Coelambus impressopunctatus, photographed by Trevor and Dilys Pendleton.

Belongs within: Dytiscidae.
Contains: Hydroporini, Bidessini, Hyphydrini.

The Hydroporinae is a clade of diving beetles distinguished from others in the family Dytiscidae by their pseudotetramerous pro- and mesotarsi (the fourth tarsal segment is much reduced, so that it appears absent). Hydroporines are divided between several tribes, with Laccornis supported as the sister group of all other hydroporines by its retention of laterotergites in the female genitalia (Miller 2001). The Methlini have the metafemur reaching the metacoxal lobe, and terga VII and VIII modified with tergum VIII posteriorly acute with dorsal and ventral lobes, the dorsal lobe forming a trifid structure.

Characters (from Miller 2001): Prosternum and prosternal process strongly declivitous; pro- and mesotarsi pseudotetramerous in both sexes; scutellum usually not externally visible; median lobe usually bilaterally symmetrical; larva with clypeal margin produced anteriorly.

    |  i. s.: Carabhydrus Watts 1978 LM87 [Carabhydrini M01]
    |           |--*C. monteithi Watts 1978 LM87
    |           |--C. andreas Zwick 1981 LM87
    |           |--C. niger Watts 1978 LM87
    |           |--C. plicatus Watts 1978 LM87
    |           `--C. stephanieae Watts et al. 2007 WM13
    |         Laccornellus M01
    |         Canthyporus M01
    |--Laccornis [Laccornini] M01
    |    |--L. difformis M01
    |    |--L. oblongus M01
    |    `--L. pacificus M01
    `--+--Methlini M01
       |    |--Methles cribratellus M01
       |    `--Celina M01
       |         |--C. aculeata R89
       |         |--C. angustata R89
       |         `--C. hubbelli M01
       `--+--Hydroporini M01
          |--Bidessini M01
          `--Hygrotini M01
               |--Coelambus M01
               |    |--C. impressopunctatus [=Hygrotus (Coelambus) impressopunctatus] M01
               |    `--+--C. nigrescens [=Hygrotus (Coelambus) nigrescens] M01
               |       |--C. patruelis [=Hygrotus (Coelambus) patruelis] M01
               |       |--C. sellatus [=Hygrotus (Coelambus) sellatus] M01
               |       `--C. unguicularis [=Hygrotus (Coelambus) unguicularis] M01
               `--+--Hyphydrini M01
                  `--+--+--Herophydrus M01
                     |  `--Hygrotus M01
                     |       |  i. s.: H. quinquelineatus C01
                     |       |--H. laccophilinus M01
                     |       `--+--H. inaequalis M01
                     |          |--H. sayi M01
                     |          `--H. versicolor M01
                     `--Hydrovatini M01
                          |--Queda M01
                          |    |--Q. hydrovatoides M01
                          |    `--Q. youngi M01
                          `--Hydrovatus Motschulsky 1855 LM87
                               |--*H. castaneus Motschulsky 1855 LM87
                               |--H. armstrongi Watts 1978 LM87
                               |--H. crassulus M01
                               |--H. fasciatus Sharp 1882 LM87
                               |--H. nigrita Sharp 1882 LM87
                               |--H. opacus Sharp 1882 LM87
                               |--H. ovalis Sharp 1882 LM87
                               |--H. parallelus Sharp 1882 LM87
                               |--H. pustulatus M01
                               `--H. rufoniger (Clark 1863) [=Hyphydrus rufoniger; incl. Hyd. politus Sharp 1882] LM87

*Type species of generic name indicated


[C01] Csiki, E. 1901. Bogarak [Coleopteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 75–120. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

[M01] Miller, K. B. 2001. On the phylogeny of the Dytiscidae (Insecta: Coleoptera) with emphasis on the morphology of the female reproductive system. Insect Systematics and Evolution 32: 45–92.

[R89] Régimbart, M. 1889. Enumération des Haliplidae, Dytiscidae et Gyrinidae recueillis par M.r le Prof. L. Balzan dans l’Amérique méridionale et description de quelques autres espèces voisines. Annali del Museo Civico di Storia Naturale di Genova, Serie 2a, 7: 256–268.

[WM13] Watts, C. H. S., & J. McRae. 2013. Limbodessus bennetti sp. nov., the first stygobitic Dytiscidae (Coleoptera) from the Pilbara region of Western Australia. Records of the Western Australian Museum 28 (2): 141–143.

Last updated: 30 January 2018.

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