Poecilostomatoida

Drawing of Paranthessius anemoniae, from here.

Belongs within: Podoplea.
Contains: Splanchnotrophidae, Clausiidae, Clausidiidae, Lichomolgidae, Pseudanthessius.

The Poecilostomatoida are a group of copepods that are generally parasitic on other marine animals.

Characters (from Nicholls 1944): First antenna not hinged in male; second antenna without an exopod; mouth parts non-suctorial, non-raptorial, mandible lacking palp.

<==Poecilostomatoida [Poecilostoma]
    `--Lichomolgoidea [Lichomolginae] K03
         |--Lichomolgidae K03
         |--Pseudanthessius K03
         `--Rhynchomolgidae K03
              |--Critomolgus anthopleurus Kim 1996 K03
              |--Doridicola longicauda K03
              `--Paranthessius Claus 1889 K03, N44 [Sabelliphilidae, Sabelliphilinae]
                   |--P. anemoniae K03
                   |--‘Heteranthessius’ dubius (Scott 1903) N44
                   |--‘Pestalichomolgus’ pectinis (Pesta 1908) N44
                   `--P. propinquus Nicholls 1944 N44

Poecilostomatoida incertae sedis:
  Pionodesmatidae BO01
  Synapticolidae MD01
  Antheacheridae MD01
  Echiurophilus [Echiurophilidae] H01
  Splanchnotrophidae H01
  Gastrodelphyidae H01
  Clausiidae HK03
  Micrallecto Stock 1971 [incl. Nannallecto Stock 1973; Micrallectidae] H09
    |--*M. uncinata Stock 1971 H01
    `--M. fusii Stock 1973 [=*Nannallecto fusii] H01
  Synaptiphilus Canu & Cuénot 1892 [Synaptiphilidae] HK03
    |--S. cantacuzenei BK77
    |--S. luteus [incl. Colaceutes mulleri Hartmann 1856] BK77
    `--S. tridens (Scott & Scott 1893) [=Remigulus tridens] BK77
  Chondracanthidae H01
    |  i. s.: Diocus lycenchelus K-MC02
    |--Pharodinae [Pharodidae] H01
    |--Lernentominae H01
    `--Chondracanthus K-M02c [Chondracanthinae H01]
         |--C. cottunculi K-M02c
         `--C. merluccii K-M02b
  Philoblennidae H01
    |--Philoblenna arabici K03
    |--Chondrocarpus Bassett-Smith 1903 H01
    |    `--*C. reticulosus H01
    `--Briarella Bergh 1876 H01
         |--*B. microcephala Bergh 1876 H01
         |--B. disphaerocephala Monod & Dollfus 1932 H01
         `--B. risbeci Monod 1928 H01
  Clausidiidae MD01
  Lernaeosoleidae [Lernaeosoleinae] MD01
  Ergasilidae [Amazonicopeidae] MD01
    |--Neoergasilus japonicus K03
    `--Ergasilus K03
         |--E. briani K03
         |--E. funduli K-M02a
         |--E. labracis K-M02d
         `--E. sieboldi K03
  Anchimolgidae MD01
  Anomoclausiidae MD01
  Anthessiidae MD01
  Bomolochidae MD01
  Chitonophilidae MD01
  Corallovexiidae MD01
  Corycaeus B26 [Corycaeidae MD01]
    |--C. carinatus B26
    |--C. elongatus B26
    |--C. ovalis B26
    `--C. speciosus B26
  Entobiidae MD01
  Erebonasteridae MD01
  Eunicicolidae MD01
  Herpyllobiidae MD01
  Intramolgidae MD01
  Lamippidae MD01
  Lubbockiidae MD01
  Mesoglicolidae MD01
  Pseudomyicola K03 [Myicolidae MD01]
    `--P. spinosus K03
  Trochicola H01 [Mytilicolidae MD01]
    `--T. entericus Dollfus 1914 H01
  Nereicolidae [Anomopsyllidae] MD01
    |--Anomopsyllus pranizoides N44
    `--Dysgenopsyllus Nicholls 1944 N44
         `--*D. reevesbyensis Nicholls 1944 N44
  Nucellicolidae MD01
  Oncaea B26 [Oncaeidae MD01]
    |--O. conifera B26
    |--O. minuta B26
    `--O. venusta B26
  Paralubbockiidae MD01
  Philichthyidae MD01
  Phyllodicolidae MD01
  Polyankylidae MD01
  Saccopsidae MD01
  Sapphirinidae MD01
  Serpulidicolidae MD01
  Shiinoidae MD01
  Spiophanicolidae MD01
  Taeniacanthus K03 [Taeniacanthidae MD01]
    |--T. balistae K-M02e
    `--T. lagocephali K03
  Tegobomolochidae MD01
  Telsidae MD01
  Thamnomolgidae MD01
  Tucca K-M02f [Tuccidae MD01]
    `--T. impressus K-M02f
  Urocopiidae MD01
  Vahiniidae MD01
  Ventriculinidae MD01
  Xarifia H94 [Xarifiidae MD01]
    |--X. diminuta Humes & Ho 1967 H94
    |--X. finitima Humes 1985 H94
    `--X. longipes Humes 1962 H94
  Xenocoelomatidae MD01

*Type species of generic name indicated

REFERENCES

[BK77] Barel, C. D. N., & P. G. N. Kramers. 1977. A survey of the echinoderm associates of the north-east Atlantic area. Zoologische Verhandelingen 156: 1-159.

