Mimosoideae

Umbrella thorn acacia Acacia tortilis, photographed by Enric G.


Belongs within: Fabales.
Contains: Acacia series Uninerves subseries Racemosae.

The Mimosoideae is a clade of leguminous plants (mostly trees or shrubs) containing the acacias and related species. Mimosoids generally have small glowers clustered into heads or spikes. The largest genus recognised in the clade has been Acacia, with about 1300 species from South America, Africa and Australia included therein; however, Acacia in this sense is now recognised as non-monophyletic and it has been divided between a number of segregate genera. Though originally named for an African species, the retypification of the genus with an Australian species has been proposed in order to retain the name for the largest of these segregates (Orchard & Maslin 2003). Acacia in this sense would include the great majority of Australian species; species from other continents would be placed in genera Vachellia and Senegalia.

Other mimosoid genera include the mesquites, Prosopis, a pantropical genus of spiny trees and shrubs that are significant parts of the vegetation of many arid regions. Members of the genus Mimosa include the sensitive plant M. pudica, so named for the way the leaves fold away when touched. Inga, the shimbillos, is a Neotropical genus of trees and shrubs producing fruit as bean-like pods containing seeds covered with a sweet white powder and enclosed within a fibrous, sweet pulp.

Characters (from Klitgaard & Lewis): Trees, shrubs, lianas, rarely aquatic herbs; root nodules generally present. Leaves compound, usually bipinnate or less frequently paripinnate, often with specialized "ant" glands, rarely with a tendril. Flowers radially symmetrical, usually small, aggregated into heads or spikes; sepals generally united at the base; petals valvate in bud, the median petal (similar in size and shape to the other four) not overlapped or overlapping the other four; stamens (3-)10-100(-100+), usually all similar in shape and size, filaments free or fused at the base, always exserted from the corolla. Seeds usually with an open pleurogram on each lateral surface.

