Setae of Hypnodendron arcuatum, from here.

Belongs within: Bryidae.

Hypnodendron is a genus of large dendroid mosses found mostly in south-east Asia and Australasia, where they are a prominent component of many wet forest floras. Molecular phylogenetic analysis has suggested that it may be paraphyletic within a clade referred to as the hypnodendroid pleurocarps (Bell & Newton 2005).

Characters (from Bell & Newton 2005): Leaf border cells not differentiated. Plants dendroid; adaxial rhizoids present, micronemata absent. Vegetative secondary modules ('branches') present; primary module heteroblasty ending in secondary module leaves; primary module growth determinate, branching from base but not from apex; perichaetia on highly reduced secondary modules, located mid-module. Dioicous.

<==Hypnodendron (Müll.Hal.) Lindb. 1861 [Hypnodendraceae] SK02
    |--H. arcuatum (see below for synonymy) SK02
    |--H. colensoi (Hooker & Wilson) Mitt. 1882 [=Isothecium colensoi] SK02
    |--H. comatulum (Geh. ex Broth.) Touw 1971 [=Mniodendron comatulum] SK02
    |--H. comosum (Labill.) Mitt. 1882 (see below for synonymy) SK02
    |    |--H. c. var. comosum [incl. Mniodendron kroneanum] SK02
    |    `--H. c. var. sieberi (Müll.Hal.) Touw 1971 [=Hypnum sieberi, Isothecium sieberi, Mniodendron sieberi] SK02
    |--H. krausei D03
    |--H. menziesii (Hooker) Paris 1895 [=Isothecium menziesii, Trachyloma menziesii] SK02
    |--H. palmeum SK02
    |--H. planifrons SK02
    |--H. spininervium (Hooker) Jaeger & Sauerb. 1879 [=Hypnum spininervium, Isothecium spininervium] SK02
    |    |--H. s. ssp. spininervium SK02
    |    `--H. s. ssp. archeri (Mitt.) Touw 1971 (see below for synonymy) SK02
    `--H. vitiense Mitt. 1873 SK02
         |--H. v. ssp. vitiense SK02
         `--H. v. ssp. australe Touw 1971 [incl. H. spininervium var. pumilum (n. n.), H. whiteleggei (n. n.)] SK02

Nomina nuda: Hypnodendron curvatocomosum Müll.Hal. ex Bailey 1886 SK02
             Hypnodendron eflagellare Ångström ex Bailey 1886 SK02
             Hypnodendron flagellare Ångström ex Bailey 1886 SK02
             Hypnodendron pterobryoides Müll.Hal. ex Bailey 1886 SK02

Hypnodendron arcuatum [=Hypnum arcuatum, Hypnodendron spininervium var. arcuatum (nom. illeg.), Racopilum arcuatum, Trachyloma arcuatum] SK02

Hypnodendron comosum (Labill.) Mitt. 1882 [=Hypnum comosum, Isothecium comosum, Mniodendron comosum, Trachyloma comosum] SK02

Hypnodendron spininervium ssp. archeri (Mitt.) Touw 1971 [=Isothecium archeri, Hypnum archeri, Trachyloma archeri; incl. Hypnodendron leiopyxis] SK02

*Type species of generic name indicated


Bell, N. E., & A. E. Newton. 2005. The paraphyly of Hypnodendron and the phylogeny of related non-hypnanaean pleurocarpous mosses inferred from chloroplast and mitochondrial sequence data. Systematic Botany 30 (1): 34-51.

[D03] Dusén, P. 1903. The vegetation of western Patagonia. In Reports of the Princeton University Expeditions to Patagonia, 1896-1899, vol. 8 – Botany (W. B. Scott, ed.) pp. 1-34. The University: Princeton (New Jersey).

[SK02] Streimann, H., & N. Klazenga. 2002. Catalogue of Australian Mosses. Flora of Australia Supplementary Series 17. Australian Biological Resources Study: Canberra.

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