Field of Science


Trichostomum brachydontium, photographed by Heino Lepp.

Belongs within: Musci.
Contains: Weissia, Barbula, Tortula.

The Pottiaceae is a large, diverse family of mosses found worldwide, divided between about eighty genera. Members of the family are commonly small and found in harsh environments such as rocks and disturbed soil, primarily in calcareous habitats.

Characters (from Smith & Smith 2004): Plants usually small, often xeromorphic. Leaves linear to spathulate or ovate, acuminate to rounded; margins entire to dentate, usually unbordered; costa usually strong, in section with one or two stereid bands; basal cells rectangular, more or less hyaline, cells above hexagonal or quadrate, often papillose, sometimes obscure. Setae very short to long, usually straight; capsules erect or inclined, globose to cylindrical, straight or curved, cleistocarpous or dehiscent; peristome absent, rudimentary, imperfect or perfect, teeth straight or spirally coiled, entire, perforated or variously cleft or divided into filiform segments.

<==Pottiaceae [Pottieae]
    |--Timmiella barbuloides FHH01
    |--Oxystegus tenuirostris N02
    |--Astomum Hampe 1837 SK02
    |--Phasconica Müll.Hal. 1882 SK02
    |--Pseudocrossidium Williams 1915 SK02
    |--Weissia SK02
    |--Barbula SK02
    |--Tortula SK02
    |--Molendoa sendtneriana J87
    |--Semibarbula orientalis (Weber) Wijk & Margad. 1959 [=Trichostomum orientale Weber 1804] SG07
    |--Pleurochaete J87
    |    |--P. luteola J87
    |    `--P. squarrosa C74
    |--Calymperastrum Stone 1986 SK02
    |    `--C. latifolium (Hampe) Stone 1986 [=Calymperes latifolium] SK02
    |--Chenia Zander 1989 [incl. Leptophascum] SK02
    |    `--C. leptophylla (Müll.Hal.) Zander 1993 (see below for synonymy) SK02
    |--Gymnostomiella Fleischer 1904 SK02
    |    `--G. vernicosa (Hooker) Fleischer 1904 [=Gymnostomum vernicosum Hooker 1836] B05 SK02
    |--Phascopsis Stone 1980 SK02
    |    `--P. rubicunda Stone 1980 SK02
    |--Stonea Zander 1989 SK02
    |    `--S. oleaginosa (Stone) Zander 1989 [=Tortula oleaginosa Stone 1978] SK02
    |--Hymenostylium Brid. 1827 J87, SK02
    |    `--H. recurvirostre (Hedw.) Dixon 1933 SK02
    |--Gymnostomum Nees & Hornsch. 1823 SK02
    |    |--G. aeruginosum Smith 1804 SK02
    |    `--G. calcareum Nees & Hornsch. 1823 (see below for synonymy) SK02
    |--Pseudosymblepharis Broth. 1924 SK02
    |    |--P. bombayensis (Müll.Hal.) Sollman 1996 [incl. P. angustata (Mitt.) Hilp. 1933] SK02
    |    |--P. schimperianus J87
    |    `--P. subduriuscula (Müll.Hal.) Chen 1941 SK02
    |--Crossidium Jur. 1882 SK02
    |    |--C. davidai Catches. 1980 SK02
    |    `--C. geheebii (Broth.) Broth. 1902 [=Barbula geheebii (nom. inv.)] SK02
    |--Hennediella Paris 1896 SK02
    |    |--H. heimii (Hedw.) Zander 1993 [=Pottia heimii (Hedw.) Hampe 1837] SK02
    |    `--H. stanfordensis (Steere) Blockeel 1990 [=Tortula stanfordensis Steere 1951] SK02
    |--Pterygoneurum Jur. 1897 SK02
    |    |--P. macleanum Warnst. 1916 [incl. P. kemsleyi Willis 1954] SK02
    |    `--P. ovatum (Hedw.) Dixon 1934 SK02
    |--Splachnobryum Müll.Hal. 1869 SK02
    |    |--S. obtusum (Brid.) Müll.Hal. 1869 (see below for synonymy) SK02
    |    `--S. wiemansii Fleischer 1904 SK02
    |--Uleobryum Broth. 1906 SK02
    |    |--U. occultum (Roth) Zander 1993 [incl. U. curtisii Stone 1984] SK02
    |    `--U. peruvianum Broth. 1906 SK02
    |--Trachycarpidium Broth. 1901 SK02
    |    |--T. brisbanicum (Müll.Hal.) Stone 1975 (see below for synonymy) SK02
    |    `--T. verrucosum SK02
    |--Hydrogonium SG07
    |    |--H. arcuatum (Griffith) Wijk. & Marg. 1958 [=Barbula arcuata Griffith 1842] B05
    |    `--H. consanguineum (Thwaites & Mitt.) Hilp. 1933 [=Tortula consanguinea Thwaites & Mitt. 1873] SG07
    |--Anoectangium Schwägr. 1811 SK02
    |    |--A. aestivum (Hedw.) Mitt. 1869 SK02
    |    |--A. bellii Broth. ex Dixon 1926 SK02
    |    `--A. thomsonii N02
    |--Hyophila Brid. 1827 SK02
    |    |--H. involuta (Hooker) Jaeger 1873 SK02 [=Gymnostomum involutum Hooker 1819 SG07; incl. H. micholitzii SK02]
    |    |--H. javanica (Nees & Blume) Brid. 1827 SK02
    |    `--H. rosea Williams 1914 SK02
    |--Calyptopogon (Mitt.) Broth. 1902 SK02
    |    `--C. mnioides (Schwägr.) Broth. 1902 (see below for synonymy) SK02
    |         |--C. m. var. mnioides SK02
