Male Irichohalticella pilosella, from Bouček (1988).

Belongs within: Chalcidoidea.
Contains: Antrocephalus.

The Haltichellinae are a subfamily of the wasp family Chalcididae, characterised by having the apex of the hind tibia almost perpendicularly truncate and with two spurs, only rarely with the ventral corner forming a short spine. Three tribes are recognised within the Haltichellinae (though the analysis by Munro et al., 2011, suggested that Haltichellini may be paraphyletic with regard to Hybothoracini [Tropimeris was not included in this analysis]). Hybothoracini and Tropimeris lack the postmarginal vein in the forewing and have the marginal vein at least slightly removed from the wing margin; Hybothoracini also have the antennae inserted very low, at the level of the mouth(Bouček 1988).

<==Haltichellinae B88
    |--Tropimeris Steffan 1948 [Tropimeridini] B88
    |    |--*T. excavata B88
    |    `--T. monodon Bouček 1958 B88
    `--Haltichellini B88
         |  i. s.: Trichoxenia Kirby 1883 [incl. Megachilochalcis Girault 1915] B88
         |           `--*T. cineraria (Walker 1871) (see below for synonymy) B88
         |         Neostomatoceras Girault 1920 B88
         |           `--*N. chalcidiformis Girault 1920 B88
         |         Nipponohockeria Habu 1960 [=Nippohockeria (l. c.)] B88
         |           `--*N. ishii B88
         |         Uga Girault 1930 [incl. Neotainania Husain & Agarwal 1982] B88
         |           |--*U. colliscutellum (Girault 1922) [=Stomatoceras colliscutellum] B88
         |           `--U. menoni Kerrich 1960 [incl. *Neotainania brevicorpis] B88
         |         Allochalcis Kiefer 1905 [incl. Trichochalcis Kiefer 1905] B88
         |           |--*A. nervosa [incl. *Trichochalcis inermis] B88
         |           |--A. poeta (Girault 1934) [=Trichoxenia poeta] B88
         |           |--A. rubrifemur (Girault 1926) [=Trichoxenia rubrifemur] B88
         |           |--A. shakespearei (Girault 1926) [=Trichoxenia shakespearei] B88
         |           `--A. subfasciata (Walker 1871) (see below for synonymy) B88
         |         Oxycoryphe Kriechbaumer 1894 [incl. Hoozania Masi 1932, Paraspirhina Cameron 1911] B88
         |           |--*O. subaenea B88
         |           |--O. acuta Bouček 1988 B88
         |           |--O. bidens Bouček 1988 B88
         |           |--O. maculipennis [=*Hoozania maculipennis] B88
         |           `--O. nitida [=*Paraspirhina nitida] B88
         |--+--Aspirrhina MH11
         |  `--Haltichella Spinola 1811 MH11, B88 (see below for synonymy)
         |       |--*H. bispinosa [=Chalcis bispinosa; incl. H. rufipes] B88
         |       |--H. macrocera Waterston 1922 B88
         |       `--H. magnidens (Girault 1917) [=Stomatoceras magnidens] B88
         `--+--+--Antrocephalus MH11
            |  `--+--Kriechbaumerella Dalla Torre 1894 MH11, B88 (see below for synonymy)
            |     |    |--*K. palpebrator [=*Coelops palpebrator] B88
            |     |    `--‘Stomatoceras’ magrettii [=*Eucepsis magrettii] B88
            |     `--Hockeria Walker 1834 MH11, B88 (see below for synonymy)
            |          |--H. bifasciata [incl. Chalcis bispinosa Kirby 1833 (preoc.), *H. bispinosa] B88
            |          |--*Afrhockeria’ basilewskyi B88
            |          |--H. dioculata (Girault 1930) [=*Hockerella dioculata] B88
            |          |--‘Halticella’ liberator [=*Stomatoceras liberator] B88
            |          `--‘Stomatoceras’ sulcatiscutellum Girault 1917 G17
            `--Hybothoracini MH11
                 |  i. s.: Nearretocera Girault 1913 [incl. Arretoceroides Girault 1915] B88
                 |           `--*N. johnstoni Girault 1913 [incl. *Arretoceroides feralis Girault 1915] B88
                 |         Xenarretocera Girault 1926 B88
                 |           `--*X. superba Girault 1926 B88
                 |         Notaspidiella Bouček 1988 B88
                 |           |--*N. tirathabae [=Irichohalticella tirathabae] B88
                 |           `--N. frater [=Hybothorax frater] B88
                 |         Nipponochalcidia Habu 1976 B88
                 |           |--*N. kajimurai [=Euchalcidia kajimurai] B88
                 |           `--N. hexcarinata (Girault 1925) B88 (see below for synonymy)
                 |         Steninvreia Bouček 1988 B88
                 |           |--*S. tricarinata (Girault 1927) [=Xenarretocera tricarinata] B88
                 |           |--S. edgari Bouček 1988 B88
                 |           `--S. petiolata Bouček 1988 B88
                 |         Invreia Masi 1927 (see below for synonymy) B88
                 |           |--*I. subaenea B88
                 |           |--‘Euchalcis’ doriae [=Euchalcidia doriae] B88
                 |           |--*Euchalcidia’ elegantula B88
                 |           `--I. soudanensis [=*Hyperchalcidia soudanensis] B88
                 |         Proconura Dodd in Girault 1915 B88
                 |           |--*P. politiventris Dodd & Girault in Girault 1915 B88
                 |           |--P. caryobori B88
                 |           |--P. dexius (Walker 1838) [=Hockeria dexius] B88
                 |           |--P. murrayi (Girault 1929) [=Xenarretocera murrayi] B88
                 |           |--P. parvula (Girault & Dodd in Girault 1915) [=Haltichella parvula] B88
                 |           |--P. seminigripes (Girault 1926) (see below for synonymy) B88
                 |           |--P. shakespearei Girault 1928 B88
                 |           `--P. v-carinata (Girault 1927) [=Xenarretocera v-carinata] B88
                 |--Notaspidium Dalla Torre 1897 MH11, B88 [=Notaspis Walker 1834 non Hermann 1804 B88]
                 |    |--*N. formiciforme [=*Notaspis formiciformis] B88
                 |    `--N. papuanum Bouček 1988 B88
                 `--+--Schwarzella Ashmead 1904 MH11, B88
                    |    `--S. americana MH11
                    `--Irichohalticella Cameron 1912 MH11, B88 [=Irichohaltichella (l. c.) B88]
                         |--*I. pilosella Cameron 1912 B88
                         |--I. atritegula (Girault 1922) [=Nearretocera atritegula] B88
                         |--I. coccinipes Girault 1928 B88
                         |--I. fuscipennis (Dodd & Girault in Girault 1915) [=Nearretocera fuscipennis] B88
                         |--I. multistriata Girault 1927 [=Irichohaltichella (l. c.) multistriata] B88
                         |--I. persimilis (Dodd in Girault 1915) [=Nearretocera persimilis] B88
                         |--I. rubricornis (Dodd in Girault 1915) [=Nearretocera rubricornis] B88
                         |--I. silvae Girault 1927 [=Irichohaltichella (l. c.) silvae] B88
                         `--I. silvifilia Girault 1927 [=Irichohaltichella (l. c.) silvifilia] B88