<==Mimosoideae [Mimosaceae, Mimoseae]
    |--Senegalia [=Acacia subg. Aculeiferum] OM03
    |--Dichrostachys cinerea PP07
    |--Desmanthus virgatus SJ73, C55 [incl. D. leptophyllus C55, D. strictus C55]
    |--Pararchidendron pruinosum B00b
    |--Entada phaseoloides B00b
    |--Adenanthera B00b
    |    |--A. abrosperma B00b
    |    `--A. pavonina P88
    |--Paraserianthes B00b
    |    |--P. lophantha B00b
    |    `--P. toona B00b
    |--Leucaena PP07
    |    |--L. glauca B88
    |    |--L. latisiliqua PP07
    |    `--L. leucocephala H06
    |--Calliandra B00b
    |    |--C. eriophylla H93
    |    |--C. houstoniana B00b
    |    |--C. portoricensis MM96
    |    `--C. surinamensis B00b
    |--Pithecellobium P88
    |    |--P. dulce HSS13
    |    |--P. elegans KIW98
    |    |--P. flexicaule RJ11
    |    |--P. monoliferum H42
    |    |--P. oppositifolium J87
    |    `--P. saman F11
    |--Neptunia M99
    |    |--N. dimorphantha B00a
    |    |--N. gracilis LK14
    |    |--N. major LK14
    |    |--N. monosperma M99
    |    |--N. oleracea GC05 [incl. Desmanthus lacustris C55, D. natans C55, Neptunia stolonifera C55]
    |    `--N. triquetra [=Desmanthus triquetrus] C55
    |--Archidendron P88
    |    |--A. ellipticum [=Pithecellobium ellipticum] P88
    |    |--A. grandiflorum B00b
    |    |--A. hirsutum B00a
    |    |--A. hooglandii H03
    |    |--A. jiringa P88
    |    |--A. lucyi B00a
    |    `--A. tenuiracemosum H03
    |--Parkia K06
    |    |--P. auriculata K06
    |    |--P. cachimboensis K06
    |    |--P. gigantean K06
    |    |--P. gigantocarpa K06
    |    |--P. igneiflora K06
    |    |--P. nitida K06
    |    |--P. pendula K06
    |    |--P. platycephala K06
    |    `--P. speciosa CK06
    |--Albizia SK02
    |    |--A. amara RaS02
    |    |--A. canescens LK14
    |    |--A. falcataria R02 [=Paraserianthes falcataria P88]
    |    |--A. lebbeck H06
    |    |--A. niopoides F11
    |    |--A. odoratissima S02
    |    |--A. procera H06
    |    |--A. pruinosa M87
    |    `--A. retusa B00b
    |--Mimosa G09
    |    |--M. asperata C55
    |    |--M. biuncifera BT87
    |    |--M. hamata PP07
    |    |--M. invisa P88
    |    |--M. jacobita G09
    |    |--M. pigra H06
    |    |--M. pudica MM96
    |    |--M. rubicaulis P03
    |    |--M. scabrella OB11
    |    |--M. senegal S99
    |    `--M. sepiaria P88
    |--Vachellia OM03
    |    |--V. ditricha LK14
    |    |--V. farnesiana HH03 [=Acacia farnesiana OM03]
    |    |--‘Mimosa’ nilotica [=Acacia nilotica; incl. A. scorpioides] OM03
    |    |    |--M. n. ssp. nilotica PP07
    |    |    `--‘Acacia’ n. ssp. indica PP07
    |    |--V. pachyphloia LK14
    |    |    |--V. p. ssp. pachyphloia LK14
    |    |    `--V. p. ssp. brevipinnula LK14
    |    |--‘Acacia’ seyal OM03
    |    |--V. suberosa LK14
    |    `--V. valida LK14
    |--Prosopis MF05
    |    |--P. alba O06
    |    |--P. caldenia MF05
    |    |--P. chilensis SN02
    |    |--P. cineraria PP07
    |    |--P. farcta HF03
    |    |--P. flexuosa RJ11
    |    |--P. juliflora PP07
    |    |    |--P. j. var. juliflora S00
    |    |    |--P. j. var. glandulosa S00
    |    |    `--P. ‘glandulosa’ var. torreyana H06
    |    |--P. nigra O06
    |    |--P. pallida B14
    |    |--P. pubescens H93
    |    |--P. strombulifera H93
    |    |--P. tamarugo H11
    |    `--P. velutina K10
    |--Inga K06
    |    |--I. adenophylla
    |    |--I. affinis
    |    |--I. coriacea
    |    |    |--I. c. var. coriacea
    |    |    `--I. c. var. leptopus [=I. leptopus]
    |    |--I. edulis
    |    |--I. fagifolia
    |    |--I. fastuosa
    |    |--I. holtonii
    |    |--I. huberi
    |    |--I. ingoides
    |    |--I. inicuil
    |    |--I. insignis
    |    |--I. laurina
    |    |--I. marginata
    |    |--I. micheliana
    |    |--I. pachycarpa
    |    |--I. pinetorum
    |    |--I. preussii
    |    |--I. ruiziana
    |    |--I. sessilis
    |    |--I. spuria
    |    |--I. uruguensis F11
    |    `--I. vera
    `--Acacia Miller 1754 [=Acacia subg. Phyllodineae, Racosperma] OM03
         |  i. s.: A. acanthocarpa C15
         |         A. accola P09
         |         A. acradenia EF04
         |         A. acuaria S95
         |         A. acuminata OS04
         |         A. adenogonia LK14
         |         A. adoxa LK14
         |           |--A. a. var. adoxa LK14
         |           `--A. a. var. subglabra LK14
         |         A. adunca M16
         |         A. alata GK00
         |         A. alexandri B00b
         |         A. alpina C15
         |         A. ampliceps LK14
         |         A. anceps B00b
         |         A. ancistrocarpa HD04
         |         A. andrewsii G04b
         |         A. aneura B88
         |           |--A. a. var. aneura MLP09
         |           |--A. a. var. intermedia MLP09 [=A. browniana var. intermedia OS04]
         |           |--A. a. var. macrocarpa MLP09
         |           `--A. a. var. pilbara MLP09
         |         A. applanata OS04
         |         A. areolata LK14
