Astereae

Lyon's pentachaeta Pentachaeta lyonii, from the Pacific Southwest Region USFWS.

Belongs within: Asteroideae.
Contains: Aster, Olearia, Calotis, Erigeron, Vittadinia, Hazardia, Bellidinae, Celmisia, Baccharis, Grindelia, Chrysothamnus, Solidago, Ericameria.

The composite-flowered tribe Astereae includes asters (Aster), goldenrods (Solidago) and related taxa. Members of the group are distinguished by the morphology of the fertile disc-flowers, in which the marginal stigmatic lines do not reach the summits of the style-branches. Within the Astereae, members of the subtribe Asterinae have a well-developed pappus, as opposed to the reduced pappus of the Bellidinae (Black & Robertson 1965). Pleurophyllum species are robust perennial herbs found on the Subantarctic islands of New Zealand (Allan 1961). Conyza, the fleabanes, are erect herbs or shrubs growing up to two metres in height, some species of which are notable as invasive weeds. Minuria is an Australian genus of mostly small undershrubs with flower-heads borne on short peduncles; the receptacle is naked and ray-flowers are borne in two or more rows (Black & Robertson 1965).

Characters (from Black & Robertson 1965): Style-branches of fertile disc-flowers flattish, with two marginal papillose stigmatic lines along lower part of branches and long or short non-stigmatic appendage at summit bearing short collecting hairs on outside; flower-heads heterogamous radiate; receptacle usually naked; pappus consisting of bristles, awns or scales, rarely absent; leaves alternate.

<==Astereae
    |--Doellingeria umbellata RU03
    |--+--Gundlachia corymbosa RU03
    |  `--Xylothamia RU03
    |       |--X. parrasana [=Haplopappus parrasanus] RU03
    |       |--‘Haplopappus’ sonoriensis RU03
    |       |--X. triantha RU03
    |       `--X. truncata RU03
    `--+--+--‘Chrysothamnus’ gramineus RU03 [incl. Petradoria discoidea H93]
       |  `--+--Sericocarpus tortifolius RU03
       |     `--+--Chrysothamnus RU03
       |        `--+--Chrysoma pauciflosculosa RU03
       |           `--Solidago RU03
       `--+--Ericameria RU03
          `--+--+--Rigiopappus leptocladus RU03
             |  `--Tracyina rostrata RU03
             `--Pentachaeta RU03
                  |--P. alsinoides [=Chaetopappa alsinoides] H93
                  |--P. aurea [=Chaetopappa aurea] H93
                  |--P. bellidiflora [=Chaetopappa bellidiflora] H93
                  |--P. exilis [=Chaetopappa exilis] H93
                  |    |--P. e. ssp. exilis H93
                  |    `--P. e. ssp. aeolica H93
                  |--P. fragilis [=Chaetopappa fragilis] H93
                  `--P. lyonii [=Chaetopappa lyonii] H93

Astereae incertae sedis:
  Asterinae BR65
    |--Aster BR65
    |--Achnophora tatei BR65
    |--Olearia BR65
    |--Calotis BR65
    |--Erigeron BR65
    |--Vittadinia BR65
    |--Podocoma BR65
    |    |--P. cuneifolia BR65
    |    `--P. nana BR65
    `--Minuria BR65
         |--M. annua [=Minuriella annua] BR65
         |--M. cunninghamii BR65
         |--M. denticulata BR65
         |--M. integerrima BR65
         |--M. leptophylla BR65
         |--M. macrorhiza LK14
         |--M. rigida BR65
         `--M. suaedifolia BR65
  Hazardia RU03
  Stenactis annua [=Aster annuus] C55
  Haplopappus M-FR08
    |--H. coronopifolius D03
    |--H. resinosus PS96
    |--H. rubiginosus [=Sideranthus rubiginosus] G05
    |--H. spinulosus A72
    |--H. tenuisectus LW70
    `--H. wigginsii A72
  Bellidinae BR65
  Pleurophyllum Hooker 1844 A61
    |--P. criniferum Hooker 1844 A61 (see below for synonymy)
    |--P. hookeri Buchan. 1884 A61 [incl. P. gilliesianum Kirk 1891 C06, P. hookerianum Kirk 1891 C06]
    `--P. speciosum Hooker 1844 A61 [incl. Aster pleurophyllum Muell. 1865 ME70]
  Celmisia A61
  Pachystegia Cheeseman 1925 A61
    `--P. insignis (Hooker) Cheeseman 1925 (see below for synonymy) A61
         |--P. i. var. insignis A61
         `--P. i. var. minor Cheeseman 1925 [=Olearia insignis var. minor Cheeseman 1916] A61
  Baccharis N10
  Grindelia S06
  Gutierrezia S06
    |--G. californica [incl. G. bracteata] H93
    |--G. lacida BT72
    |--G. microcephala H93
    `--G. sarothrae CL86
  Hysterionica S06
  Nardophyllum S06
  Chiliophyllum S06
  Egletes S06
  Leucopsidium S06
  Sommerfeldtia S06
  Chiliotrichium diffusum S06, CL85
  Conyza S06
    |--C. albida GK00
    |--C. apurensis J87
    |--C. bilboana H93
    |--C. bonariensis Y98
    |--C. canadensis AGF98
    |    |--C. c. var. canadensis J87
    |    `--C. c. var. pusilla J87
    |--C. coulteri H93
    `--C. floribunda H93
  Chrysocoma puberula S06, CV06
  Heterothalamus nivalis S06, D03

Pachystegia insignis (Hooker) Cheeseman 1925 [=Olearia insignis Hooker 1855; incl. O. marginata Colenso 1883] A61

Pleurophyllum criniferum Hooker 1844 A61 [=Aster criniferum (Hooker) Muell. 1865 ME70; incl. P. hombronii Decne 1853 A61, Albinia oresigenesa Homb. & Jacq. ex Decne 1853 A61, P. oresigenesum C06]

*Type species of generic name indicated

REFERENCES

[AGF98] Abd El-Ghani, M. M., & A. G. Fahmy. 1998. Composition of and changes in the spontaneous flora of Feiran Oasis, S Sinai, Egypt, in the last 60 years. Willdenowia 28: 123–134.