[B26] Bigelow, H. B. 1926. Plankton of the offshore waters of the Gulf of Maine. Bulletin of the Bureau of Fisheries 40 (2): 1-509.

[BH88] Boxshall, G. A., & A. G. Humes. 1988. A new genus of Lichomolgidae (Copepoda: Poecilostomatoida) associated with a phoronid in Hong Kong. Bulletin of the British Museum (Natural History), Zoology Series 54 (6): 301-307.

[BO01] Boxshall, G. A., & S. Ohtsuka. 2001. Two new families of copepods (Copepoda: Siphonostomatoida) parasitic on echinoderms. Journal of Crustacean Biology 21 (1): 96-105.

[HK03] Ho, J.-S., & I.-H. Kim. 2003. New clausiid copepods (Poecilostomatoida) associated with polychaetes of Korea, with cladistic analysis of the family Clausiidae. Journal of Crustacean Biology 23 (3): 568-581.

[H94] Humes, A. G. 1994. Two species of Paramolgus (Copepoda: Poecilostomatoida: Lichomolgidae) associated with the scleractinian Pavona in New Caledonia with a key to the females of Paramolgus. Beaufortia 44 (1): 1-9.

[H01] Huys, R. 2001. Splanchnotrophid systematics: A case of polyphyly and taxonomic myopia. Journal of Crustacean Biology 21 (1): 106-156.

[K03] Kim, I.-H. 2003. Copepodid stages of Critomolgus anthopleurus (Copepoda, Poecilostomatoida, Rhynchomolgidae). Journal of Crustacean Biology 23 (3): 558-567.

[K-M02a] Klein-MacPhee, G. 2002a. Smelts. Family Osmeridae. In Bigelow and Schroeder’s Fishes of the Gulf of Maine (B. B. Collette & G. Klein-MacPhee, eds) 3^rd ed. pp. 162-170. Smithsonian Institute Press: Washington.

[K-M02b] Klein-MacPhee, G. 2002b. Silver hakes. Family Merlucciidae. In Bigelow and Schroeder’s Fishes of the Gulf of Maine (B. B. Collette & G. Klein-MacPhee, eds) 3^rd ed. pp. 217-222. Smithsonian Institute Press: Washington.

[K-M02c] Klein-MacPhee, G. 2002c. Fathead sculpins. Family Psychrolutidae. In Bigelow and Schroeder’s Fishes of the Gulf of Maine (B. B. Collette & G. Klein-MacPhee, eds) 3^rd ed. pp. 357-358. Smithsonian Institute Press: Washington.

[K-M02d] Klein-MacPhee, G. 2002d. Temperate basses. Family Moronidae. In Bigelow and Schroeder’s Fishes of the Gulf of Maine (B. B. Collette & G. Klein-MacPhee, eds) 3^rd ed. pp. 374-389. Smithsonian Institute Press: Washington.

[K-M02e] Klein-MacPhee, G. 2002e. Triggerfishes. Family Balistidae. In Bigelow and Schroeder’s Fishes of the Gulf of Maine (B. B. Collette & G. Klein-MacPhee, eds) 3^rd ed. pp. 590-592. Smithsonian Institute Press: Washington.

[K-M02f] Klein-MacPhee, G. 2002f. Porcupinefishes. Family Diodontidae. In Bigelow and Schroeder’s Fishes of the Gulf of Maine (B. B. Collette & G. Klein-MacPhee, eds) 3^rd ed. pp. 601-602. Smithsonian Institute Press: Washington.

[K-MC02] Klein-MacPhee, G., & B. B. Collette. 2002. Eelpouts. Family Zoarcidae. In Bigelow and Schroeder’s Fishes of the Gulf of Maine (B. B. Collette & G. Klein-MacPhee, eds) 3^rd ed. pp. 466-474. Smithsonian Institute Press: Washington.

[MD01] Martin, J. W., & G. E. Davis. 2001. An updated classification of the Recent Crustacea. Natural History Museum Los Angeles County, Science Series 39: 1-124.

[N44] Nicholls, A. G. 1944. Littoral Copepoda from South Australia. (II) Calanoida, Cyclopoida, Notodelphyoida, Monstrilloida and Caligoida. Records of the South Australian Museum 8 (1): 1-62.