Seaside brookweed Samolus valerandi, copyright Christian Fischer.

Belongs within: Ericales.
Contains: Primula, Androsace, Dodecatheon, Lysimachia, Myrsine, Ardisia.

Though its members have been included by recent classifications in the Ericales, the name 'Primulales' is used here for a clade containing the Primulaceae and Myrsinaceae. Members of the clade usually have spirally arranged leaves, and sympetalous flowers with stamens borne opposite to petals ( Species of Primulaceae are herbs bearing capsular fruits whereas Myrsinaceae are trees or shrubs whose fruits are drupes (Allan 1961).

Synapomorphies (from, as Primulaceae): Inflorescence racemose; petals and stamens from common primordia; stamens equal to and opposite petals, staminodes opposite calyx, represented by at least a vascular trace; nectary present; gynoecium with 5 connate carpels, opposite corolla, placentation free-central, style short, hollow, stigma more or less capitate; ovules at least partly immersed in swollen placenta, micropyle bistomal, outer integument about 2 cells across, inner integument 3-4 cells across, endothelium present, tanniniferous; seeds angled; endotesta crystalliferous; endosperm nuclear, copious.

    |--Theophrastaceae T00
    |    |--Clavija K01
    |    |--Theophrasta jussieui JK80
    |    |--Clavijopsis staubi Schindehütte 1907 CBH93
    |    `--Jacquinia JK80
    |         |--J. aurantiaca JK80
    |         |--J. eggersii J87
    |         `--J. purgena JK80
    |--Maesa [Maesaceae] T00
    |    |--M. chisia D07
    |    |--M. dependens B00
    |    |--M. haplobotrys B00
    |    |--M. indica C55b
    |    |--M. macrophylla SN88
    |    |--M. montana SN88
    |    `--M. muelleri B00
    |--Primulaceae T00
    |    |--Coris monspeliensis T00, C55a
    |    |--Primula JK80
    |    |--Androsace O88
    |    |--Omphalogramma elwesiana O88
    |    |--Hottonia palustris JK80
    |    |--Asterolinon linum-stellatum (Linnaeus) Duby 1844 JK80, PL04
    |    |--Dodecatheon H93
    |    |--Cortusa matthioli JK80
    |    |--Glaux maritima JK80
    |    |--Centunculus minimus H93
    |    |--Lysimachia JK80
    |    |--Nummularia Hill 1756 KC01
    |    |--Trientalis JK80
    |    |    |--T. arctica H93
    |    |    |--T. europaea C55a
    |    |    `--T. latifolia H93
    |    |--Cyclamen JK80
    |    |    |--C. coum V09
    |    |    |--C. graecum Link 1835 PL04
    |    |    |--C. hederaefolium C55b
    |    |    `--C. persicum JK80
    |    |--Anagallis JK80
    |    |    |--A. arvensis Linnaeus 1753 PL04
    |    |    |    |--A. a. var. arvensis GK00
    |    |    |    `--A. a. var. caerulea GK00
    |    |    |--A. foemina AGF98
    |    |    |--A. linifolia H93
    |    |    `--A. phaenicea C55b
    |    `--Samolus Linnaeus 1753 A61
    |         |--S. caespitosus GK00
    |         |--S. ebracteatus H93
    |         |--S. junceus GK00
    |         |--S. parviflorus H93
    |         |--S. repens (Forster & Forster) Pers. 1805 A61 (see below for synonymy)
    |         |    |--S. r. var. repens KM08
    |         |    `--S. r. var. paucifolius KM08
    |         `--S. valerandi JK80
    `--Myrsinaceae [Myrsinales, Myrsineae, Myrsinoideae] T00
         |--Cybianthus YY22
         |--Grenacheria YY22
         |--Hymenandra YY22
         |--Monoporus YY22
         |--Weigeltia YY22
         |--Geisanthus YY22
         |--Discocalyx YY22
         |--Grammadenia YY22
         |--Stylogyne YY22
         |--Conomorpha YY22
         |--Amblyanthopsis GDC02
         |--Myrsine HD03
         |--Myrsinopsis succinea Conwentz 1886 CBH93
         |--Pleiomerites reticulatus Ettingshausen 1868 CBH93
         |--Pleiomeropsis rottensis Weyland 1938 CBH93
         |--Ardisia JK80
         |--Tapeinosperma pseudojambosa H90
         |--Peckia Vell. 1825 KC01
         |--Suttonia Rich. 1832 KC01
         |--Elingamita Baylis 1951 A61
         |    `--*E. johnsonii Baylis 1951 A61
         |--Aegiceras [Aegicerataceae] T00
         |    |--A. corniculatum LK14
         |    `--A. majus H09
         |--Wallenia J87
         |    |--W. aquifolia JS87
         |    |--W. ekmanii J87
         |    `--W. formonensis Judd 1986 JS87
         |--Embelia SR07
         |    |--E. australiana H90
         |    |--E. curvinervia B00
         |    |--E. philippinensis K03
         |    `--E. tsjeriam-cottam SR07
         |--Amblyanthus GDC02
         |    |--A. glandulosus GDC02
         |    |--A. multiflorus GDC02
         |    |--A. obovatus Giri, Das & Chowdhery 2002 GDC02
         |    `--A. praetervisus GDC02
         `--Rapanea JK80
              |--R. ferruginea SWK87
              |--R. howittiana H90
              |--R. melanophloeus JK80
              |--R. minutifolia H03
              |--R. porosa B00
              |--R. subsessilis H90
              `--R. variabilis B00

Samolus repens (Forster & Forster) Pers. 1805 A61 [=Sheffieldia repens Forster & Forster 1776 A61; incl. Sa. littoralis C06, Sa. repens var. procumbens Knuth 1905 A61, Sa. repens var. strictus Cockayne 1916 A61]

*Type species of generic name indicated


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Last updated: 26 June 2020.

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