Nuculanoidea

Oval yoldia Yoldia myalis, photographed by L. Schroeder.


Belongs within: Bivalvia.
Contains: Nuculana.

The Nuculanoidea are a group of infaunal bivalves whose earliest representatives are recorded from the Ordovician. Their shells are posteriorly elongate, though the degree of elongation varies from only slight to prominently so. Members of the family Nuculanidae possess a resilifer and have a partially internal ligament, whereas the Malletiidae lack a resilifer and their ligament is predominantly external (Cox et al. 1969).

Characters (from Cox et al. 1969): Shell elongate posteriorly, with or without resilifer, pallial sinus usually present.

<==Nuculanoidea
    |  i. s.: Portlandia isonota GW02
    |         Yoldiella lucida GW02
    |--Neilonella [Neilonellidae] GW02
    |    `--N. subovata GW02
    |--Malletiidae H00
    |    |--Malletia Desmoulins 1832 F27a
    |    |    |  i. s.: M. chilensis GW02
    |    |    |         M. gigantea GW02
    |    |    |         M. inequalis GW02
    |    |    |         M. obtusata GW02
    |    |    `--M. (Minormalletia Dall 1908) F27a
    |    |         |--M. (*M.) arciformis F27a
    |    |         `--M. (M.) tenera Marwick 1926 F27a
    |    `--Palaeoneilo Hall & Whitfield 1869 (see below for synonymy) H00
    |         |--*P. constricta Conrad 1842 [=Nuculites constricta] H00
    |         |--P. anthraconeiloides (Chao 1927) ZP86
    |         |--*Synek’ antiquus Barrande 1881 [=*Filius antiquus] H00
    |         |--*Straba’ bohemica Prantl & Růžička 1954 H00
    |         |--P. elliptica (Goldfuss 1837) (see below for synonymy) H00
    |         |--*Olegija’ eugenii Chernyshev 1948 H00
    |         |--‘Straba (*Strabina)’ formosa Prantl & Růžička 1954 H00
    |         |--P. galatea (d’Orbigny 1849) H00
    |         |--P. guangdongensis Zhang & Pojeta 1986 ZP86
    |         |--‘Straba (*Strabiella)’ holynensis Prantl & Růžička 1954 H00
    |         |--*Anthraconeilopsis’ kansana Tasch 1953 H00
    |         |--*Goniodon’ ohioensis Herrick 1888 H00
    |         |--P. oweni ZP86
    |         |--P. phillipsii H00
    |         `--*Anthraconneilo’ taffiana Girty 1911 H00
    `--Nuculanidae [Yoldiidae, Yoldiinae] GW02
         |--Spinula F27a
         |--Nuculana MG-H11
         |--Poroleda Hutton 1893 P61
         |    `--*P. lanceolata (Hutton 1885) [incl. P. pertubata Iredale 1924] P61
         |--Zealeda Marwick 1924 F27a
         |    |--*Z. hamata Marwick 1924 F27a
         |    `--Z. mutabilis Marwick 1926 F27a
         |--Thestyleda Iredale 1929 P61
         |    |--*T. ramsayi [=Leda ramsayi] P61
         |    `--T. investigator Dell 1952 P61
         |--Nucinella Wood 1850 P61
         |    |--*N. miliaris [=Nucula miliaris] P61
         |    `--N. maorianus (Hedley 1904) P61
         |--Austrotindaria Fleming 1948 P61
         |    |--*A. wrighti Fleming 1948 P61
         |    |--A. benthicola Dell 1956 P61
         |    `--A. flemingi Dell 1956 P61
         |--Ledella Verrill & Bush 1897 P61
         |    |--*L. messanensis [=Leda messanensis] P61
         |    |--L. finlayi Powell 1935 P61
         |    |--L. herdmanni Dell 1953 P61
         |    `--L. librata Dell 1952 P61
         |--Ovaleda Iredale 1925 P61
         |    |--*O. tellinaeformis (Hedley 1901) [=Sarepta tellinaeformis] F27a
         |    |--O. aucklandica (Marshall 1918) [=Sarepta aucklandica] F27a
         |    |--O. hamiltoni Powell 1935 P61
         |    `--O. powelli Dell 1950 P61
         |--Neilo Adams 1854 P61
         |    |--*N. australis (Quoy & Gaimard 1835) [=Nucula australis] P61
         |    |--N. awamoana Finlay 1926 F27a
         |    |--N. cymbula (Woods 1917) [=Malletia (Neilo) cymbula] F27a
         |    |--N. ‘elongata’ (Marshall 1917) [=Malletia elongata non Etheridge 1902] F27b
         |    |--N. randsi (Etheridge 1892) [=Nuculana (Yoldia) randsi; incl. Leda elongata] F27b
         |    |--N. rugata Dell 1956 P61
         |    |--N. sinangula Finlay 1926 F27a
         |    |--N. sublaevis Marwick 1926 F27a
         |    `--N. waitaraensis Marwick 1926 F27a
         |--Rollieria Cossmann 1920 (see below for synonymy) H00
         |    |--*R. palmae (Sowerby 1824) [=Nucula palmae] H00
         |    `--R. bronni (Andler 1858) (see below for synonymy) H00
         `--Yoldia MG-H11
              |--Y. amygdalea GW02
              |--Y. ensifera Dall 1897 M12
              |--Y. limaluta (Say 1831) SS07
              |--Y. myalis GW02
              |--Y. thraciaeformis Storer 1838 R78
              `--Y. truncata (Brown 1827) [=Nucula truncata non Nilsson 1827, Leda truncata] F27b