         |         A. argyrodendron B00b
         |         A. arida LK14
         |         A. armata TW07
         |         A. aroma HH03
         |         A. arrecta MLP09
         |         A. ataxacantha HK98
         |         A. atkinsiana EF04
         |         A. aulacocarpa B88
         |         A. ayersiana MLP09
         |         A. baileyana H06
         |         A. bancroftii B00b
         |         A. baueri MB08
         |         A. barrettiorum LK14
         |         A. berlandieri FZ77
         |         A. betchei B00b
         |         A. bidwilli C15
         |         A. biflora GK00
         |         A. binata G04b
         |         A. bivenosa EF04
         |         A. brachybotrya B00b
         |         A. brachyclada G04b
         |         A. brachyphylla G04b
         |         A. browniana OS04
         |         A. burrowii BW05
         |         A. buxifolia C15
         |         A. bynoeana M16
         |         A. calamifolia B00b
         |         A. cambagei B00b
         |         A. castanostegia G04b
         |         A. catechu HH03
         |         A. catenulata M04
         |           |--A. c. ssp. catenulata MLP09
         |           `--A. c. ssp. occidentalis MLP09
         |         A. caven MF05
         |         A. celastrifolia RL05
         |         A. chisholmi Ca16
         |         A. chrysochaeta LK14
         |         A. cibaria Ca16
         |         A. citrinoviridis MLP09
         |         A. cochlearis GK00
         |         A. colei OM03
         |           |--A. c. var. colei LK14
         |           `--A. c. var ileocarpa LK14
         |         A. colletioides S95
         |         A. complanata Ca16
         |         A. conferta B00b
         |         A. constricta B14
         |         A. coolgardiensis G04a
         |           |--A. c. ssp. coolgardiensis G04a
         |           `--A. c. ssp. effusa G04a
         |         A. coriacea SM90
         |           |--A. c. ssp. coriacea MLP09
         |           `--A. c. ssp. pendens MLP09
         |         A. coulteri OM03
         |         A. cowleana LK14
         |         A. crassicarpa OM03
         |         A. cuneata B88
         |         A. cunninghamii C15
         |         A. cyclops H93
         |         A. cyperophylla G08
         |         A. dacrydioides LK14
         |         A. dasyphylla M16
         |         A. dawsoni Baker 1897 C08
         |         A. dealbata [=A. decurrens var. dealbata] H93
         |         A. deanei B00a
         |         A. decora Ca16
         |         A. decurrens OM03
         |           |--A. d. var. decurrens B88
         |           `--A. d. var. pauciglandulosa B88
         |         A. deficiens G04b
         |         A. delibrata LK14
         |         A. deltoidea LK14
         |           |--A. d. ssp. deltoidea LK14
         |           `--A. d. ssp. ampla LK14
         |         A. difficilis LK14
         |         A. discolor B88
         |           |--A. d. var. discolor Ch16a
         |           |--A. d. var. angustifolia non A. penninervis var. angustifolia Maiden 1906 Ch16a
         |           `--A. d. var. fraseri [incl. A. maritima] Ch16a
         |         A. disparrima B00b
         |         A. dissimilis LK14
         |         A. distans MLP09
         |         A. dodoneaefolia C15
         |         A. doratoxylon Ca16
         |         A. drepanolobium Ru-S02
         |         A. dunnii LK14
         |         A. elata H06
         |         A. elongata P09
         |         A. erinacea OS04
         |         A. eriopoda DK14
         |         A. evenulosa G04b
         |         A. excelsa Ca16
         |         A. extensa RL05
         |         A. fecunda MLP09
         |         A. ferruginea RaS02
         |         A. filamentosa LK14
         |         A. filicifolia B00b
         |         A. fimbriata B00b
         |         A. flavescens B00b
         |         A. floribunda B88
         |         A. froggattii LK14
         |         A. galioides Ca16
         |         A. gardneri LK14
         |         A. genistifolia B00b
         |         A. georginae N91
         |         A. glauca MM96
         |         A. glaucescens N91
         |         A. gonocarpa LK14
         |         A. gonoclada Ca16
         |         A. gracillima LK14
         |         A. granitica B00b
         |         A. greggii DD06
         |         A. gregorii ML13
         |         A. halliana B00b
         |         A. hamersleyensis EF04
         |         A. hammondii LK14
         |         A. harpophylla M04
         |         A. hemignosta LK14
         |         A. hemiteles G04b
         |         A. hemsleyi LK14
         |         A. heterochroa G04b
         |           |--A. h. ssp. heterochroa G04b
         |           `--A. h. ssp. robertii G04b
         |         A. heterophylla C55
         |         A. hilliana MLP09
         |         A. hippuroides LK14
         |         A. hispidula C15
         |         A. holcocarpa Ca16
         |         A. holosericea B12
