Dipterocarpaceae

Stand of Dipterocarpus alatus, photographed by Tony Rodd.

Belongs within: Malvales.

The Dipterocarpaceae are a pantropical family of forest trees. Some species in Asia are economically significant hardwood sources.

Characters (from the Angiosperm Phylogeny Website): Triterpenoid dipterocarpol and sesquiterpene oleoresins present; cork also outer cortical; cambium storied; tyloses present; cortical bundles present; secretory cavities in pith; nodes also 5:5; petiole geniculate; hairs tufted, peltate, etc.; leaves spiral and two-ranked; inflorescence axillary, often branched; androecium initiation centrifugal, anthers more or less versatile, with prolonged connective, median carpel abaxial, ovules apical, stigma slightly lobed or not; calyx thinnish, enlarging somewhat in fruit; seed usually 1, testa vascularized; cotyledons often folded.

<==Dipterocarpaceae
    |  i. s.: Shorea W01
    |           |--S. albida H03
    |           |--S. curtisii W01
    |           |--S. macroptera W01
    |           |--S. parvistipulata K03
    |           |--S. paucifolia W01
    |           `--S. robusta LJ03
    |         Hopea VP93
    |           |--H. mollissima VP93
    |           `--H. papuana C78
    |         Anisoptera WM09
    |           |--A. marginata WM09
    |           `--A. thurifera C78
    |--Pakaramaea [Pakaramaeoideae] T00
    |--Monotoideae T00
    |    |--Monotes T00
    |    |--Marquesia T00
    |    `--Pseudomonotes T00
    `--Dipterocarpus PA-W02 [Dipterocarpoideae T00]
         |--D. alatus PA-W02
         |--D. costatus BAN98
         |--D. grandiflorus P88
         |--D. kerrii Z02
         `--D. oblongifolius P88

*Type species of generic name indicated

REFERENCES

[BAN98] Baum, D. A., W. S. Anderson & R. Nyffeler. 1998. A durian by any other name: taxonomy and nomenclature of the core Malvales. Harvard Papers in Botany 3 (2): 315-330.

[C78] Clunie, N. M. U. 1978. The vegetation. In Handbooks of the Flora of Papua New Guinea vol. 1 (J. S. Womersley, ed.) pp. 1-11. Melbourne University Press: Carlton South (Australia).

[H03] Heads, M. 2003. Ericaceae in Malesia: Vicariance biogeography, terrane tectonics and ecology. Telopea 10 (1): 311-449.

[K03] Kulip, J. 2003. An ethnobotanical survey of medicinal and other useful plants of Muruts in Sabah, Malaysia. Telopea 10 (1): 81-98.

[LJ03] Lim, Y. W., & H. S. Jung. 2003. Irpex hydnoides, sp. nov. is new to science, based on morphological, cultural and molecular characters. Mycologia 95 (4): 694-699.

[PA-W02] Pang, K.-L., M. A. Abdel-Wahab, S. Sivichai, H. M. El-Sharouney & E. B. G. Jones. 2002. Jahnulales (Dothideomycetes, Ascomycota): A new order of lignicolous freshwater ascomycetes. Mycological Research 106 (9): 1031-1042.

[P88] Polunin, I. 1988. Plants and Flowers of Malaysia. Times Editions: Singapore.

[T00] Thorne, R. F. 2000. The classification and geography of the flowering plants: Dicotyledons of the class Angiospermae (subclasses Magnoliidae, Ranunculidae, Caryophyllidae, Dilleniidae, Rosidae, Asteridae, and Lamiidae). The Botanical Review 66: 441-647.

[VP93] Vu V. D., Pham M. G., Nguyen N. C., D. Tuoc, P. Arctander & J. MacKinnon. 1993. A new species of living bovid from Vietnam. Nature 363: 443-445.

[WM09] Wang, H., M. J. Moore, P. S. Soltis, C. D. Bell, S. F. Brockington, R. Alexandre, C. C. Davis, M. Latvis, S. R. Manchester & D. E. Soltis. 2009. Rosid radiation and the rapid rise of angiosperm-dominated forests. Proceedings of the National Academy of Sciences of the USA 106 (10): 3853-3858.

[W01] Watling, R. 2001. The relationships and possible distributional patterns of boletes in south-east Asia. Mycological Research 105 (12): 1440-1448.

[Z02] Zherikhin, V. V. 2002. Pattern of insect burial and conservation. In History of Insects (A. P. Rasnitsyn & D. L. J. Quicke, eds) pp. 17-63. Kluwer Academic Publishers: Dordrecht.