Marsh bedstraw Galium palustre agg., from here.

Belongs within: Paederieae.

Galium, bedstraw, is a cosmopolitan genus of small herbs. Many species in the genus have noticeably 'sticky' stems with small recurved hairs. These include the cleavers Galium aparine, which is native to temperate Eurasia but has become naturalised in many other parts of the world.

Characters (from Black & Robertson 1965): Weak herbs with quadrangular stems, apparently whorled sessile or subsessile leaves and usually whitish floweres. Calyx absent; corolla rotate, with scarcely any tube and four spreading lobes; fruit small, dry.

<==Galium Linnaeus 1753 A61
    |--G. sect. Aparinoides (Jordan) Gren. 1851 B06
    |    |--G. innocuum B06
    |    |--G. karakulense B06
    |    |--G. palustre B06
    |    |--G. shanense Bhattacharjee 2006 B06
    |    `--G. trifidum B06
    |--G. sect. Asperigalium Bhattacharjee 2006 B06
    |    |--G. (sect. *Asperigalium) asperifolium B06
    |    |--G. acutum B06
    |    |--G. campylotrichum B06
    |    |--G. confertum B06
    |    |--G. duthiei Bhattacharjee 2006 B06
    |    |    |--G. d. var. duthiei B06
    |    |    `--G. d. var. glabriusculum Bhattacharjee 2006 B06
    |    |--G. himachalense B06
    |    |--G. megacyttarion B06
    |    |--G. pilosissimum B06
    |    |--G. poluninii B06
    |    |--G. pseudoaserifolium B06
    |    |--G. subfalcatum B06
    |    |--G. tetraphyllum B06
    |    `--G. vicaryii B06
    |--G. sect. Hirtiflorum Bhattacharjee 2006 B06
    |    |--G. (sect *Hirtiflorum) hirtiflorum B06
    |    `--G. subtrinervum B06
    `--G. sect. Trachygalium B06
         |--G. ceratophylloides B06
         `--G. kaganense Bhattacharjee 2006 B06

Galium incertae sedis:
  G. album BM00
  G. aparine BR65
  G. australe BR65
  G. brevifolium PT98
    |--G. b. ssp. brevifolium PT98
    `--G. b. ssp. insulare PT98
  G. brevipes J87
  G. capitatum Bory & Chaubard 1832 PL04
  G. corrudoefolium C74
  G. cruciata TG88
  G. divaricatum BR65
  G. exile O88
  G. fuegianum D03
  G. gaudichaudii [incl. G. ciliare] BR65
    |--G. g. var. gaudichaudii BR65
    `--G. g. var. muriculatum BR65
  G. latifolium EBS98
  G. mollugo Linnaeus 1753 PL04
  G. murale BR65
  G. perpusillum (Hooker) Allan 1961 (see below for synonymy) A61
  G. propinquum Cunn. 1839 (see below for synonymy) A61
  G. pumilum C55a
  G. pusillosetosum O88
  G. relbum D03
  G. saccharatum C55b
  G. saxatile PAE98
  G. serpylloides O88
  G. setaceum PT98
  G. spurium C55b
  G. tenerum [incl. G. aparine var. minor] BR65
  G. tenuicaule Cunn. 1839 [incl. G. triloba Colenso 1888] A61
  G. tricorne BR65
  G. vaillantii C55b
  G. verum BBM02

Nomen nudum: Galium (sect. Hirtiflorum) falconeri B06

Galium perpusillum (Hooker) Allan 1961 [=Asperula perpusilla Hooker 1853; incl. A. aristifera Colenso 1889, A. fragrantissima Armstrong 1882] A61

Galium propinquum Cunn. 1839 [incl. G. propinquum var. elongata Hooker 1853, G. erythrocaulon Colenso 1884, G. propinquum var. glabrata Hooker 1853, G. propinquum var. hispidula Hooker 1853, G. umbrosum Sol. in Forster 1786 (n. n.)] A61

*Type species of generic name indicated


[A61] Allan, H. H. 1961. Flora of New Zealand vol. 1. Indigenous Tracheophyta: Psilopsida, Lycopsida, Filicopsida, Gymnospermae, Dicotyledones. R. E. Owen, Government Printer: Wellington (New Zealand).