    |         `--‘Streptopogon’ m. var. prostratus [=Tortula prostrata] SK02
    |--Goniomitrium Hooker & Wilson 1846 SK02
    |    `--G. acuminatum Hooker & Wilson 1846 SK02
    |         |--G. a. ssp. acuminatum [incl. Physcomitrium goniomitrium] SK02
    |         `--G. a. ssp. enerve (Hooker & Wilson) Fife & Seppelt 2001 (see below for synonymy) SK02
    |--Microbryum Schimp. 1860 SK02
    |    |--M. brevicaule (Taylor) Zander 1993 (see below for synonymy) SK02
    |    |--M. davallianum (Smith) Zander 1993 [=Pottia davalliana (Smith) Jensen 1923] SK02
    |    `--M. starckeanum (Hedw.) Zander 1993 (see below for synonymy) SK02
    |--Leptodontium (Müll.Hal.) Hampe ex Lindb. 1864 SK02
    |    |--L. interruptum (Mitt.) Broth. 1902 SK02
    |    |--L. paradoxum Stone & Scott 1981 SK02
    |    `--L. viticulosoides (Palisot de Beauvois) Wijk & Margad. 1960 (see below for synonymy) SK02
    |--Triquetrella Müll.Hal. 1897 SK02
    |    |--T. fragilis Müll.Hal. 1897 SK02
    |    |--T. papillata (Hooker & Wilson) Broth. 1902 (see below for synonymy) SK02
    |    |--T. richardsiae Müll.Hal. 1897 SK02
    |    `--T. tasmanica (Broth.) Granzow 1989 (see below for synonymy) SK02
    |--Tortella (Lind.) Limpr. 1888 SK02
    |    |--T. dakinii Willis 1955 SK02
    |    |--T. flavovirens (Bruch ex Muell.) Broth. 1902 SK02
    |    |--T. inclinata (Hedw.) Limpr. 1888 [=Barbula inclinata; incl. Tortula nervosa (preoc.)] SK02
    |    |--T. knightii (Mitt.) Broth. 1902 (see below for synonymy) SK02
    |    `--T. novae-valesiae Broth. 1916 SK02
    |--Tetrapterum Hampe ex Jaeger 1869 SK02
    |    |--T. brachypelma (Müll.Hal.) Broth. 1924 (see below for synonymy) SK02
    |    |--T. cylindricum (Taylor) Jaeger 1869 (see below for synonymy) SK02
    |    |--T. sullivanii (Müll.Hal.) Broth. 1924 (see below for synonymy) SK02
    |    |--T. tetrapteroides (Müll.Hal.) Broth. 1924 (see below for synonymy) SK02
    |    `--T. weymouthii (Müll.Hal.) Broth. 1924 [=Phascum weymouthii, Astomum weymouthii, Systegium weymouthii] SK02
    |--Aloina Kindb. 1882 SK02
    |    |--A. aloides (Schultz) Kindb. 1883 SK02
    |    |    |--A. a. var. aloides SK02
    |    |    `--A. a. var. ambigua (Bruch & Schimp.) Craig 1939 SK02
    |    |--A. bifrons (De Not.) Delgad. 1973 SK02
    |    |--A. rigida (Hedw.) Limpr. 1888 SK02
    |    `--A. sullivaniana (Müll.Hal.) Broth. 1902 (see below for synonymy) SK02
    |--Didymodon Hedw. 1801 SK02
    |    |--D. australasiae (Hooker & Greville) Zander 1978 (see below for synonymy) SK02
    |    |--D. dubius (Schwägr.) Paris 1904 (see below for synonymy) SK02
    |    |--D. rubiginosus (Müll.Hal.) Broth. 1902 (see below for synonymy) SK02
    |    |--D. tophaceus (Brid.) Lisa 1837 (see below for synonymy) SK02
    |    |--D. torquatus (Taylor) Catches. 1980 (see below for synonymy) SK02
    |    `--D. vinealis (Brid.) Zander 1978 SK02
    |--Bryoerythrophyllum Chen 1941 SK02
    |    |--B. binnsii (R.Br.bis) Wijk & Margad. 1959 (see below for synonymy) SK02
    |    |--B. dentatum N02
    |    |--B. jamesonii (Taylor) Crum 1957 SK02
    |    |--B. recurvirostre (Hedw.) Chen 1941 (see below for synonymy) SK02
    |    |--B. recurvum N02
    |    |--B. rotundatum (Lindb. & Arnell) Chen 1941 SK02
    |    `--B. wallichii [incl. Barbula zygodontifolia] SK02
    |--Acaulon Müll.Hal. 1847 SK02
    |    |--A. chrysacanthum Stone 1976 SK02
    |    |--A. eremicola Stone 1979 SK02
    |    |--A. granulosum Stone 1988 SK02
    |    |--A. integrifolium Müll.Hal. 1855 (see below for synonymy) SK02
    |    |--A. leucochaete Stone 1976 SK02
    |    |--A. mediterraneum Limpr. 1885 [=A. muticum var. mediterraneum] SK02
    |    |--A. muticum FHH01
    |    `--A. triquetrum (Spruce) Müll.Hal. 1847 SK02
    `--Trichostomum Bruch 1829 SK02
         |--T. austrocrispum (Beckett) Zander 1993 (see below for synonymy) SK02
         |--T. bombayense N02
         |--T. brachydontium Bruch 1829 (see below for synonymy) SK02
         |--T. crispulum Bruch 1829 SK02
         |--T. diminutum [=Timmiella diminuta] SK02
         |--T. eckelianum Zander 1993 (see below for synonymy) SK02
         |--T. leptotheca Müll.Hal. 1868 (n. d.) SK02
         |--T. pallidum [=Ditrichum pallidum] SK02
         |--T. polyphyllum [=Ptychomitrium polyphyllum] SK02
         |--T. sublamprothecium J87
         `--T. tenuirostre J87