Allochalcis subfasciata (Walker 1871) [=Halticella subfasciata; incl. Stomatoceras unrubripunctus Girault 1924] B88

Haltichella Spinola 1811 MH11, B88 [=Halticella Stephens 1829 B88; incl. Haltichellodes Steffan 1955 B88, Microchalcis Kieffer 1905 B88]

Hockeria Walker 1834 MH11, B88 [incl. Afrhockeria Steffan 1955 B88, Afrochalcis Schmitz 1946 B88, Centrochalcis Cameron 1905 B88, Hockerella Girault 1930 B88, Hypochalcis Girault 1915 B88, Stomatoceras Kirby 1833 B88]

Invreia Masi 1927 [incl. Euchalcidia Masi 1927, Hyperchalcidia Steffan 1951, Parinvreia Steffan 1951, Peltochalcidia Steffan 1948] B88

Kriechbaumerella Dalla Torre 1894 MH11, B88 [=Coelops Kriechbaumer 1894 non Blyth 1849 B88; incl. Eucepsis Steffan 1953 B88]

Nipponochalcidia hexcarinata (Girault 1925) B88 [=Nearretocera hexcarinata B88, Xenarretocera hexcarinata G26]

Proconura seminigripes (Girault 1926) [=Xenarretocera seminigripes; incl. Nearretocera longipedicel Girault 1926] B88

*Trichoxenia cineraria (Walker 1871) [=Halticella cineraria; incl. *Megachilochalcis auricomosus Girault 1915, M. auricomosa, M. marginatipennis Girault 1926] B88

*Type species of generic name indicated


[B88] Bouček, Z. 1988. Australasian Chalcidoidea (Hymenoptera): A biosystematic revision of genera of fourteen families, with a reclassification of species. CAB International: Wallingford (UK).

[G17] Girault, A. A. 1917. Descriptiones hymenopterorum chalcidoidicarum variorum cum observationibus. V. Privately published (reprinted Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 116-131).

[G26] Girault, A. A. 1926. New pests from Australia. Privately published (reprinted Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 199).

[MH11] Munro, J. B., J. M. Heraty, R. A. Burks, D. Hawks, J. Mottern, A. Cruaud, J.-Y. Rasplus & P. Jansta. 2011. A molecular phylogeny of the Chalcidoidea (Hymenoptera). PLoS One 6 (11): e27023.

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