Belongs within: Parasitengonina.
The Johnstonianidae are a group of relatively large parasitengonine mites found in moist litter habitats, characterised by nude or sparsely barbed idionotal setae and the presence of a large ventral or lateral spine on the palptibia (Walter et al. 2009).
<==Johnstonianidae
| i. s.: Crossothrombium Womersley 1939 H98
| `--C. parkhousei Womersley 1939 H98
| Hirstithrombium Oudemans in Thor & Willmann 1947 H98
| `--H. australiense (Hirst 1928) [=Diplothrombium australiense, Johnstoniana australiense] H98
|--Charadracarus K90 [Charadracarinae We91]
|--Polydiscus [Polydiscinae] K90
`--Johnstonianinae We91
|--Centrotrombidium Kramer 1896 SB63
| |--*C. schneideri Kramer 1896 [incl. Diplothrombium walteri Schweizer 1922, C. walteri] SB63
| `--C. australasiae Womersley 1942 H98
|--Diplothrombium Berlese 1910 SB63
| |--*D. longipalpe (Berlese 1887) [=Trombidium longipalpe] SB63
| |--D. eximium S22
| |--D. micidium WL09
| `--D. monoense WL09
`--Johnstoniana George 1909 H98 [incl. Rohaultia Oudemans 1911 SB63]
|--*J. errans (Johnston 1852) [=Rhyncholophus errans] SB63
|--‘Rohaultia’ biungulum AL37
|--J. dubiosa Schweizer 1951 SB63
|--J. insignia (Berlese 1916) [=Diplothrombium eximium insignius; incl. D. longipes Schweizer 1922] SB63
|--J. parva WWO01
|--J. rapax WWO01
|--J. tuberculata Schweizer 1951 [=J. tuberculatua] SB63
|--J. ventripilosa Wi91
`--J. vitzthumi Womersley 1939 H98
*Type species of generic name indicated
REFERENCES
[AL37] André, M., & É. Lamy. 1937. Les Idées actuelles sur la Phylogénie des Acariens. Published by the author: Paris.
[H98] Halliday, R. B. 1998. Mites of Australia: A checklist and bibliography. CSIRO Publishing: Collingwood.
[K90] Kethley, J. 1990. Acarina: Prostigmata (Actinedida). In: Dindal, D. L. (ed.) Soil Biology Guide pp. 667–756. John Wiley & Sones: New York.
[S22] Schweizer, J. 1922. Beitrag zur Kenntnis der terrestrischen Milbenfauna der Schweiz. Verhandl. Naturf. Ges. Basel 33: 23–112, 4 pls.
[SB63] Schweizer, J. & C. Bader. 1963. Die Landmilben der Schweiz (Mittelland, Jura und Alpen): Trombidiformes Reuter, mit 217 Arten und Unterarten und 193 Originalzeichnungen. Denkschriften der Schweizerischen Naturforschenden Gesellschaft [Mémoires de la Société Helvétique des Sciences Naturelles] 84 (2): i–vi, 209–378.
[WL09] Walter, D. E., E. E. Lindquist, I. M. Smith, D. R. Cook & G. W. Krantz. 2009. Order Trombidiformes. In: Krantz, G. W., & D. E. Walter (eds) A Manual of Acarology 3rd ed. pp. 233–420. Texas Tech University Press.
[We91] Welbourn, W. C. 1991. Phylogenetic studies of the terrestrial Parasitengona. In: Dusbábek, F., & V. Bukva (eds) Modern Acarology: Proceedings of the VIII International Congress of Acarology, held in České Budĕjovice, Czechoslovakia, 6–11 August 1990 vol. 2 pp. 163–170. SPB Academic Publishing: The Hague.
[Wi91] Witte, H. 1991. The phylogenetic relationships within the Parasitengonae. In: Dusbábek, F., & V. Bukva (eds) Modern Acarology: Proceedings of the VIII International Congress of Acarology, held in České Budĕjovice, Czechoslovakia, 6–11 August 1990 vol. 2 pp. 171–182. SPB Academic Publishing: The Hague.
[WWO01] Wohltmann, A., H. Witte & R. Olomski. 2001. Organismal patterns causing high potential for adaptive radiation in Parasitengonae (Acari: Prostigmata). In: Halliday, R. B., D. E. Walter, H. C. Proctor, R. A. Norton & M. J. Colloff (eds) Acarology: Proceedings of the 10th International Congress pp. 83–99. CSIRO Publishing: Melbourne.
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