[B06] Bhattacharjee, R. 2006. New taxa of Galium L. (Rubiaceae) from the Indian subcontinent. Bulletin of the Botanical Survey of India 48 (1–4): 59–72.

[BR65] Black, J. M., & E. L. Robertson. 1965. Flora of South Australia. Part IV. Oleaceae–Compositae. W. L. Hawes, Government Printer: Adelaide.

[BBM02] Bonte, D., L. Baert & J.-P. Maelfait. 2002. Spider assemblage structure and stability in a heterogeneous coastal dune system (Belgium). Journal of Arachnology 30 (2): 331–343.

[BM00] Bremer, B., & J.-F. Manen. 2000. Phylogeny and classification of the subfamily Rubioideae (Rubiaceae). Plant Systematics and Evolution 225: 43–72.

[C55a] Candolle, A. de. 1855a. Géographie Botanique Raisonée: Ou exposition des faits principaux et des lois concernant la distribution géographique des plantes de l’époque actuelle vol. 1. Librairie de Victor Masson: Paris.

[C55b] Candolle, A. de. 1855b. Géographie Botanique Raisonée: Ou exposition des faits principaux et des lois concernant la distribution géographique des plantes de l’époque actuelle vol. 2. Librairie de Victor Masson: Paris.

[C74] Coineau, Y. 1974. Éléments pour une monographie morphologique, écologique et biologique des Caeculidae (Acariens). Mémoires du Muséum National d'Histoire Naturelle, nouvelle série, Série A, Zoologie 81: 1–299, 224 pls.

[D03] Dusén, P. 1903. The vegetation of western Patagonia. In: Scott, W. B. (ed.) Reports of the Princeton University Expeditions to Patagonia, 1896–1899 vol. 8. Botany pp. 1–34. The University: Princeton (New Jersey).

[EBS98] Elliott, K. J., L. R. Boring & W. T. Swank. 1998. Changes in vegetation structure and diversity after grass-to-forest succession in a southern Appalachian watershed. American Midland Naturalist 140: 219–232.

[J87] Judd, W. S. 1987. Floristic study of Morne La Visite and Pic Macaya National Parks, Haiti. Bulletin of the Florida State Museum—Biological Sciences 32 (1): 1–136.

[O88] Ohba, H. 1988. The alpine flora of the Nepal Himalayas: an introductory note. In: Ohba, H., & S. B. Malla (eds)The Himalayan Plants vol. 1.  The University Museum, University of Tokyo, Bulletin 31: 19–46.

[PAE98] Pakeman, R. J., J. P. Attwood & J. Engelen. 1998. Sources of plants colonizing experimentally disturbed patches in an acidic grassland, in eastern England. Journal of Ecology 86: 1032–1041.

[PT98] Panitsa, M., & D. Tzanoudakis. 1998. Contribution to the study of the Greek flora: flora and vegetation of the E Aegean islands Agathonisi and Pharmakonisi. Willdenowia 28: 95–116.

[PL04] Pohl, G., & I. Lenski. 2004. Zur Verbreitung und Vergesellschaftung von Pennisetum orientale Rich. in Nordeuböa (Griechenland) (Poaceae, Paniceae). Senckenbergiana Biologica 83 (2): 209–223.

[TG88] Tepfer, D., A. Goldmann, N. Pamboukdjian, M. Maille, A. Lepingle, D. Chevalier, J. Dénarié & C. Rosenberg. 1988. A plasmid of Rhizobium meliloti 41 encodes catabolism of two compounds from root exudate of Calystegium sepium. Journal of Bacteriology 170 (3): 1153–1161.

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