Trigonotis peduncularis, copyright Dalgial.

Belongs within: Boraginaceae.
Contains: Pectocarya, Plagiobothrys, Amsinckia, Hackelia, Trichodesmeae, Myosotis, Cryptantha, Cynoglossum.

The Cynoglosseae were recognised by Långström & Chase (2002) as a clade within the Boraginaceae containing taxa with heterocolpate pollen, an undivided style with one stigma, and usually well-developed appendages or wings on the nutlets. A previously recognised division between tribes Cynoglosseae, Eritrichieae and Myosotideae was not supported by Långström & Chase's (2002) molecular analysis. Members of the clade include the stickseeds of the genus Lappula, a Northern Hemisphere genus of annual herbs producing nutlet fruits covered more or less long, barbed prickles (Hickman 1993).

See also: Sending forget-me-nots.

<==Cynoglosseae [Eritrichieae]
    |--Omphalodes verna LC02
    `--+--Trichodesmeae LC02
       `--+--Lappula LC02
          |    |--L. barbata GR98
          |    |--L. microcarpa O88
          |    |--L. redowskii H93
          |    |    |--L. r. var. redowskii H93
          |    |    `--L. r. var. cupulata [incl. L. redowskii var. desertorum] H93
          |    `--L. squarrosa [incl. L. echinata] H93
          `--+--Myosotis LC02
             |--+--Pardoglossum cheirifolium LC02
             |  |--Cryptantha LC02
             |  `--+--Lindelofia longiflora LC02
             |     `--Cynoglossum LC02
             `--Trigonotis LC02
                  |--T. brevipes LC02
                  |--T. multicaulis O88
                  |--T. ovalifolia O88
                  |--T. peduncularis LC02
                  |--T. rotundifolia O88
                  `--T. tibetica O88

Cynoglosseae incertae sedis:
  Actinocarya LC02
  Afrotysonia LC02
  Antiotrema LC02
  Diploloma LC02
  Gyrocaryum LC02
  Harpagonella palmeri LC02, H93
  Heliocarya LC02
  Heterocaryum LC02
  Krynitzkia LC02
  Kuschakewiczia LC02
  Mattia LC02
  Mattiastrum LC02
  Paracaryum LC02
  Pectocarya LC02
  Rindera LC02
  Solenanthus LC02
  Thyrocarpus LC02
  Trachelanthus LC02
  Tysonia LC02
  Anoplocaryum LC02
  Asperugo procumbens LC02, C55
  Bothriospermum LC02
  Craniospermum LC02
  Echiochospermum LC02
  Eremocarya LC02
  Lepechiniella LC02
  Myosotidium Hooker 1859 LC02, A61
    `--M. hortensia (Decne) Baill. 1891 (see below for synonymy) A61
  Nesocaryum LC02
  Omphalolappula Brand 1931 LC02, BR65
    `--O. concava (Mueller) Brand 1931 (see below for synonymy) BR65
  Oreocarya LC02
  Plagiobothrys LC02
  Pseudomertensia LC02
  Rochelia stellulata LC02, C55
  Schistocaryum LC02
  Tianschaniella LC02
  Tretocarya LC02
  Amsinckia LC02
  Hackelia LC02
  Echinospermum LC02
    |--E. deflexum C55
    `--E. lappula C55
  Microula LC02
    |--M. pustulosa O88
    |--M. sikkimensis O88
    `--M. tibetica O88
  Eritrichium LC02
    |--E. canum O88
    |--E. fulvum C55
    |--E. minimum O88
    |--E. nanum H09
    |    |--E. n. ssp. nanum H09
    |    `--E. n. ssp. jankae H09
    `--E. thomsonii O88

Myosotidium hortensia (Decne) Baill. 1891 [=Myosotis hortensia Decne in Deless. 1846; incl. Cynoglossum nobile Hooker 1858, Myosotidium nobile Hook. 1859] A61

Omphalolappula concava (Mueller) Brand 1931 [=Echinospermum concavum Mueller 1860, Cynoglossum concavum Kuntze 1891, Lappula concava Mueller 1882] BR65

*Type species of generic name indicated


[A61] Allan, H. H. 1961. Flora of New Zealand vol. 1. Indigenous Tracheophyta: Psilopsida, Lycopsida, Filicopsida, Gymnospermae, Dicotyledones. R. E. Owen, Government Printer: Wellington (New Zealand).

[BR65] Black, J. M., & E. L. Robertson. 1965. Flora of South Australia. Part IV. Oleaceae–Compositae. W. L. Hawes, Government Printer: Adelaide.

[C55] Candolle, A. de. 1855. Géographie Botanique Raisonée: Ou exposition des faits principaux et des lois concernant la distribution géographique des plantes de l’époque actuelle vol. 2. Librairie de Victor Masson: Paris.

[GR98] Greuter, W., & T. Raus (eds.) 1998. Med-Checklist Notulae, 17. Willdenowia 28: 163–174.

[H09] Heltmann, H. 2009. Der Königstein (Piatra Craiului), die Perle der Burzenländer Gebirge. Mauritiana 20 (3): 515–527.

[H93] Hickman, J. C. (ed.) 1993. The Jepson Manual: Higher Plants of California. University of California Press: Berkeley (California).

[LC02] Långström, E., & M. W. Chase. 2002. Tribes of Boraginoideae (Boraginaceae) and placement of Antiphytum, Echiochilon, Ogastemma and Sericostoma: a phylogenetic analysis based on atpB plastid DNA sequence data. Plant Systematics and Evolution 234: 137–153.

[O88] Ohba, H. 1988. The alpine flora of the Nepal Himalayas: an introductory note. In: Ohba, H., & S. B. Malla (eds) The Himalayan Plants vol. 1. The University Museum, University of Tokyo, Bulletin 31: 19–46.

Last updated: 27 October 2019.

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