Sympiesis

Female Sympiesis marylandensis, photographed by Chris Maier.

Belongs within: Eulophini.

Sympiesis is a genus of wasps whose larvae are ectoparasitic on the larvae of leaf-mining Lepidoptera. As currently defined, Sympiesis encompasses a wide range of morphologies. For instance, males may have the antennae either branched or unbranched (Bouček 1988).

See also: Sympiesis at Catalogue of Organisms.

Characters (from Bouček 1988): Antenna with scapes at most only slightly exceeding ocelli, flagellum not distinctly clavate, in male often with three branches, female funicle four-segmented; scutellum not transverse, without sublateral grooves, bare except for four setae; postmarginal vein ending far from wing apex; gaster dorsally collapsing.

Sympiesis Förster 1856 (see below for synonymy) B88
    |--*S. sericeicornis [=Eulophus sericeicornis] B88
    |--S. agromyzae Gahan 1913 GM79
    |    |--S. a. agromyzae G17b
    |    `--S. a. pennsylvanicus Girault 1917 G17b
    |--S. australiensis (Girault 1913) [=*Diaulomella australiensis] B88
    |--S. bimaculatus Crawford 1913 GM79 [incl. S. bimaculatipennis Girault 1917 G25, GM79]
    |--S. boasi (Girault 1913) [=*Sympiesonecremnus boasi] B88
    |--S. campbellensis (Kerrich & Yoshimoto 1964) [=Ardalus campbellensis] B88
    |--S. coloni Girault 1917 G17c
    |--S. consona (Girault 1915) [=*Pardiaulomella consonus] B88
    |--S. cyaneipurpurea (Girault 1915) [=*Diaulomorphella cyaneipurpurea] B88
    |--S. dandoloni Girault 1917 G17a
    |--S. dolichogaster Ashmead 1888 (see below for synonymy) B88
    |--S. felti Crawford 1911 GM79
    |--S. flavipes Ashmead 1886 GM79
    |--S. fucosa (Girault 1915) [=Opheliminus fucosus] B88
    |--S. grotii (Girault 1913) [=*Opheliminus grotii, O. grotiusi Girault 1913] B88
    |--S. guttatipennis Girault 1917 G17b
    |--S. hawaiiensis (Ashmead 1901) [=Ophelinus (l. c. for Ophelimus) hawaiiensis] B88
    |--S. lexingtonensis Girault 1917 G17e
    |--S. longfellowi (Girault 1913) [=Opheliminus longfellowi] B88
    |--S. longiventris (Girault 1913) [=*Pseudopheliminus longiventris, Asympiesiella longiventris] B88
    |--S. miltoni Girault 1917 G17c
    |--S. maculipes Crawford 1913 GM79
    |--S. marilandia Girault 1917 G17d
    |--S. marylandensis Girault 1917 GM79
    |--S. mauiensis (Ashmead 1901) [=Ophelinus (l. c. for Ophelimus) mauiensis] B88
    |--S. nigrifemora Ashmead 1888 GM79
    |--S. nigripes Ashmead 1888 GM79 [incl. S. massasoit Crawford 1913 G25, GM79]
    |--S. pulcherrima (Dodd in Girault 1915) [=Opheliminus pulcherrimus] B88
    |--S. rex Girault 1917 G17a
    |--S. saintpierrei (Girault 1913) (see below for synonymy) B88
    |--S. spadiceipes (Girault 1915) [=*Pardiaulomyia spadiceipes] B88
    |--S. speciosa (Girault & Dodd in Girault 1915) [=*Pronecremnus speciosus] B88
    |--S. stigmatipennis Girault 1917 G17b
    |--S. stigmatus Girault 1917 G17b
    |--S. superba (Girault 1915) [=Asympiesiella superbus] B88
    |--S. tischeriae Ashmead 1888 GM79
    |--S. uroplatae Howard 1885 GM79 [incl. S. unicarinata Ashmead 1894 G17b, GM79]
    `--S. wordsworthi Girault 1917 G17c

Sympiesis Förster 1856 [incl. Aympiesiella Girault 1913, Cladosympiesis Graham 1959, Diaulomella Girault 1913, Diaulomorphella Girault 1915, Moroceras Erdös 1954, Necremnomyia Girault 1913, Opheliminus Girault 1913, Pardiaulomella Girault 1915, Pardiaulomyia Girault 1915, Pronecremnus Girault & Dodd in Girault 1915, Pseudopheliminus Girault 1913, Sympiesonecremnus Girault 1913, Teleogmus Förster 1856] B88

Sympiesis dolichogaster Ashmead 1888 [incl. Asympiesiella nelsonensis var. gracilis Girault 1915, A. india Girault 1916, *A. nelsonensis Girault 1913, Sympiesis nelsonensis Girault 1914] B88

Sympiesis saintpierrei (Girault 1913) [=*Necremnomyia saintpierrei; incl. Sympiesonecremnus aenella Dodd in Girault 1915, N. variscutum Girault 1915] B88

*Type species of generic name indicated

REFERENCES

[B88] Bouček, Z. 1988. Australasian Chalcidoidea (Hymenoptera): A biosystematic revision of genera of fourteen families, with a reclassification of species. CAB International: Wallingford (UK).

[G17a] Girault, A. A. 1917a. New chalcid flies. Privately published (reprinted Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 72-77).

[G17b] Girault, A. A. 1917b. Descriptiones stellarum novarum. Privately published (reprinted Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 80-101).

[G17c] Girault, A. A. 1917c. Descriptiones hymenopterorum chalcidoidicarum variorum cum observationibus. III. Privately published (reprinted Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 102-111).

[G17d] Girault, A. A. 1917d. Chalcidoidea nova Marilandensis. Privately published (reprinted Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 112-113).

[G17e] Girault, A. A. 1917e. Descriptiones hymenopterorum chalcidoidicarum variorum cum observationibus. V. Privately published (reprinted Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 116-131).

[G25] Girault, A. A. 1925. Indications (in new insects) of ruling power and laws in nature. Privately published (reprinted Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 184-186).

[GM79] Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 1-400.