [incl. Ceratolepis Chapuis 1869, Coptodryas Schedl 1948 non Hopkins 1915, Coptosomus Schedl 1952, Loganius Chapuis 1869, Minulus Eggers 1912] W86

Coccotrypes Eichhoff 1878 [incl. Cryphaloides Formenek 1908, Dendrurgus Eggers 1923, Hyphaene Hagedorn 1913 (n. n.), Poecilips Schaufuss 1897, Spermatoplex Hopkins 1915, Thamnurgides Hopkins 1915] W86

Coccotrypes carpophagus [=Bostrichus carpophagus; incl. *Cryphaloides donisthorpei, *Hyphaene guineensis (n. n.)] W86

Cosmoderes Eichhoff 1878 [incl. Dendriops Schedl 1953, Erioschidias Schedl 1938, Vitaderes Beeson 1941 (n. n.)] W86

Cryphalus Erichson 1836 [incl. Acryphalus Tsai & Li 1963, Allarthrum Hagedorn 1912, Cryptarthrum Blandford 1896, Ericryphalus Hopkins 1915, Ernocryphalus Murayama 1958, Jugocryphalus Tsai & Li 1963, Piperius Hopkins 1915, Pseudocryphalus Ferrari 1869, Taenioglyptes Bedel 1888] W86

Ernoporus Thomson 1859 [incl. Cryphalops Reitter 1889, Euptilius Schedl 1940, Stephanorhopalus Hopkins 1915] W86

Glostatus Schedl 1939 [incl. Apoglostatus Schedl 1957, Ctonocryphus Schedl 1941, Paraglostaus Schedl 1964, Rhopalocryphus Nunberg 1967] W86

Lanurgus Eggers 1920 [incl. Landolphianus Schedl 1950, Micraciops Schedl 1953, Pseudohylocurus Nunberg 1961] W86

Pseudothysanoes Blackman 1920 [incl. Aphanocleptus Wood 1960, Bostrichips Schedl 1951, Chalcohyus Blackman 1943, Cryptocleptes Blackman 1920 (preoc.), Cryptulocleptus Wood 1962, Gretschkinia Sokanovskii 1959, Neoglostatus Schedl 1978] W86

Pycnarthrum Eichhoff 1878 [incl. Monebius Hopkins 1914, Nemobius Chapuis 1869 (preoc.), Nomebius Navas 1915] W86

Pycnarthrum hispidum [=Hypoborus hispidus; incl. *P. gracile, *Nemobius lambottei, *Monebius lambottei, *Nomebius lambottei] W86

Scolytoplatypus Schaufuss 1893 [incl. Spongocerus Blandford 1893, Strophionocerus Sampson 1921, Taeniocerus Blandford 1893; Scolytoplatypini, Scolytoplatypodini, Spongocerinae, Taeniocerini] W86

Scolytus Geoffroy 1762 [=Coptogaster Illiger 1807, Eccoptogaster Gyllenhal 1813, Ekkoptogaster Herbst 1793; incl. Archaeoscolytus Butovitsch 1929, Confusoscolytus Tsai, Yin & Huang 1962, Pinetoscolytus Butovitsch 1929, Pygmaeoscolytus Butovitsch 1929, Ruguloscolytus Butovitsch 1929, Scolytochelus Reitter 1913, Spinuloscolytus Butovitsch 1929, Tubuloscolytus Butovitsch 1929] W86

Scolytus multistriatus [=Ips multistriatus, *Scolytochelus multistriatus, *Spinuloscolytus multistriatus] W86

*Scolytus scolytus [=Bostrichus scolytus, *Coptogaster scolytus, *Eccoptogaster scolytus, *Ekkoptogaster scolytus] W86

Stephanopodius Schedl 1941 [incl. Cryphalomimetes Browne 1963, Cryphalomimus Browne 1962 non Eggers 1927] W86

Taphrorychus Eichhoff 1878 [incl. Pseudopoecilips Murayama 1957, Saliciphilus Sokanovskii 1954, Taphroterus Schedl 1965] W86

Tiarophorus Schreiner 1882 [incl. Hypaspistes Hagedorn 1908 (preoc.), Orthaspistes Hagedorn 1909, Pseudothamnurgus Eggers 1912] W86

Webbia Hopkins 1915 [incl. Prowebbia Browne 1962, Pseudowebbia Browne 1961, Xelyborus Schedl 1939 (n. n.)] W86

*Type species of generic name indicated

REFERENCES

[GE05] Grimaldi, D., & M. S. Engel. 2005. Evolution of the Insects. Cambridge University Press: New York.

[JKW03] Jacobs, K., T. Kirisits & M. J. Wingfield. 2003. Taxonomic re-evaluation of three related species of Graphium, based on morphology, ecology and phylogeny. Mycologia 95 (4): 714-727.

[MHG04] Maynard, G. V., J. G. Hamilton & J. F. Grimshaw. 2004. Quarantine—phytosanitary, sanitary and incursion management: an Australian entomological perspective. Australian Journal of Entomology 43 (3): 318-328.

[W86] Wood, S. L. 1986. A reclassification of the genera of Scolytidae (Coleoptera). Great Basin Naturalist Memoirs 10: 1-126.

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