Nomina nuda: Hyophila angustifolia Müll.Hal. ex Kindb. 1889 SK02
             Hyophila muelleri Watts & Whitel. 1902 SK02
             Hyophila pugionifolia Müll.Hal. ex Kindb. 1888 [=Weissia pugionifolia Müll.Hal. ex Bailey 1890] SK02
             Leptodontium australe Müll.Hal. 1901 SK02
             Tortella perhumilis Broth. ex Bailey 1896 SK02
             Trichostomum albovaginatum Hampe ex Müll.Hal. 1901 SK02
             Trichostomum weymouthii Müll.Hal. 1901 [=Desmatodon weymouthii Watts & Whitel. 1902] SK02
             Triquetrella albicuspis Broth. ex Watts 1906 SK02
             Triquetrella triquetra Müll.Hal. ex Kindb. 1889 SK02

Acaulon integrifolium Müll.Hal. 1855 [=Sphaerangium integrifolium; incl. A. apiculatum, Phascum apiculatum (nom. illeg.), Sphaerangium apiculatum, Acaulon apiculatum var. aristatum, A. integrifolium var. aristatum (Willis) Willis 1957, A. austromuticum, A. sullivanii, S. sullivanii, Phascum apiculatum var. turgidum, Acaulon apiculatum var. turgidum, Sphaerangium apiculatum var. turgidum] SK02