         |         A. homalophylla W95
         |         A. horrida G17
         |         A. huegelii GK00
         |         A. humifusa LK14
         |         A. hypermeces LK14
         |         A. imbricata M16
         |         A. implexa H06
         |         A. inaequilatera EF04
         |         A. incurva GK00
         |         A. ingramii B00b
         |         A. intertexta Ch16b
         |         A. intricata G04b
         |         A. irrorata B00b
         |         A. iteaphylla GFS97
         |         A. ixiophylla Benth. 1842 [incl. A. fuliginea Baker 1906, A. glutinosa] M16
         |         A. jasperensis LK14
         |         A. julibrissin [=Albizzia julibrissin] C15
         |         A. julifera Ca16
         |         A. juncifolia B00b
         |         A. juniperina C08
         |           |--A. j. var. juniperina H87
         |           `--A. j. var. brownei H87 [incl. A. acicularis Ch16b]
         |         A. karroo G10
         |         A. kelleri LK14
         |         A. kempeana G04a
         |         A. kenneallyi LK14
         |         A. kimberleyensis LK14
         |         A. koa HH03
         |           |--A. k. var. koa HH03
         |           `--A. k. var. latifolia HH03
         |         A. laccata LK14
         |         A. lamprocarpa LK14
         |         A. lanigera C08
         |           |--A. l. var. lanigera C08
         |           `--A. l. var. venulosa C08
         |         A. laricina GK00
         |         A. lasiocalyx OS04
         |         A. lasiocarpa SM06
         |           |--A. l. var. lasiocarpa OS04
         |           `--A. l. var. sedifolia OS04
         |         A. latronum B03
         |         A. laurifolia M94
         |         A. leiocalyx B00b
         |         A. leiophylla B00b
         |         A. lentiginea LK14
         |         A. leprosa C15
         |         A. leptocarpa B00b
         |         A. leptoclada P09
         |         A. leptopetala [incl. A. murrayana] M16
         |         A. leptophleba LK14
         |         A. leptostachya Ca16
         |         A. leucoclada B00b
         |         A. leucophloea RaS02
         |         A. ligulata B05
         |         A. linearis [=A. mucronata var. linearis] M16
         |         A. linifolia N91
         |         A. linophylla G04a
         |         A. longifolia H02
         |         A. lophantha C15 [=Albizzia lophantha C15; incl. Albizia distachya H93]
         |         A. lunata C08
         |         A. lycopodiifolia LK14
         |         A. lysiphloia LK14
         |         A. macradenia B00b
         |         A. macracantha OA02
         |         A. macrostachya GC05
         |         A. maideni C15
         |         A. maitlandii EF04
         |         A. mangium OM03
         |         A. marramamba EF04
         |         A. mearnsii [incl. A. mollissima] TW07
         |         A. merrallii G04b
         |         A. microbotrya OS04
         |         A. minyura MLP09
         |         A. mitodes LK14
         |         A. moirii G04b
         |           |--A. m. ssp. moirii G04b
         |           `--A. m. ssp. recurvistipula G04b
         |         A. montana Benth. 1842 M16
         |         A. monticola LK14
         |         A. montis-usti CV06
         |         A. mucronata M16
         |         A. multisiliqua LK14
         |         A. myrtifolia GK00
         |         A. nematophylla B00b
         |         A. neovernicosa B14
         |         A. neriifolia GCC02
         |         A. nervosa GK00
         |         A. neurocarpa LK14
         |         A. neurophylla G04a
         |         A. nigripilosa S95
         |         A. nubica B88
         |         A. nuperrima LK14
         |         A. obtusata C15
         |           |--A. o. var. obtusata M16
         |           `--A. o. var. hamiltoni M16
         |         A. obtusifolia B00b
         |         A. oligoneura LK14
         |         A. orthocarpa HD04
         |         A. orthotricha LK14
         |         A. oswaldii MM09
         |         A. oxycedrus H87
         |         A. pachyacra CC03
         |         A. papyrocarpa B00b
         |         A. paradoxa H06
         |         A. paraneura MLP09
         |         A. parramattensis B00b
         |         A. paula LK14
         |         A. pellita LK14
         |         A. pendula B88
         |         A. pennata HK98
         |         A. pennatula HH03
         |         A. pentadenia GK00
         |         A. peregrinalis OM03
         |         A. phlebophylla C15
         |         A. pityoides C15
         |         A. planifrons RaS02
         |         A. platycarpa G14
         |         A. plectocarpa B12
         |         A. podalyriaefolia C15
         |         A. poliochroa G04b
         |         A. polybotrya B00b
         |           |--A. p. var. polybotrya Ca16
         |           `--A. p. var. foliolosa Ca16
         |         A. polystachya B00b
         |         A. prainii G04b