Palaeoneilo Hall & Whitfield 1869 [=Palaeaneilo (l. c.), Paleoneilo (l. c.); incl. Anthraconeilopsis Tasch 1953, Anthraconneilo Girty 1911, Filius Barrande 1881, Goniodon Herrick 1888, Olegija Chernyshev 1948, Straba Prantl & Růžička 1954, Straba (Strabiella Prantl & Růžička 1954), Straba (Strabina Prantl & Růžička 1954), Synek Barrande 1881] H00

Palaeoneilo elliptica (Goldfuss 1837) [=Nucula elliptica, N. muensteri Goldfuss 1840 (nom. van.); incl. Palaeoneilo hamiltoni Wilckens 1927, P. oviformis Troedsson 1951, Nucula patagonidica Leanza 1942, P. patagonidica, Leda quenstedti Tate 1870, L. trapezoidalis Monke 1888, Nucula tunicata Quenstedt 1856] H00

Rollieria Cossmann 1920 [incl. Isoleda Rollier 1923, Isonuculana Cox 1925, Nuculopsis Rollier 1912 non Girty 1911] H00

Rollieria bronni (Andler 1858) [=Leda bronni, Nuculana (Rollieria) bronni; incl. Tellina aequilatera Koch & Dunker 1837 (n. d.), L. aequilatera, Nucula fallax Terquem & Piette 1865, Nucula stoppani Winkler 1886] H00

*Type species of generic name indicated

REFERENCES

Cox, L. R., N. D. Newell et al. 1969. Treatise on Invertebrate Paleontology pt N. Bivalvia vol. 1. The Geological Society of America, Inc. and The University of Kansas.

[F27a] Finlay, H. J. 1927a. A further commentary on New Zealand molluscan systematics. Transactions and Proceedings of the New Zealand Institute 57: 320–485.

[F27b] Finlay, H. J. 1927b. New specific names for austral Mollusca. Transactions and Proceedings of the New Zealand Institute 57: 488–533.

[GW02] Giribet, G., & W. Wheeler. 2002. On bivalve phylogeny: a high-level analysis of the Bivalvia (Mollusca) based on combined morphology and DNA sequence data. Invertebrate Biology 121 (4): 271–324.

[H00] Hodges, P. 2000. The Early Jurassic Bivalvia from the Hettangian and Lower Sinemurian of south-west Britain. Part 1. Monograph of the Paleontographical Society 614: 1–64.

[MG-H11] McEnnulty, F. R., K. L. Gowlett-Holmes, A. Williams, F. Althaus, J. Fromont, G. C. B. Poore, T. D. O’Hara, L. Marsh, P. Kott, S. Slack-Smith, P. Alderslade & M. V. Kitahara. 2011. The deepwater megabenthic invertebrates on the western continental margin of Australia (100–1100 m depths): composition, distribution and novelty. Records of the Western Australian Museum Supplement 80: 1–191.

[M12] Morton, B. 2012. The biology and functional morphology of Nucula pusilla (Bivalvia: Protobranchia: Nuculidae) from Western Australia, Australia: primitive or miniature simplicity? Records of the Western Australian Museum 27 (2): 85–100.

[P61] Powell, A. W. B. 1961. Shells of New Zealand: An illustrated handbook 4th ed. Whitcombe and Tombs Limited: Christchurch.

[R78] Reid, R. G. B. 1978. The systematic, adaptive and physiological significance of proteolytic enzyme distribution in bivalves. Veliger 20 (3): 260–265.

[SS07] Struck, T. H., N. Schutt, T. Kusen, E. Hickman, C. Bleidorn, D. McHugh & K. M. Halanych. 2007. Annelid phylogeny and the status of Sipuncula and Echiura. BMC Evolutionary Biology 7: 57.

[ZP86] Zhang, R., & J. Pojeta, Jr. 1986. New bivalves from the Datang Stage, Lower Carboniferous, Guangdong Province, China. Journal of Paleontology 60 (3): 669–679.

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