Aloina sullivaniana (Müll.Hal.) Broth. 1902 [=Barbula sullivaniana, Tortula sullivaniana; incl. B. lamellosa (n. n.), T. lamellosa (n. n.)] SK02

Bryoerythrophyllum binnsii (R.Br.bis) Wijk & Margad. 1959 [=Didymodon binnsii, Erythrobarbula binnsii; incl. Barbula glaucula Broth. 1916 (preoc.)] SK02

Bryoerythrophyllum recurvirostre (Hedw.) Chen 1941 [=Barbula recurvirostris, Didymodon recurvirostris, Erythrobarbula recurvirostris, Erythrophyllum recurvirostre; incl. Barbula glabriuscula, Didymodon rubellus (nom. illeg.), Trichostomum subrubellum] SK02

Calyptopogon mnioides (Schwägr.) Broth. 1902 [=Streptopogon mnioides, Tortula mnioides; incl. Barbula crispata, Calyptopogon crispatus, Streptopogon crispatus, Tortula crispata, Barbula wilhelmii, Calyptopogon wilhelmii, Streptopogon wilhelmii, Tortula wilhelmii] SK02

Chenia leptophylla (Müll.Hal.) Zander 1993 [=Phascum leptophyllum Müll.Hal. 1888, *Leptophascum leptophyllum; incl. Tortula evanescens Broth. 1916, T. rhizophylla (Sak.) Iwats. & Saito 1972] SK02

Didymodon australasiae (Hooker & Greville) Zander 1978 [=Tortula australasiae, Barbula australasiae, Trichostomopsis australasiae; incl. Trichostomum calcicola, Barbula calcicola (Hampe) Broth. 1902, Tortula calcicola, Trichostomum fuscescens, To. fuscescens (nom. inv.), Barbula menziesiana (nom. inv.), B. nanocaulis, B. rufiseta] SK02

Didymodon dubius (Schwägr.) Paris 1904 [=Syrrhopodon dubius; incl. Codonoblepharon schwaegrichenii (nom. illeg.), Zygodon schwaegrichenii (nom. illeg.)] SK02

Didymodon rubiginosus (Müll.Hal.) Broth. 1902 [=Trichostomum rubiginosum, Erythrophyllum rubiginosum, Tortula rubiginosa] SK02

Didymodon tophaceus (Brid.) Lisa 1837 [incl. Barbula wildii, Didymodon wildii (Broth.) Broth. 1902, Tortula wildii (n. n.), Trichostomum wildii] SK02

Didymodon torquatus (Taylor) Catches. 1980 [=Barbula torquata, Tortula torquata; incl. B. chrysochaete Müll.Hal. 1898, B. chrysopus Müll.Hal. 1898, Tortula crispifolia, B. crispifolia (Mitt.) Jaeger 1873, B. luehmannii, Didymodon luehmannii (Broth. & Geh.) Catches. 1980, B. subtorquata, D. subtorquatus (Müll.Hal. & Hampe) Catches. 1980, Tortula subtorquata] SK02

Goniomitrium acuminatum ssp. enerve (Hooker & Wilson) Fife & Seppelt 2001 [=G. enerve Hooker & Wilson 1846, Physcomitrium enerve] SK02

Gymnostomum calcareum Nees & Hornsch. 1823 [=Weissia calcarea (preoc.); incl. G. calcareum var. australe, G. calcareum var. longifolium, Eucladium tasmanicum] SK02

Leptodontium viticulosoides (Palisot de Beauvois) Wijk & Margad. 1960 [incl. Trichostomum subdenticulatum, L. whitteronii Broth. ex Stone & Scott 1981 (nom. inv.)] SK02

Microbryum brevicaule (Taylor) Zander 1993 [=Gymnostomum brevicaule, Pottia brevicaulis (Taylor) Müll.Hal. 1849] SK02

Microbryum starckeanum (Hedw.) Zander 1993 [=Pottia starckeana (Hedw.) Müll.Hal. 1849; incl. Anacalypta cespitulosa, P. cespitulosa, P. caespitulosa (l. c.)] SK02