[A61] Allan, H. H. 1961. Flora of New Zealand vol. 1. Indigenous Tracheophyta: Psilopsida, Lycopsida, Filicopsida, Gymnospermae, Dicotyledones. R. E. Owen, Government Printer: Wellington (New Zealand).

[A72] Atchley, W. R. 1972. The chromosome karyotype in estimation of lineage relationships. Systematic Zoology 21 (2): 199–209.

[BT72] Baker, E. W., & D. M. Tuttle. 1972. New species and further notes on the Tetranychoidea mostly from the southwestern United States (Acarina: Tetranychidae and Tenuipalpidae). Smithsonian Contributions to Zoology 116: 1–37.

[BR65] Black, J. M., & E. L. Robertson. 1965. Flora of South Australia. Part IV. Oleaceae–Compositae. W. L. Hawes, Government Printer: Adelaide.

[C55] Candolle, A. de. 1855. Géographie Botanique Raisonée: Ou exposition des faits principaux et des lois concernant la distribution géographique des plantes de l’époque actuelle vol. 2. Librairie de Victor Masson: Paris.

[C06] Cheeseman, T. F. 1906. Manual of the New Zealand Flora. John Mackay, Government Printer: Wellington.

[CL85] Cokendolpher, J. C., & D. Lanfranco L. 1985. Opiliones from the Cape Horn Archipelago: new southern records for harvestmen. Journal of Arachnology 13: 311–319.

[CL86] Collins, E. I., & R. W. Lichvar. 1986. Vegetation inventory of current and historic black-footed ferret habitat in Wyoming. Great Basin Naturalist Memoirs 8: 85–93.

[CV06] Craven, P., & P. Vorster. 2006. Patterns of plant diversity and endemism in Namibia. Bothalia 36 (2): 175–189.

[D03] Dusén, P. 1903. The vegetation of western Patagonia. In: Scott, W. B. (ed.) Reports of the Princeton University Expeditions to Patagonia, 1896–1899 vol. 8. Botany pp. 1–34. The University: Princeton (New Jersey).

[GK00] Gibson, N., & G. J. Keighery. 2000. Flora and vegetation of the Byenup-Muir reserve system, south-west Western Australia. CALMScience 3 (3): 323–402.

[G05] Grissell, E. E. 2005. A review of North American species of Microdontomerus Crawford (Torymidae: Hymenoptera). Journal of Hymenoptera Research 14 (1): 22–65.

[H93] Hickman, J. C. (ed.) 1993. The Jepson Manual: Higher Plants of California. University of California Press: Berkeley (California).

[J87] Judd, W. S. 1987. Floristic study of Morne La Visite and Pic Macaya National Parks, Haiti. Bulletin of the Florida State Museum—Biological Sciences 32 (1): 1–136.

[LW70] Lowe, C. H., J. W. Wright, C. J. Cole & R. L. Bezy. 1970. Natural hybridization between the teiid lizards Cnemidophorus sonorae (parthenogenetic) and Cnemidophorus tigris (bisexual). Systematic Zoology 19 (2): 114–127.

[LK14] Lyons, M. N., G. J. Keighery, L. A. Gibson & T. Handasyde. 2014. Flora and vegetation communities of selected islands off the Kimberley coast of Western Australia. Records of the Western Australian Museum Supplement 81: 205–244.

[ME70] Moore, L. B., & E. Edgar. 1970. Flora of New Zealand vol. 2. Indigenous Tracheophyta: Monocotyledones except Gramineae. A. R. Shearer, Government Printer: Wellington (New Zealand).

[M-FR08] Morais-Filho, J. C., & G. Q. Romero. 2008. Microhabitat use by Peucetia flava (Oxyopidae) on the glandular plant Rhyncanthera dichotoma (Melastomataceae). Journal of Arachnology 36 (2): 374–378.

[N10] Norrbom, A. L. 2010. Tephritidae (fruit flies, moscas de frutas). In: Brown, B. V., A. Borkent, J. M. Cumming, D. M. Wood, N. E. Woodley & M. A. Zumbado (eds) Manual of Central American Diptera vol. 2 pp. 909–954. NRC Research Press: Ottawa.

[PS96] Pike, K. S., P. Starý, R. Miller, D. Allison, L. Boydston, G. Graf & T. Miller. 1996. New species and host records of aphid parasitoids (Hymenoptera: Braconidae: Aphidiinae) from the Pacific northwest, U.S.A. Proceedings of the Entomological Society of Washington 98 (3): 570–591.

[RU03] Roberts, R. P., & L. E. Urbatsch. 2003. Molecular phylogeny of Ericameria (Asteraceae, Astereae) based on nuclear ribosomal 3´ ETS and ITS sequence data. Taxon 52: 209–228.

[S06] Stuckert, T. 1906. Distribución geográfica de la flora Argentina. Géneros de la familia de las compuestas. Anales del Museo Nacional de Buenos Aires, serie 3, 6: 303–309.

[Y98] Yannitsaros, A. 1998. Additions to the flora of Kithira (Greece) I. Willdenowia 28: 77–94.

Last updated: 21 December 2019.