         |         A. pravissima B00b
         |         A. pruinocarpa EF04
         |         A. pruinosa C15
         |         A. pubescens M87
         |         A. pulchella SM06
         |           |--A. p. var. pulchella OS04
         |           |--A. p. var. glaberrima OS04
         |           `--A. p. var. goadbyi GK00
         |         A. pumila B15
         |         A. pycnantha B88
         |         A. pycnophylla M16
         |         A. pyrifolia HD04
         |         A. quadrimarginea G04a
         |         A. quinquenervia G04b
         |         A. quornensis B00b
         |         A. raddiana HF03 [=A. tortilis ssp. raddiana AGF98]
         |         A. ramulosa G04a
         |         A. retinervis LK14
         |         A. retivenea LK14
         |         A. rigens B00b
         |         A. rivalis B88
         |         A. robeorum MLP09
         |         A. robusta BCS02
         |         A. robynsiana CV06
         |         A. rostellifera [incl. A. subbinervia] M16
         |         A. rupicola B00b
         |         A. salicina N91
         |           |--A. s. var. salicina C15
         |           |--A. s. var. varians C15
         |           `--A. s. var. wayae C15
         |         A. saligna OM03
         |         A. scirpifolia Z94
         |         A. sclerophylla B00b
         |         A. sclerosperma EF04
         |         A. senegal PP07
         |         A. sentis Ca16
         |         A. sericata LK14
         |         A. shirleyi B00b
         |         A. singula G04b
         |         A. smallii HH03
         |         A. smeringa LK14
         |         A. sophorae B88 [=A. longifolia var. sophorae H02]
         |         A. sowdenii TW07
         |         A. spectabilis C08
         |         A. spectra LK14
         |         A. stellaticeps LK14
         |         A. stenophylla MM09
         |         A. stenoptera RL05
         |         A. stereophylla S95
         |         A. stigmatophylla LK14
         |         A. stipuligera LK14
         |         A. stowardii EF04
         |         A. stricta C15
         |         A. suaveolens H87
         |         A. subcontorta MLP09
         |         A. sulcata G04b
         |           |--A. s. var. sulcata G04b
         |           `--A. s. var. platyphylla G04b
         |         A. sutherlandi Ca16
         |         A. synchronicia LK14
         |         A. tenuispica LK14
         |         A. tenuissima LK14
         |         A. terminalis SN08
         |         A. tetraneura G04b
         |         A. tortilis HF03
         |         A. torulosa C15
         |         A. trachycarpa MLP09
         |         A. trachyphloia B00b
         |         A. translucens LK14
         |         A. triptera C15
         |         A. tumida VT13
         |           |--A. t. var. tumida LK14
         |           |--A. t. var. extenta LK14
         |           |--A. t. var. kulparn DK14
         |           `--A. t. var. pilbarensis MLP09
         |         A. uncinella G04b
         |         A. undulifolia C15
         |         A. unifissilis G04b
         |         A. urophylla GK00
         |         A. varia GK00
         |         A. verniciflua C15
         |         A. verticillata SN08
         |         A. vestita C15
         |         A. victoriae M04
         |         A. vincentii LK14
         |         A. viscidula C08
         |         A. visco HH03
         |         A. wickhamii LK14
         |         A. willdenowiana GK00
         |         A. willingii LK14
         |         A. xanthina B00b
         |         A. xiphophylla EF04
         |         A. yorkrakinensis G04b
         |         A. zanzibarica Ru-S02
         |--A. ser. Uninerves M16
         |    |--A. subser. Racemosae M16
         |    |--A. (subser. Armatae) lineata M16
         |    `--A. subser. Brevifoliae M16
         |         |--A. aspera M16
         |         `--A. meissneri M16
         |--A. (ser. Calamiformes subser. Uninerves) gonophylla M16
         |--A. ser. Juliflorae M16
         |    |--A. (ser. Juliflorae subser. Dimidiatae) cincinnata M16
         |    |--A. (subser. Falcatae) auriculiformis M16
         |    `--A. subser. Stenophyllae M16
         |         |--A. burkittii C15
         |         `--A. currani Maiden 1916 M16
         |--A. ser. Plurinerves M16
         |    |--A. (ser. Plurinerves subser. Dimidiatae) binervata M16
         |    `--A. subser. Nervosae M16
         |         |--A. cyclopis M16
         |         |--A. homoclada M16
         |         `--A. melanoxylon M16
         `--A. ser. Pungentes M16
              |--A. (ser. Pungentes subser. Plurinerves) striatula M16
              `--A. ser. Pungentes subser. Uninerves M16
                   |--A. carnei Maiden 1916 M16
                   |--A. genistoides M16
                   |--A. quadrisulcata M16
                   `--A. tetragonophylla M16

Nomen nudum: Acacia latipes Gibson & Keighery 2000 GK00

*Type species of generic name indicated

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