Styphelioideae

Dracophyllum muscoides, photographed by Jardin Lautaret.


Belongs within: Ericaceae.
Contains: Styphelieae.

The Styphelioideae are a clade of heath-like plants previously recognised as the family Epacridaceae, but now regarded as a derived subgroup of the Ericaceae. Styphelioids are distinguished by their lignified leaf cells.

Characters (from Kron et al. 2002): Shrubs, small trees or climbers. Leaves evergreen, alternate, usually entire; veins often more or less palmate, parallel, or reduced to one. Leaf epidermal cells lignified; bracts and bracteoles scarious. Inflorescence variable, axillary or terminal. Flowers tetra- or pentamerous, actinomorphic. Calyx persistent; corolla persistent, sympetalous, usually tubular or campanulate, lobes small to large. Stamens 2, 4 or 5, usually epipetalous; anthers bisporangiate, inverting early in development, mostly monothecal, dehiscing by longitudinal slits, lacking a fibrous endothecium and disintegration tissue. Pollen in monads, tetrads, or shed as reduced tetrads with one or more cells aborting, without viscin threads. Ovary 1-11-locular, with axile or apical placentation, superior; style impressed or not; stigma capitate or lobed. Fruit drupaceous or a loculicidal capsule; seeds with variable testa; embryo with two cotyledons.

<==Styphelioideae [Epacridaceae, Epacridoideae]
    |--Prionoteae KJ02
    |    |--Prionotes cerinthoides KJ02
    |    `--Lebetanthus KJ02
    |         |--L. americanus D03
    |         `--L. myrsinites KJ02
    `--+--Styphelieae KJ02
       |--Archeria [Archerieae] KJ02
       |    |--A. comberi KJ02
       |    `--A. traversii W91
       |--Oligarrheneae KJ02
       |    |--Oligarrhena micrantha KJ02
       |    `--Needhamiella pumilio KJ02
       |--Cosmelieae KJ02
       |    |--Cosmelia rubra KJ02
       |    |--Sprengelia KJ02
       |    |    |--S. incarnata KJ02
       |    |    `--S. monticola KJ02
       |    `--Andersonia KJ02
       |         |--A. caerulea GK00
       |         |--A. parvifolia OS04
       |         `--A. sprengelioides KJ02
       |--Epacrideae [Epacridinae] KJ02
       |    |--Rupicola sprengeloides KJ02
       |    |--Lysinema ciliatum KJ02
       |    |--Budawangia KJ02
       |    |--Woollsia pungens KJ02
       |    `--Epacris KJ02
       |         |--E. alpina W91
       |         |--E. impressa KJ02
       |         |--E. longiflora KJ02
       |         `--E. pauciflora A27
       `--Richeeae KJ02
            |--Richea [Richeoideae] KJ02
            |    |--R. dracophylla KJ02
            |    `--R. pandanifolia KJ02
            |--Sphenotoma KJ02
            |    |--S. capitatum GK00
            |    |--S. dracophylloides KJ02
            |    `--S. gracile GK00
            `--Dracophyllum KJ02
                 |  i. s.: D. arboreum A27
                 |         D. latifolium A27
                 |         D. longifolium KJ02
                 |         D. milliganii KJ02
                 |         D. paludosum A27
                 |         D. politum W91
                 |         D. pubescens W91
                 |         D. recurvum A27
                 |         D. rosmarinifolium A27
                 |         D. scoparium A27
                 |         D. sinclairii MM04
                 |         D. traversii W91
                 |         D. uniflorum H03
                 |         D. urvilleanum A27
                 |--D. (subg. Cordophyllum) involucratum H03
                 `--D. (subg. Oreothamnus) H03
                      |--D. minimum H03
                      `--D. muscoides H03

*Type species of generic name indicated

REFERENCES

[A27] Andersen, J. C. 1927. Popular names of New Zealand plants. Transactions and Proceedings of the New Zealand Institute 57: 905-977.

[D03] Dus̩n, P. 1903. The vegetation of western Patagonia. In Reports of the Princeton University Expeditions to Patagonia, 1896-1899, vol. 8 РBotany (W. B. Scott, ed.) pp. 1-34. The University: Princeton (New Jersey).

[GK00] Gibson, N., & G. J. Keighery. 2000. Flora and vegetation of the Byenup-Muir reserve system, south-west Western Australia. CALMScience 3 (3): 323-402.

[H03] Heads, M. 2003. Ericaceae in Malesia: Vicariance biogeography, terrane tectonics and ecology. Telopea 10 (1): 311-449.

[KJ02] Kron, K. A., W. S. Judd, P. F. Stevens, D. M. Crayn, A. A. Anderberg, P. A. Gadek, C. J. Quinn & J. L. Luteyn. 2002. Phylogenetic classification of Ericaceae: Molecular and morphological evidence. Botanical Review 68: 335-423.

[MM04] Martin, N. A., & L. A. Mound. 2004. Host plants for some New Zealand thrips (Thysanoptera: Terebrantia). New Zealand Entomologist 27: 119-123.

[OS04] Obbens, F. J., & L. W. Sage. 2004. Vegetation and flora of a diverse upland remnant of the Western Australian wheatbelt (Nature Reserve A21064). Journal of the Royal Society of Western Australia 87 (1): 19-28.

[W91] Williams, P. A. 1991. Subalpine and alpine vegetation of granite ranges in western Nelson, New Zealand. New Zealand Journal of Botany 29: 317-330.

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