Lyon's pentachaeta Pentachaeta lyonii, from the Pacific Southwest Region USFWS.

Belongs within: Asteroideae.
Contains: Aster, Olearia, Calotis, Erigeron, Vittadinia, Hazardia, Bellidinae, Celmisia, Baccharis, Grindelia, Chrysothamnus, Solidago, Ericameria.

The composite-flowered tribe Astereae includes asters (Aster), goldenrods (Solidago) and related taxa. Members of the group are distinguished by the morphology of the fertile disc-flowers, in which the marginal stigmatic lines do not reach the summits of the style-branches. Within the Astereae, members of the subtribe Asterinae have a well-developed pappus, as opposed to the reduced pappus of the Bellidinae (Black & Robertson 1965). Pleurophyllum species are robust perennial herbs found on the Subantarctic islands of New Zealand (Allan 1961). Conyza, the fleabanes, are erect herbs or shrubs growing up to two metres in height, some species of which are notable as invasive weeds. Minuria is an Australian genus of mostly small undershrubs with flower-heads borne on short peduncles; the receptacle is naked and ray-flowers are borne in two or more rows (Black & Robertson 1965).

Characters (from Black & Robertson 1965): Style-branches of fertile disc-flowers flattish, with two marginal papillose stigmatic lines along lower part of branches and long or short non-stigmatic appendage at summit bearing short collecting hairs on outside; flower-heads heterogamous radiate; receptacle usually naked; pappus consisting of bristles, awns or scales, rarely absent; leaves alternate.

    |--Doellingeria umbellata RU03
    |--+--Gundlachia corymbosa RU03
    |  `--Xylothamia RU03
    |       |--X. parrasana [=Haplopappus parrasanus] RU03
    |       |--‘Haplopappus’ sonoriensis RU03
    |       |--X. triantha RU03
    |       `--X. truncata RU03
    `--+--+--‘Chrysothamnus’ gramineus RU03 [incl. Petradoria discoidea H93]
       |  `--+--Sericocarpus tortifolius RU03
       |     `--+--Chrysothamnus RU03
       |        `--+--Chrysoma pauciflosculosa RU03
       |           `--Solidago RU03
       `--+--Ericameria RU03
          `--+--+--Rigiopappus leptocladus RU03
             |  `--Tracyina rostrata RU03
             `--Pentachaeta RU03
                  |--P. alsinoides [=Chaetopappa alsinoides] H93
                  |--P. aurea [=Chaetopappa aurea] H93
                  |--P. bellidiflora [=Chaetopappa bellidiflora] H93
                  |--P. exilis [=Chaetopappa exilis] H93
                  |    |--P. e. ssp. exilis H93
                  |    `--P. e. ssp. aeolica H93
                  |--P. fragilis [=Chaetopappa fragilis] H93
                  `--P. lyonii [=Chaetopappa lyonii] H93

Astereae incertae sedis:
  Asterinae BR65
    |--Aster BR65
    |--Achnophora tatei BR65
    |--Olearia BR65
    |--Calotis BR65
    |--Erigeron BR65
    |--Vittadinia BR65
    |--Podocoma BR65
    |    |--P. cuneifolia BR65
    |    `--P. nana BR65
    `--Minuria BR65
         |--M. annua [=Minuriella annua] BR65
         |--M. cunninghamii BR65
         |--M. denticulata BR65
         |--M. integerrima BR65
         |--M. leptophylla BR65
         |--M. macrorhiza LK14
         |--M. rigida BR65
         `--M. suaedifolia BR65
  Hazardia RU03
  Stenactis annua [=Aster annuus] C55
  Haplopappus M-FR08
    |--H. coronopifolius D03
    |--H. resinosus PS96
    |--H. rubiginosus [=Sideranthus rubiginosus] G05
    |--H. spinulosus A72
    |--H. tenuisectus LW70
    `--H. wigginsii A72
  Bellidinae BR65
  Pleurophyllum Hooker 1844 A61
    |--P. criniferum Hooker 1844 A61 (see below for synonymy)
    |--P. hookeri Buchan. 1884 A61 [incl. P. gilliesianum Kirk 1891 C06, P. hookerianum Kirk 1891 C06]
    `--P. speciosum Hooker 1844 A61 [incl. Aster pleurophyllum Muell. 1865 ME70]
  Celmisia A61
  Pachystegia Cheeseman 1925 A61
    `--P. insignis (Hooker) Cheeseman 1925 (see below for synonymy) A61
         |--P. i. var. insignis A61
         `--P. i. var. minor Cheeseman 1925 [=Olearia insignis var. minor Cheeseman 1916] A61
  Baccharis N10
  Grindelia S06
  Gutierrezia S06
    |--G. californica [incl. G. bracteata] H93
    |--G. lacida BT72
    |--G. microcephala H93
    `--G. sarothrae CL86
  Hysterionica S06
  Nardophyllum S06
  Chiliophyllum S06
  Egletes S06
  Leucopsidium S06
  Sommerfeldtia S06
  Chiliotrichium diffusum S06, CL85
  Conyza S06
    |--C. albida GK00
    |--C. apurensis J87
    |--C. bilboana H93
    |--C. bonariensis Y98
    |--C. canadensis AGF98
    |    |--C. c. var. canadensis J87
    |    `--C. c. var. pusilla J87
    |--C. coulteri H93
    `--C. floribunda H93
  Chrysocoma puberula S06, CV06
  Heterothalamus nivalis S06, D03

Pachystegia insignis (Hooker) Cheeseman 1925 [=Olearia insignis Hooker 1855; incl. O. marginata Colenso 1883] A61

Pleurophyllum criniferum Hooker 1844 A61 [=Aster criniferum (Hooker) Muell. 1865 ME70; incl. P. hombronii Decne 1853 A61, Albinia oresigenesa Homb. & Jacq. ex Decne 1853 A61, P. oresigenesum C06]

*Type species of generic name indicated


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[N10] Norrbom, A. L. 2010. Tephritidae (fruit flies, moscas de frutas). In: Brown, B. V., A. Borkent, J. M. Cumming, D. M. Wood, N. E. Woodley & M. A. Zumbado (eds) Manual of Central American Diptera vol. 2 pp. 909–954. NRC Research Press: Ottawa.

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Last updated: 21 December 2019.

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