Splachnobryum obtusum (Brid.) Müll.Hal. 1869 [incl. S. baileyi Broth. 1888, S. geheebii Fleischer 1904, S. indicum Hampe & Müll.Hal. 1872] SK02

Tetrapterum brachypelma (Müll.Hal.) Broth. 1924 [=Phascum brachypelma, Astomum brachypelma, Systegium brachypelma] SK02

Tetrapterum cylindricum (Taylor) Jaeger 1869 [=Astomum cylindricum, Phascum tetragonum var. cylindricum, Systegium cylindricum, Tetrapterum capense var. cylindricum; incl. T. australe (nom. inv.)] SK02

Tetrapterum sullivanii (Müll.Hal.) Broth. 1924 [=Phascum sullivanii, Astomum sullivanii (preoc.), Pleuridium sullivanii, Systegium sullivanii] SK02

Tetrapterum tetrapteroides (Müll.Hal.) Broth. 1924 [=Phascum tetrapteroides, Astomum tetrapteroides, Systegium tetrapteroides] SK02

Tortella knightii (Mitt.) Broth. 1902 [=Tortula knightii, Barbula knightii; incl. B. nanotortuosa, Tortella nanotortuosa] SK02

Trachycarpidium brisbanicum (Müll.Hal.) Stone 1975 [=Acaulon brisbanicum, Astomum brisbanicum, Sphaerangium brisbanicum; incl. As. wattsii] SK02

Trichostomum austrocrispum (Beckett) Zander 1993 [=Weissia austrocrispa (Beckett) Stone 1980; incl. Pleuridium lanceolatum (R.Br.bis) Paris 1898] SK02

Trichostomum brachydontium Bruch 1829 [incl. Tortella aristatula Broth. ex Watts & Whitel. 1902 (n. n.), Tr. aristatulum Broth. 1916] SK02

Trichostomum eckelianum Zander 1993 [incl. Tortella cirrhata Broth. 1902, Tr. cirrhatum (preoc.), To. subflavovirens Broth. & Watts 1915] SK02

Triquetrella papillata (Hooker & Wilson) Broth. 1902 [=Didymodon papillatus, Leptodontium papillatum; incl. Triquetrella filiformis, Zygodon preissianus, T. preissiana (Hampe) Müll.Hal. 1897, Z. preissii (l. c.), Leskea rubricaulis, Triquetrella scabra, Zygodon scaber (n. n.)] SK02

Triquetrella tasmanica (Broth.) Granzow 1989 [=Anomodon tasmanicus Broth. 1900, Anoectangium tasmanicum (nom. inv.); incl. Triquetrella curvifolia] SK02

*Type species of generic name indicated


[B05] Bhattacharyya, P. K. 2005. Bryophyte flora of Barddhaman district, West Bengal. Bulletin of the Botanical Survey of India 47 (1-4): 11-42.

[C74] Coineau, Y. 1974. Éléments pour une monographie morphologique, écologique et biologique des Caeculidae (Acariens). Mémoires du Muséum National d'Histoire Naturelle, nouvelle série, Série A, Zoologie 81: 1-299, 224 pls.

[FHH01] Frey, W., M. Hofmann & H. H. Hilger. 2001. The gametophyte-sporophyte junction: Unequivocal hints for two evolutionary lines of archegoniate land plants. Flora 196: 431-445.

[J87] Judd, W. S. 1987. Floristic study of Morne La Visite and Pic Macaya National Parks, Haiti. Bulletin of the Florida State Museum – Biological Sciences 32 (1): 1-136.

[N02] Negi, H. R. 2002. Abundance and diversity of moss communities of Chopta-Tunganath in the Garhwal Himalaya. Journal of the Bombay Natural History Society 99 (3): 418-433.

[SG07] Singh, S. K. & J. P. Ghosh. 2007. Bryo-diversity in Indian Botanic Garden, Howrah, West Bengal. Bulletin of the Botanical Survey of India 49 (1-4): 155-164.

Smith, A. J. E., & R. Smith. 2004. The Moss Flora of Britain and Ireland, 2nd ed. Cambridge University Press.

[SK02] Streimann, H., & N. Klazenga. 2002. Catalogue of Australian Mosses. Flora of Australia Supplementary Series 17. Australian Biological Resources Study: Canberra.

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