Goniaceritae

Specimen of Plagiophorus, from here.

Belongs within: Staphylinidae.
Contains: Natypleurina, Reichenbachia.

The Goniaceritae are a group of pselaphine beetles with clearly separated hind coxae, short trochanters, three-segmented tarsi with the first segment smaller than the other two, asymmetrical or single tarsal claws, and generally complete parameres in the male genitalia (Nomura & Sakchoowong).

<==Goniaceritae
    |--Iniocyphini NG05
    |    |--Natypleurina NG05
    |    `--Sunorfa [Iniocyphina] NG05
    |         `--S. clavata (Raffray 1895) NG05
    |--Proterini NG05
    |    |--Mechanicus cribratus Raffray 1893 NG05
    |    |--Proterus punctatus NG05
    |    `--Bythinoderes grabowskyi Reitter 1883 NG05
    |--Atychodea [Tychini] NG05
    |    |--A. lenticornis Reitter 1883 NG05
    |    |--A. raffrayi Reitter 1883 NG05
    |    |--A. simoniana Reitter 1883 NG05
    |    `--A. singularis Reitter 1883 NG05
    |--Arnylliini NG05
    |    |--Awas Löbl 1994 NG05
    |    |    |--*A. giraffa Löbl 1994 NG05
    |    |    `--A. rajah Nomura 2004 NG05
    |    `--Harmophorus Motschulsky 1851 [incl. Arnyllium Reitter 1884] NG05
    |         |--H. ciliatum (Raffray 1894) NG05
    |         |--H. cultratum (Raffray 1894) NG05
    |         |--H. ensipes (Reitter 1883) NG05
    |         |--H. parviceps (Reitter 1883) NG05
    |         `--H. pectinatum (Reitter 1883) NG05
    |--Brachyglutina [Brachyglutini, Pselaptina] NG05
    |    |--Ephymata mucronata (Raffray 1895) NG05
    |    |--Trissemus NG05
    |    |--*Noduliceps latifrons NG05
    |    |--Reichenbachia NG05
    |    |--Atenisodus Raffray 1904 NG06
    |    |    `--A. longicornis (Raffray 1895) NG05
    |    |--Eupines King 1866 NG05
    |    |    |--E. atomus (Schaufuss 1887) NG05
    |    |    `--E. sphaerica (Motchulsky 1851) NG05
    |    |--Comatopselaphus Schaufuss 1882 NG05
    |    |    |--C. opacicollis Schaufuss 1882 NG05
    |    |    `--C. puncticollis Raffray 1895 NG05
    |    |--Rybaxis NG05
    |    |    |--R. amica Reitter 1883 NG05
    |    |    |--R. imperatrix Schaufuss 1882 NG05
    |    |    |--R. nubile Reitter 1882 NG05
    |    |    `--R. sarawakensis Schaufuss 1882 NG05
    |    `--Batraxis Reitter 1881 NG05
    |         |--B. brevis Raffray 1895 NG05
    |         |--B. curvispina Raffray 1892 NG05
    |         |--B. elegans Raffray 1895 NG05
    |         |--B. foveata Raffray 1895 NG05
    |         |--B. gracilicornis Raffray 1895 NG05
    |         |--B. instabilis Raffray 1895 NG05
    |         |--B. obliqua Raffray 1895 NG05
    |         |--B. pumilio Raffray 1895 NG05
    |         |--B. quadrata Raffray 1895 NG05
    |         |--B. truncaticornis Raffray 1895 NG05
    |         `--B. variabilis Raffray 1890 NG05
    `--Plagiophorus Motschulsky 1851 [incl. Cyathiger King 1865; Cyathigerini] NG05
         |--P. baumeisteri (Reitter 1883) NG05
         |--P. besucheti (Burckhardt 1985) NG05
         |--P. borneensis (Reitter 1883) NG05
         |--P. bulbifer (Bryant 1915) NG05
         |--P. cribratus (Raffray 1895) NG05
         |--P. decamerus (Burckhardt 1985) NG05
         |--P. dispar (Bryant 1915) NG05
         |--P. elongatus (Bryant 1915) NG05
         |--P. endecamerus (Burckhardt 1985) NG05
         |--P. heterocerus (Raffray 1895) NG05
         |--P. impar (Raffray 1895) NG05
         |--P. inexpectus (Raffray 1895) NG05
         |--P. magnicornis ((Burckhardt 1985) NG05
         |--P. malayanus (Bryant 1915) NG05
         |--P. matangensis (Bryant 1915) NG05
         |--P. mirandus (Bryant 1915) NG05
         |--P. murphyi (Burckhardt 1985) NG05
         |--P. myrmecophilus (Bryant 1915) NG05
         |--P. octomerus (Burckhardt 1985) NG05
         |--P. penangensis (Burckhardt 1985) NG05
         |--P. pseudoheptamerus (Burckhardt 1985) NG05
         |--P. raffrayi (Bryant 1915) NG05
         |--P. rufus (Raffray 1895) NG05
         |--P. sarawakensis (Bryant 1915) NG05
         |--P. schaufussi Reitter 1883 NG05
         |--P. sericeus (Raffray 1895) NG05
         |--P. simoni (Reitter 1883) NG05
         |--P. variolosus (Raffray 1895) NG05
         |--P. vicinus (Burckhardt 1985) NG05
         `--P. viduus (Raffray 1895) NG05

*Type species of generic name indicated

REFERENCES

[NG05] Nomura, S., & I. Abd Ghani. 2005. Faunistic notes on the pselaphine species of the supertribes Goniaceritae, Pselaphitae and Clavigeritae from Malaysia and Singapore (Coleoptera: Staphylinidae: Pselaphinae). Serangga 10 (1-2): 1-36.